 | | | | | | | | | | | | | | | | | | | | | | | | | | | # Species: 2887 |
|  | |  | | With 94 genera and 1285 species , this family is the largest of the scorpion families. This family is widespread around the world (not found in Antarctica and New Zealand), and are found in tropical, subtropical and partly in temperate habitats. 26.07.22: Lowe & Kovarik (2022) has synonymized the family Microcharmidae Lourenco, 1996 with Buthidae. The two genera (Microcharmus andNeoprotobuthus) and 17 species are all transfered to Buthidae. August 2019: The family Microcharmidae has been resurrected by Lourenco et al., 2019 and belinging taxa have been transferred back from ButhidaeInformation about the family Microcharmidae. Kovarik (2007) has published a revision of the genusHottentotta. Note that the medical importantMesobuthus tamulus and two other AsianMesobuthus have been transfered toHottentotta. Kovarik's revision can be downloadedHERE (browse down to issue 58). Paraorthochirus has been synonymized withOrthochirus by Navidpour et al., 2008). The members of this family are small and medium sized scorpions with a triangular sternum (some genera have a more pentagonal sternum). In many genera (e.g.,Androctonus,Apistobuthus,Parabuthus, and other Old World genera) the cauda is usually strong and powerful and the pedipalps are often very slender whereas other genera have elongate appendages, especially the males (e.g.,Centruroides,Lychas,Isometrus and related genera). Many species are yellow or brown (or variations of these colors), but black forms are also present. Some species have quite intricate and attractive patterns (e.g.,Lychas andIsometrus) and colours (e.g.,Centruroides andUroplectes). Sizes range from about 20 mm (likeMicrotityus andMicrobuthus) to over 120 mm (as inAndroctonus, someCentruroides,Apistobuthus and others). An indentification key to the genusCompsobuthus was published byKovarik (2012) [Free full text]. Several species in this family are highly toxic, but fewer than 20 can be lethal to man. The most notorious species are found in the genusAndroctonus, Centruroides, Hottentotta, Leiurus, Parabuthus andTityus. SeeScorpions of medical importance page for further information. Unlike most other scorpion families, the Buthidae are of both scientific and public interest due to their medical importance in many parts of the world. Several species have also made their way into the hobby. Many of these are safe to be kept by experienced collectors, but the most dangerous species should be avoided. Androctonus crassicauda habitat picture from Kuwait are locatedHERE. | Aegaeobuthus Kovarik, 2019* (4)
A. cyprius (Gantenbein & Kropf, 2000)*
A. gallianoi (Ythier, 2018)*
A. gibbosus (Brulle, 1832)
A. nigrocinctus (Ehrenberg, 1828)*
| Afrolychas Kovarik, 2019* (2)
A. braueri (Kraepelin, 1896)
A. burdoi (Simon, 1882)
| Afroisometrus Kovarik, 1997 (1)
A. minshullae (Fitzpatrick, 1994)
| Akentrobuthus Lamoral, 1976 (2)
A. atakora Vignoli & Prendini, 2008*
A. leleupi Lamoral, 1976 | Alayotityus Armas, 1973 (8)
A. delacruzi Armas, 1973*
A. feti Teruel, 2004*
A. granma Armas, 1984
A. juraguaensis Armas, 1973
A. lapidicola Teruel, 2002*
A. nanus Armas, 1973
A. pallidus Teruel, 2002*
A. sierramaestrae Armas, 1973
| Androctonus Ehrenberg, 1828 (43)
A. aeneas C. L. Koch, 1839*
A. afghanus Lourenço & Qi, 2006*
A. agrab Ythier & Lourenço, 2022*
A. ajjer Ythier, Sadine, Alioua & Lourenço, 2025*
A. aleksandrplotkini Lourenço & Qi, 2007*
A. amoreuxi(Audouin,1826)
A. australis(Linnaeus, 1758)
A. baluchicus (Pocock, 1900)*
A. barbouri (Werner, 1932)*
A. bartolozzii Rossi & Merendino, 2016*
A. bicolor Ehrenberg, 1828
A. bourdoni Vachon, 1948*
A. burkinensis Ythier, 2021*
A. cacahuati Lourenço, 2023*
A. cholistanus Kovarik & Ahmed, 2013*
A. crassicauda (Olivier, 1807)
A. dekeyseri Lourenço, 2005*
A. donairei Rossi, 2015*
A. eburneus (Pallary, 1928)*
A. finitimus (Pocock, 1897)
A. gonneti Vachon, 1948*
A. hoggarensis (Pallary, 1929)
A. ishtar Yagmur, Kachel, Al-Khazali, Al-Jubouri & Ali 2025*
A. kunti Yagmur, 2023*
A. liouvillei (Pallary, 1924)*
A. maelfaiti Lourenço, 2005*
A. mauritanicus (Pocock, 1902)
A. maroccanus Lourenço, Ythier & Leguin, 2009*
A. pallidus Lourenço, Duhem & Cloudsley-Thompson, 2012*
A. robustus Kovarik & Ahmed, 2013*
A. rostami Barahoei, Mirshamsi, Amiri, Moeinadin & Rakhshani, 2025*
A. santi Lourenço, 2015*
A. sergenti Vachon, 1948
A. simonettai Rossi, 2015*
A. sistanus Barahoei & Mirshamsi, 2022*
A. sumericus Al-Khazali & Yagmur, 2023*
A. tenuissimus Teruel, Kovarik & Turiel, 2013*
A. tibesti Lourenço & El-Hennawy, 2022*
A. tigrai Lourenço, Rossi & Sadine 2015*
A. tihamicus Alqahtani, Yagmur & Badry 2023*
A. togolensis Lourenço, 2008*
A. tropeai Rossi, 2015*
A. turkiyensis Yagmur, 2021*
| Anomalobuthus Kraepelin, 1900 (6)
A. krivochatskyi Teruel, Kovarik & Fet, 2018*
A. lowei Teruel, Kovarik & Fet, 2018*
A. pavlovskyi Teruel, Kovarik & Fet, 2018*
A. rickmersi Kraepelin, 1900
A. talebii Teruel, Kovarik, Navidpour & Fet, 2014*
A. zarudnyi (Birula, 1911)*
| Apistobuthus Finnegan, 1932 (2)
A. pterygocercus Finnegan, 1932
A. susanae Lourenço, 1998
| Australobuthus Locket, 1990 (1)
A. xerolimniorum Locket, 1990
| Babycurus Karsch, 1886 (14)
B. ansorgei Hirst, 1911
B. brignolii Lourenço & Rossi, 2017* [nomen dubium]
B. buettneri Karsch, 1886
B. centrurimorphus Karsch, 1886*
B. dunlopi Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015*
B. gigas Kraepelin, 1896
B. jacksoni (Pocock, 1890)
B. kirki (Pocock, 1890)*
B. melanicus Kovarik, 2000*
B. multisubaculeatus Kovarik, 2000*
B. pictus Pocock, 1896*
B. solegladi Lourenço, 2005*
B. taramassoi Borelli, 1919*
B. wituensis Kraepelin, 1913
| Baloorthochirus Kovarik, 1996 (1)
B. becvari Kovarik, 1996
| Barbaracurus Kovarik, Lowe & Stahlavsky, 2018 (12)
B. feti Kovarik, Lowe &Stahlavsky & Hurre, 2019*
B. exquisitus (Lowe, 2000)*
B. hofereki Kovarik, 2024*
B. kabateki Kovarik, Lowe, Stahlavsky & Just, 2022*
B. prudenti (Lourenço, 2013)*
B. somalicus (Hirst, 1907)*
B. sofomarensis (Kovarik, Lowe, Seiter, Pliskova &Stahlavsky, 2015)*
B. subpunctatus (Borelli, 1925)*
B. ugartei (Kovarik, 2000)*
B. winklerorum Kovarik, Lowe &Stahlavsky, 2018*
B. yemenensis Kovarik, Lowe &Stahlavsky, 2018*
B. zambonellii (Borelli, 1902)*
| | Birulatus Vachon, 1974 (4)
B. astartiae Stathi & Lourenço, 2003*
B. haasi Vachon, 1974
B. israelensis Lourenço, 2002*
B. jordanensis Lourenço, Al-Saraireh, Afifeh, Baker, Bader-Katbeh & Amr, 2021*
| Buthacus Birula, 1908 (36)
B. agarwali Zambre & Lourenço, 2010*
B. ahaggar Lourenço, Kourim & Sadine, 2017*
B. algerianus Lourenço, 2006*
B. amitaii Cain, Gefen & Prendini, 2021*
B. arava Cain, Gefen & Prendini, 2021*
B. arenicola (Simon, 1885)
B. armasi Lourenço, 2013*
B. bicolor Afifeh, Al-Saraireh, Baker, Amr & Lourenço, 2022*
B. birulai Lourenço, 2006*
B. clevai Lourenço, 2001*
B. deserticus Sadine, Souilem, Lourenço & Ythier, 2024*
B. elmenia Lourenço & Sadine, 2017*
B. foleyi Vachon, 1948
B. frontalis Werner, 1936 (nomen dubium)
B. fuscata Pallary, 1929*
B. golovatchi Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. leptochelys (Ehrenberg, 1829)
B. levyi Cain, Gefen & Prendini, 2021*
B. maroccanus Lourenço, 2006*
B. mahraouii Lourenço, 2004*
B. nigerianus Lourenço & Qi, 2006*
B. nigroaculeatus Levy et al., 1973*
B. nitzani Levy, Amitai & Shulov, 1973
B. occidentalis Vachon, 1953*
B. pakistanensis Lourenço & Qi, 2006*
B. sadinei Ythier, 2022*
B. samiae Lourenço & Sadine, 2015*
B. spatzi (Birula, 1911)
B. spinatus Lourenço, Bissati & Sadine, 2016*
B. stockmanni Kovarik, Lowe & Stahlavsky, 2016*
B. striffleri Lourenço, 2004*
B. tadmorensis (Simon, 1892)*
B. villiersi Vachon, 1949
B. williamsi Lourenço & Leguin, 2009*
B. yotvatensis Levy, Amitai & Shulov, 1973
B. ziegleri Lourenço, 2000*
| Butheoloides Hirst, 1925 (19)
B. annieae Lourenço, 1986
B. aymerichi Lourenço, 2002*
B. charlotteae Lourenço, 2000*
B. cimrmani Kovarik, 2003*
B. granulatus Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. grosseri Kovarik, 2016*
B. hirsti Lourenço, 1996
B. littoralis Lourenço, Touloun & Boumezzough, 2011*
B. maroccanus Hirst, 1925
B. milloti Vachon, 1948
B. monodi Vachon, 1950
B. nuer Kovarik, 2015*
B. occidentalis Lourenço, Slimani & Berahou, 2003*
B. polisi Lourenço, 1996
B. savanicola Lourenço, 2013*
B. schwendingeri Lourenço, 2002*
B. slimanii Lourenço, 2010*
B. vanderberghi Lourenço, 2015*
B. wilsoni Lourenço, 1995
| Butheolus Simon, 1882 (6)
B. andersoni (Pocock, 1895)*
B. gallagheri Vachon, 1980
B. hallani Lourenço & Rossi, 2017*
B. harrisoni Lowe, 2018*
B. thalassinus Simon, 1882
B. villosus Hendrixson, 2006*
| Buthiscus Birula, 1905 (2)
B. bicalcaratus Birula, 1905
B. ifoghas Ythier & Lourenco, 2023*
| Buthoscorpio Werner, 1936 (5)
B. chinnarensis Aswathi, Sureshan & Lourenço, 2015*
B. indicus Lourenço, 2012*
B. politus (Pocock, 1899)
B. rayalensis Javed, Rao, Mirza, Sanap & Tampal, 2010*
B. sarasinorum (Karsch, 1891)
| Buthus Leach, 1815 (81)
B. adrianae Rossi, 2013
B. ahaggar Ythier, Sadine, Haddadi & Lourenco 2021*
B. ajax (C. L. Koch, 1839)*
B. alacanti Teruel & Turiel 2020*
B. albengai Lourenço, 2003*
B. amri Lourenço, Yagmur & Duhem 2010*
B. apiatus Lourenço, El Bouhissi & Sadine, 2020*
B. atlantis Pocock, 1889
B. aures Lourenço & Sadine 2016*
B. awashensis Kovarik, 2011*
B. baeticus Teruel & Turiel 2020*
B. balmensis Ythier & Laborieux, 2022*
B. barcaeus Birula, 1909*
B. berberensis Pocock, 1900*
B. bobo Ythier, 2021*
B. bonito Lourenço & Geniez, 2005*
B. boumalenii Touloun & Boumezzough, 2011*
B. boussaadi Lourenço, Chichi & Sadine, 2018*
B. brignolii Lourenço, 2003*
B. castellano Teruel & Turiel 2022*
B. centroafricanus Lourenço, 2016*
B. chambiensis Kovarik, 2006*
B. confluens Lourenço, Touloun & Boumezzough, 2012*
B. danyii Rossi, 2017*
B. delafuentei Teruel & Turiel 2020*
B. draa Lourenço & Slimani, 2004*
B. dunlopi Kovarik, 2006*
B. duprei Rossi & Tropea, 2016*
B. egyptiensis Lourenço, 2012*
B. elhennawyi Lourenço, 2005*
B. elizabethae Lourenço, 2005*
B. elmoutaouakili Lourenço & Qi, 2006*
B. elongatus Rossi, 2012*
B. gabani Ythier, 2021*
B. garcialorcai Teruel & Turiel 2020*
B. gonzalezdelavegai Gonzalez-Moline & de Armas, 2024*
B. goyffoni Abidi, Sadine & Lourenco 2021*
B. halius (C. L. Koch, 1839)*
B. hassanini Lourenço, Duhem & Cloudsley-Thompson, 2012*
B. iaspis Teruel & Turiel 2022*
B. ibericus Lourenço & Vachon, 2004*
B. intermedius (Ehrenberg, 1829)*
B. intumescens (Ehrenberg in Hemprich & Ehrenberg, 1829)*
B. israelis Shulov & Amitai, 1959*
B. jianxinae Lourenço, 2005*
B. karoraensis Rossi & Tropea, 2016*
B. kunti Yagmur, Koc & Lourenço, 2011*
B. labuschagnei Lourenço, 2015*
B. lienhardi Lourenço, 2003*
B. lourencoi Rossi, Tropea & Yagmur, 2013*
B. maamora Ythier, 2023*
B. malhommei Vachon, 1949*
B. manchego Teruel & Turiel 2020*
B. mardochei Simon, 1878*
B. mariefranceae Lourenço, 2003*
B. maroccanus Birula, 1903
B. montanus Lourenço & Vachon, 2004*
B. nabataeus Lourenço, Abu Afifeh & Al-Saraireh, 2021*
B. nigrovesiculosus Hirst, 1925*
B. occidentalis Lourenço, Sun & Zhu, 2009*
B. occitanus (Amoreux, 1789)
B. orientalis Lourenço & Simon, 2012*
B. oudjanii Lourenço, 2017*
B. paris (C. L. Koch, 1839)*
B. parroti Vachon, 1949*
B. pedrosousai Teruel & Turiel 2021*
B. pococki Kovarik, Stahlavsky & Elmi, 2020*
B. prudenti Lourenço & Leguin, 2012*
B. pusillus Lourenço, 2013*
B. pygmaeus Kovarik, Lowe, Stahlavsky & Elmi, 2025*
B. pyrenaeus Ythier, 2021*
B. rochati Lourenço, 2003*
B. saharicus Sadine, Bissati & Lourenço, 2015*
B. serrano Teruel & Turiel 2020*
B. somalilandus Kovarik, Stahlavsky & Elmi, 2020*
B. tassili Lourenço, 2002*
B. trinacrius Lourenço & Rossi, 2013*+
B. tunetanus (Herbst, 1800)*
B. turkana Ythier & Lourenço, 2022*
B. yemenensis Lourenço, 2008*
B. zeylensis Pocock, 1900*
| Centruroides Marx, 1890 (104)
C. alayoni Armas, 1999*
C. altagraciae Teruel, de Armas & Kovarik, 2015*
C. anchorellus Armas, 1976
C. arctimanus Armas, 1976
C. baergi Hoffmann, 1932*
C. baldazoi Ponce-Saavedra, Linares-Guillen & Quijano-Ravell, 2022*
C. balsasensis Ponce & Francke, 2004*
C. bani Armas & Marcano Fondeur, 1987
C. baracoae Armas, 1976*
C. barbudensis (Pocock, 1898)
C. berstoni Goodman, Prendini, Francke & Esposito, 2021*
C. bertholdii (Thorell, 1876)
C. bicolor (Pocock, 1898)
C. bonito Quijano-Ravell, Teruel & Ponce-Saavedra, 2016*
C. caral Armas & Trujillo, 2013
C. caribbeanus Teruel & Myers, 2017*
C. chamela Ponce-Saavedra & Francke, 2011*
C. chamulaensis Hoffmann, 1932
C. catemacoensis Goodman, Prendini, Francke & Esposito, 2021*
C. chanae Goodman, Prendini, Francke & Esposito, 2021*
C. chiapanensis Hoffmann, 1932
C. concordia Armas & Teruel, 2021*
C. cuauhmapan Goodman, Prendini, Francke & Esposito, 2021*
C. edwardsii (Gervais, 1843)*
C. elegans (Thorell, 1876)
C. exilicauda (Wood, 1863)
C. exilimanus Teruel & Stockwell, 2002*
C. exsul (Meise, 1933)
C. fallassisimus Armas & Trujillo, 2010*
C. farri Armas, 1976
C. flavopictus (Pocock, 1898)
C. franckei Santibanez-Lopez & Contreras-Felix, 2013*
C. fulvipes (Pocock, 1898)
C. galano Teruel, 2001*
C. gracilis (Latreille, 1804)
C. granosus (Thorell, 1876)*
C. griseus (C. L. Koch, 1844)
C. guanensis Franganillo, 1930
C. hamadryas Goodman, Prendini, Francke & Esposito, 2021*
C. hentzi (Banks, 1900)
C. hirsuticauda Teruel, 2011*
C. hirsutipalpus Ponce-Saavedra & Francke, 2009*
C. hoffmanni Armas, 1996
C. huichol Teruel, Ponce-Saavedra & Quijano-Ravell, 2015*
C. infamatus (C.L. Koch, 1844)
C. insulanus (Thorell, 1876)
C. ixil Trujillo & Armas, 2016*
C. jaragua Armas, 1999*
C. jorgeorum Santiago-Blay, 2009*
C. koesteri Kraepelin, 1912
C. lauriadnae Ponce-Saavedra & Francke, 2019*
C. lenca Armas & Cubas-Rodriguez, 2025*
C. limbatus (Pocock, 1898)
C. limpidus (Karsch, 1879)
C. luceorum Armas, 1999*
C. lucidus Teruel, Armas & Kovarik, 2015*
C. marcanoi Armas, 1981
C. margaritatus (Gervais, 1841)
C. mascota Ponce-Saavedra & Francke, 2011*
C. meisei Hoffmann, 1939*
C. melanodactylus Teruel, 2001*
C. navarroi Teruel, 2001*
C. nigrescens (Pocock, 1898)
C. nigrimanus (Pocock, 1898)
C. nigropunctatus Teruel, 2006*
C. nigrovariatus (Pocock, 1898)
C. nitidus (Thorell, 1876)
C. noxius Hoffman, 1932
C. ochraceus (Pocock, 1898)
C. orizaba Armas & Martin-Frias, 2003*
C. ornatus Pocock, 1902*
C. pallidiceps Pocock, 1902
C. panamensis Arias & Esposito, 2014*
C. platnicki Armas, 1981
C. pococki Sissom & Francke, 1983
C. polito Teruel, 2007*
C. poncei Teruel, Kovarik, Baldazo-Monsivais & Hoferek, 2015*
C. possanii Gonzalez-Santillan, Galan-Sanchez & Valdez-Velazquez, 2019*
C. rileyi Sissom, 1995
C. robertoi Armas, 1976
C. rodolfoi Santibanez-Lopez & Contreras-Felix, 2013*
C. romeroi Quijano-Ravell, De Armas, Francke & Ponce-Saavedra, 2019*
C. ruana Quijano-Ravell & Ponce-Saavedra, 2016*
C. sanandres Armas, Sarmiento & Florez 2012*
C. sasae Santiago-Blay, 2009*
C. schmidti Sissom, 1995
C. sculpturatus Ewing, 1928*
C. serrano Santibanez-Lopez & Ponce-Saavedra, 2009*
C. simplex (Thorell, 1876)*
C. sissomi Armas, 1996
C. spectatus Teruel, 2006*
C. stockwelli Teruel, 2001*
C. suffusus Pocock, 1902
C. tapachulaensis Hoffmann, 1932*
C. tecomanus Hoffmann, 1932*
C. tenuis Thorell, 1876*
C. terueli de Armas & Cubas-Rodriguez, 2023*
C. testaceus (DeGeer, 1778)
C. thorellii (Kraepelin, 1891)
C. tuxtla Armas, 1999*
C. underwoodi Armas, 1976
C. villegasi Baldazo-Monsivaiz, Ponce-Saavedra & Flores-Moreno, 2013*
C. vittatus (Say, 1821)
C. yucatanensis Goodman, Prendini, Francke & Esposito, 2021*
| Charmus Karsch, 1879 (5)
C. brignolii Lourenço, 2000*
C. dakshini Joshi, Deshpande, Ukale, Gowande, Bilat, Kovarik, Mottaz, Stahlavsky, Bastawade, Monod & Sulakhe, 2025*
C. laneus Karsch, 1879
C. saradieli Kovarik, Lowe, Ranawana, Hoferek & Jayarathne 2016*
C. sinhagadensis Tikader & Bastawade, 1983
| Chaneke Francke, Teruel & Santibanez-Lopez, 2014* (4)
C. aliciae (Armas & Frias, 1998)*
C. baldazoi Kovarik, Teruel & Lowe, 2016*
C. fogoso Francke, Teruel & Santibanez-Lopez, 2014*
C. hofereki Kovarik, Teruel & Lowe, 2016*
| Cicileiurus Teruel, 2007* (1)
C. monticola Teruel, 2007*
| Cicileus Vachon, 1948 (5)
C. exilis (Pallary, 1928)
C. cloudsleythompsoni Lourenço, 1999*
C. hoggarensis Lourenço & Rossi, 2015*
C. latellai Lourenço & Rossi, 2015*
C. montanus Lourenço & Rossi, 2015*
| Compsobuthus Vachon, 1949 (57)
C. abyssinicus (Birula, 1903)
C. air Lourenço & Rossi, 2018*
C. acutecarinatus (Simon, 1882)
C. afghanus Kovarik & Ahmed, 2007*
C. andresi Lourenço, 2004*
C. arabicus Levy, Amitai & Shulov, 1973
C. armenicus Lourenço & Leguin, 2015*
C. atrostriatus (Pocock, 1897)*
C. berlandi Vachon, 1950
C. birulai Lourenço, Leguin & Duhem, 2010*
C. boucheti Lourenço, Duhem & Cloudsley-Thompson, 2012*
C. brevimanus (Werner, 1936)
C. carmelitis Levy, Amitai & Shulov, 1973
C. eritreaensis Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
C. egyptiensis Lourenço, Sun & Zhu, 2009*
C. garyi Lourenço & Vachon, 2001*
C. henrii Ythier & Ullrich, 2025*
C. humaae Amir, Kamaluddin & Kahn, 2005* (nomen dubium)
C. jakesi Kovarik, 2003*
C. jordanensis Levy, Amitai & Shulov, 1973
C. kabateki Kovarik, 2003*
C. kaftani Kovarik, 2003*
C. khaybari Abu Afifeh, Aloufi & Al-Saraireh, 2021*
C. klaptoczi (Birula, 1909)
C. krali Kovarik, 2012*
C. levyi Kovarik, 2012*
C. longipalpis Levy, Amitai & Shulov, 1973
C. mahazat Ythier & Lourenço, 2023*
C. maidensis Kovarik, 2018*
C. maindroni (Kraepelin, 1900)
C. manzonii (Borelli, 1915)
C. matthiesseni (Birula, 1905)
C. nematodactylus Lowe, 2009*
C. pakistanus Kovarik & Ahmed, 2007*
C. pallidus Hendrixson, 2006*
C. persicus Navidpour, Soleglad, Fet & Kovarik, 2008*
C. petriolii Vignoli, 2005*
C. plutenkoi Kovarik, 2003*
C. polisi Lowe, 2001*
C. rugosulus (Pocock, 1900)
C. satpuraensis Waghe, Gangalmale & Khandekar, 2022*
C. schmiedeknechti Vachon, 1949*
C. seicherti Kovarik, 2003*
C. setosus Hendrixson, 2006*
C. simoni Lourenço, 1999*
C. sindicus Kovarik & Ahmed, 2011*
C. somalilandus Kovarik, 2012*
C. starhai Kovarik, 2025*
C. tassili Lourenço, 2010*
C. tofti Lourenço, 2001*
C. tombouctou Lourenço, 2009*
C. turieli Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. ullrichi Kovarik, Lowe, Stockmann & Stahlavsky, 2020*
C. vachoni Sisson, 1994
C. vannii Rossi, 2017*
C. werneri (Birula, 1908)
C. yagi Barahoei, 2025*
| Congobuthus Lourenço, 1999* (1)
C. fagei Lourenço, 1999* | Darchenia Vachon, 1977 (1)
D. bernadettae Vachon, 1977
| Egyptobuthus Lourenço, 1999* (1)
E. vaissadei Lourenço, 1999*
| Femtobuthus Lowe, 2010* (1)
F. shutuae Lowe, 2010* | Fetilinia Lowe & Kovarik, 2021* (1)
F. dentator Lowe & Kovarik, 2021*
| | Gint Kovarik, Lowe, Pliskova & Stahlavsky, 2013* (14)
G. abshiri Kovarik, Elmi & Stahlavsky, 2024*
G. amoudensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. banfasae Kovarik & Lowe, 2019*
G. calviceps (Pocock, 1900)
G. childsi Kovarik, 2018*
G. dabakalo Kovarik & Mazuch, 2015*
G. derbiae Kovarik, Elmi & Stahlavsky, 2024*
G. gaitako Kovarik, Lowe, Pliskova & Stahlavsky, 2013*
G. gubanensis Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018*
G. insolitus (Borelli, 1925)* (nomen dubium)
G. marialuisae Rossi, 2015* (nomen dubium)
G. monicae Rossi, 2015* (nomen dubium)
G. puntlandus Kovarik & Mazuch, 2015*
G. sahil Kovarik, Elmi & Stahlavsky, 2024*
| Grosphus Simon, 1880 (14)
G. ambre Lourenco, Wilme & Waeber, 2018*
G. angulatus Lowe & Kovarik, 2022*
G. darainensis Lourenço, Goodman & Ramilijaona, 2004*
G. goudoti Lourenço & Goodman, 2006*
G. halleuxi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. hirtus Kraepelin, 1900
G. madagascariensis (Gervais, 1843)
G. mandena Lourenço, 2005*
G. mayottensis Lourenço & Goodman, 2009*
G. polskyi Lourenço, Qi & Goodman, 2007*
G. rakotoariveloi Lourenço, Wilme, Soarimalala & Waeber, 2017*
G. simoni Lourenço, Goodman & Ramilijaona, 2004*
G. tavaratra Lourenco, Soarimalala & Goodman, 2009*
G. voahangyae Lourenço & Wilme 2015*
| Hemibuthus Pocock, 1900 (2)
H. crassimanus (Pocock, 1897)
H. umarii Amir, Kamaluddin & Khan, 2004*
| Hemilychas Hirst, 1911 (1)
H. alexandrinus (Hirst, 1911)
| Heteroctenus Pocock, 1893* (10)
H. abudi (Armas & Marcano Fondeur, 1987)*
H. aridicola (Teruel & Armas, 2012)*
H. bonettii (Armas 1999)*
H. garridoi (Armas, 1974)*
H. gibarae (Teruel, 2006)*
H. granulimanus (Teruel, 2006)*
H. junceus (Herbst, 1800)*
H. melloleitaoi (Teruel & Armas, 2006)*
H. princeps (Karsch, 1879)*
H. turieli Teruel & Yong, 2023*
| Hottentotta Birula, 1908 (62)
H. akbarii Yagmur, Moradi, Tabatabaei & Jafari, 2022*
H. alticola (Pocock, 1895)
H. arenaceus (Purcell, 1902)
H. asimii Amir, Kamaluddin & Khan, 2004*(nomen dubium)
H. buchariensis (Birula, 1897)*
H. caboverdensis Lourenço & Ythier, 2006*
H. conspersus (Thorell, 1876)
H. finneganae Kovarik, 2007*
H. flavidulus Teruel & Rein, 2010*
H. franzwerneri (Birula, 1914)
H. fuscitruncus (Caporiacco, 1936)
H. gambelaensis Kovarik, 2015*
H. gentili (Pallary, 1924)*
H. gibaensis Kovarik, 2015*
H. hatamtiorum Amiri, Prendini, Hussen, Aliabadian, Siahsarvie & Mirshamsi, 2024*
H. haudensis Kovarik & Lowe, 2021*
H. hoggarensis Lourenço & Leguin, 2014*
H. hottentotta (Fabricius, 1787)
H. jabalpurensis Kovarik, 2007*
H. jalalabadensis Kovarik, 2007*
H. jayakari (Pocock, 1895)
H. judaicus (Simon, 1872)
H. juliae Kovarik, Yagmur & Fet, 2019*
H. keralaensis Aswhati, Sureshan & Lourenço, 2016*
H. khoozestanus Navidpour, Kovarik, Soleglad & Fet, 2008*
H. krivokhatskyi Kovarik, Yagmur & Fet, 2019*
H. lacroixi Ythier & Dupre, 2021*
H. leetzi Ythier, 2024*
H. lorestanus Navidpour, Nayebzadeh, Soleglad, Fet, Kovarik & Kayedi, 2010*
H. mazuchi Kovarik, 2013*
H. mesopotamicus Lourenço & Qi, 2007*
H. minax (L. Koch, 1875)
H. minusalta Vachon, 1959*
H. navidpouri Kovarik, Yagmur & Moradi, 2018*
H. nigrimontanus Kovarik & Lowe, 2021*
H. niloticus (Birula, 1928)*
H. novaki Kovarik, 2015*
H. pachyurus (Pocock, 1897)*
H. pellucidus Lowe, 2010*
H. penjabensis (Birula, 1897)*
H. polystictus (Pocock, 1896)
H. pooyani Moradi, Yagmur, Akbari & Jafari, 2022*
H. reddyi Lourenço, 2015*
H. rugiscutis (Pocock, 1897)*
H. salei (Vachon, 1980)*
H. saulcyi (Simon, 1880)
H. saxinatans Lowe, 2010*
H. scaber (Ehrenberg, 1828)
H. schach (Birula, 1905)
H. sistanensis Kovarik, Yagmur & Moradi, 2018*
H. socotrensis (Pocock, 1889)
H. somalicus Kovarik, 2018*
H. songi (Lourenço, Qi & Zhu, 2005)*
H. sousai Turiel, 2014*
H. stockwelli Kovarik, 2007*
?H. syrticus (Borelli, 1914) (nomen dubium)
H. tamulus (Fabricius, 1798)*
H. trailini Kovarik, 2013*
H. trilineatus (Peters, 1861)
H. ugandaensis Kovarik, 2013*
H. vinchu Mirza, Ambekar & Kulkarni, 2019*
H. zagrosensis Kovarik, 1997
| Iranobuthus Kovarik, 1997 (1)
I. krali Kovarik, 1997
| Ischnotelson Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (2)
I. guanambiensis (Lenarducci, Pinto-da-Rocha & Lucas, 2005)*
I. peruassu Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
| Isometroides Keyserling, 1885 (1)
I. vescus (Karsch, 1880)
| Isometrus Ehrenberg, 1828 (18)
I. adviteeya Deshpande, Gowande, Dandekar, Joshi, Bastawade & Sulakhe, 2024*
I. amboli Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. anamalaiensis Deshpande, Gowande, Dandekar, Joshi, Bastawade & Sulakhe, 2024*
I. dnyandeoi Bandgar, Kininge, Bhosale, Bandgar & Bhosale, 2024*
I. formosus Pocock, 1894 (nomen dubium)
I. isadensis Tikader & Bastawade, 1983* (nomen dubium)
I. kovariki Sulakhe, Dandekar, Mukherjee, Pandey, Ketkar, Padhye & Bastawade, 2020*
I. lithophilis Deshpande, Gowande, Dandekar, Joshi, Bastawade & Sulakhe, 2024*
I. longitelson Deshpande, Gowande, Bastawade & Sulakhe, 2022*
I. maculatus (DeGeer, 1778)
I. nakshatra Sulakhe, Deshpande, Gowande, Dandekar & Ketkar, 2022*
I. palani Deshpande, Gowande, Dandekar, Joshi, Bastawade & Sulakhe, 2024*
I. sankeriensis Tikander & Bastawade, 1983
I. tamhini Sulakhe, Dandekar, Padhye & Bastawade, 2020*
I. thenmala Deshpande, Gowande, Dandekar, Joshi, Bastawade & Sulakhe, 2024*
I. thurstoni Pocock, 1893
I. thwaitesi Pocock, 1897
I. wayanadensis Sulakhe, Deshpande, Gowande, Dandekar & Ketkar, 2022*
| Jaguajir Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017 (3)
J. agamemnon (C. L. Koch, 1839)*
J. pintoi (Mello-Leitao, 1932)*
J. rochae (Borelli, 1910)*
| Janalychas Kovarik, 2019* (9)
J. albimanus (Henderson, 1919)
J. farkasi (Kovarik, 1997)
J. granulatus Mirza, 2020*
J. heurtaultae (Kovarik, 1997)
J. keralaensis Mirza, 2020*
J. laevifrons (Pocock, 1897)
J. shoplandi (Oates, 1888)
J. srilankensis (Lorenco, 1997)
J. tricarinatus (Simon, 1884)
| Karasbergia Hewitt, 1913 (1)
K. methueni Hewitt, 1913 | Kraepelinia Vachon, 1974 (1)
K. palpator (Birula, 1903) | Langxie Tang, Jia & Liu, 2023* (1)
L. feti Tang, Jia & Liu, 2023* | Lanzatus Kovarik, 2001* (3)
L. huluul Kovarik & Lowe, 2021*
L. somalicus Kovarik, 2001*
L. somalilandus Kovarik & Lowe, 2016* | Leiurus Ehrenberg, 1828 (25)
L. abdullahbayrami Yagmur, Koc & Kunt, 2009*
L. arabicus Lowe, Yagmur & Kovarik, 2014*
L. ater Lourenço, 2019*
L. aylaensis Abu Afifeh, Al-Saraireh & Amr, 2025*
L. brachycentrus (Ehrenberg, 1829)*
L. dekeyseri Lourenço, 2020*
L. gubanensis Kovarik & Lowe, 2020*
L. hadb Al-Qahtni, Al-Salem, Alqahtani & Badry, 2023*
L. haenggii Lowe, Yagmur & Kovarik, 2014*
L. heberti Lowe, Yagmur & Kovarik, 2014*
L. hebraeus (Birula, 1908)*
L. hoggarensis Lourenço, Kourim & Sadine, 2018*
L. jordanensis Lourenço, Modry & Amr, 2002*
L. kuwaiti Lourenço, 2020*
L. libycus (Birula, 1908)*
L. macroctenus Lowe, Yagmur & Kovarik, 2014*
L. maculatus Lourenço, 2022*
L. nigellus Abu Afifeh, Aloufi & Al-Saraireh, 2023*
L. nigerianus Lourenço, 2021*
L. quinquestriatus (Ehrenberg, 1828)
L. saharicus Lourenço, 2020*
L. sinai Badry, Saleh, Lourenço & Ythier, 2023*
L. savanicola Lourenço, Qi & Cloudsley Thompson, 2006*
L. somalicus Lourenço, & Rossi, 2016*
L. tamajeq Lourenço, & Ythier, 2024*
| Liobuthus Birula, 1898 (1)
L. kessleri Birula, 1898 | Lissothus Vachon, 1948 (3)
L. bernardi Vachon, 1948
L. chaambi Lourenço & Sadine, 2014*
L. occidentalis Vachon, 1950 | Lychas C.L. Koch, 1845 (33)
L. aberlenci Lourenço, 2013*
L. armasi Kovarik, 2013*
L. armillatus (Gervais, 1841)*
L. asper (Pocock, 1891)
L. biharensis Tikader & Bastawade, 1983
L. brehieri Lourenço, 2017*
L. buchari Kovarik, 1997
L. cernickai Kovarik, 2013*
L. chanthaburiensis Ythier & Lourenco, 2022*
L. flavimanus (Thorell, 1888)
L. gravelyi Henderson, 1913
L. hendersoni (Pocock, 1897)
L. hillyardi Kovarik, 1997
L. inexpectatus Lourenço, 2011*
L. jakli Kovarik, 2023*
L. kamshetensis Tikader & Bastawade, 1983
L. kharpadi Bastawade, 1986
L. krali Kovarik, 1995
L. lourencoi Kovarik, 1997
L. marmoreus (C.L. Koch, 1844)
L. mjobergi Kraepelin, 1916
L. mucronatus (Fabricius, 1798)
L. nigristernis (Pocock, 1899)
L. obsti Kraepelin, 1913
L. perfidus (Keyserling, 1885)
L. rackae Kovarik, 1997
L. rugosus (Pocock, 1897)
L. santoensis Lourenço, 2009*
L. scaber (Pocock, 1893)
L. scutilus C. L. Koch, 1845
L. serratus (Pocock, 1891)
L. shelfordi (Borelli, 1904)
L. variatus (Thorell, 1876)
| | Mauritanobuthus Qi & Lourenço, 2007* (1)
M. geniezi Qi &Lourenço, 2007*
| Mesobuthus Vachon, 1950 (31)
M. afghanus (Pocock, 1889)*
M. barszczewskii (Birula, 1904)
M. birulai Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. bogdoensis (Birula, 1896)*
M. crucittii Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. eupeus (C.L. Koch, 1839)
M. faiki Yagmur, Kovarik & Fet, 2024*
M. farleyi Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. fomichevi Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. galinae Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. haarlovi Vachon, 1958*
M. iranus (Birula, 1917)*
M. kaftani Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. kirmanensis (Birula, 1900)*
M. macmahoni (Pocock, 1900)
M. marusiki Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. mesopotamicus Penther, 1912*
M. mirshamsii Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. navidpouri Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. persicus (Pocock, 1899)*
M. philippovitschi (Birula, 1905)*
M. phillipsi (Pocock, 1889)*
M. rahsenae Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. rakhshanii Barahoei, 2023*
M. thersites (C. L. Koch, 1839)*
M. turcicus Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. vesiculatus (Pocock, 1899)
M. vignolii Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
M. yagmuri Kovarik & Fet, 2022*
M. zarudnyi Nouvruzov, Kovarik & Fet, 2022*
M. zonsteini Kovarik, Fet, Gantenbein, Graham, Yagmur, Stahlavsky, Poverenni & Nouvruzov, 2022*
| Mesotityus Gonzalez-Sponga, 1981 (1)
M. vondangeli Gonzalez-Sponga, 1981 | Microbuthus Kraepelin, 1898 (8)
M. fagei Vachon, 1949
M. flavorufus Lourenço & Duhem, 2007*
M. gardneri Lowe, 2010*
M. kristensenorum Lowe, 2010*
M. litoralis (Pavesi, 1885)
M. maroccanus Lourenço, 2002*
M. saharicus Lourenço, 2023*
M. satyrus Lowe, Kovarik, Stockmann & Stahlavsky, 2018*
| Microcharmus Lourenço, 1995 (17)
M. andrei Lourenço, Waeber & Wilme, 2019*
M. antongil Lourenço, Waeber & Wilme, 2019*
M. aridus Lourenço & Wilme, 2025*
M. bemaraha Lourenço, Goodman & Fisher, 2006*
M. cloudsleythompsoni Lourenço, 1995
M. confluenciatus Lourenço, Goodman & Fisher, 2006*
M. djangoa Lourenço, Waeber & Wilme, 2019*
M. duhemi Lourenço, Goodman & Fisher, 2006*
M. fisheri Lourenço, 1998
M. hauseri Lourenço, 1996
M. jussarae Lourenço, 1996*
M. maculatus Lourenço, Goodman & Fisher, 2006*
M. madagascariensis Lourenço, 1999*
M. pauliani (Lourenço, 2004)*
M. sabineae Lourenço, 1996
M. variegatus Lourenço, Goodman & Fisher, 2006*
M. violaceous Lourenço, Goodman & Fisher, 2006*
| Microtityus Kjellesvig-Waering, 1966 (41)
M. adriki Moreno-Gonzalez, Bertani & Carvalho, 2024*
M. angelaerrosae Gonzales-Sponga, 2001*
M. barahona Armas & Teruel, 2012*
M. biordi Gonzalez-Sponga, 1970
M. bivicentorum Botero-Trujillo, Erazo-Moreno & Perez, 2009*
M. borincanus Teruel, Rivera & Sanchez, 2014*
M. capayaensis Gonzalez-Sponga, 2001*
M. consuelo Armas & Marcano Fondeur, 1987
M. desuzeae Gonzalez-Sponga, 2001*
M. difficilis Teruel & Armas, 2006*
M. dominicanensis Santiago-Blay, 1985
M. eustatia Armas, 2018*
M. farleyi Teruel, 2000*
M. flavescens Teruel, 2001*
M. franckei Botero-Trujillo & Noriega, 2008*
M. fundorai Armas, 1974
M. guantanamo Armas, 1984
M. iviei Armas, 1999*
M. jaumei Armas, 1974
M. joseantonioi Gonzalez-Sponga, 1981
M. kovariki Teruel & Infante, 2007*
M. lantiguai Armas & Marcano Fondeur, 1992
M. litoralensis Gonzalez-Sponga, 2001*
M. lourencoi Armas & Teruel, 2012*
M. minimus Kovarik & Teruel, 2014*
M. paucidentatus Armas & Marcano Fondeur, 1992
M. prendinii Armas & Teruel, 2012*
M. pusillus Teruel & Kovarik, 2012*
M. reini Armas & Teruel, 2012*
M. rickyi Kjellesvig-Waering, 1966
M. santosi Teruel, Rivera & Sanchez, 2014*
M. sevciki Gonzalez-Sponga, 2001*
M. solegladi Armas & Teruel, 2012*
M. starri Lourenço & Huber, 1999*
M. trinitensis Armas, 1974
M. vanzolinii Lorenco & Eickstedt, 1983
M. vieques Teruel, Rivera & Santos, 2015*
M. virginiae Armas, 1999*
M. vulcanicus Teruel, 2019*
M. waeringi Francke & Sissom, 1980
M. yaracuyanus Gonzalez-Sponga, 2001*
| Neobuthus Hirst, 1911 (21)
N. amoudensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. awashensis Kovarik & Lowe, 2012*
N. berberensis Hirst, 1911
N. cloudsleythompsoni Lourenço, 2001*
N. dhobo Kovarik, Elmi & Stahlavsky, 2024*
N. erigavoensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. eritreaensis Lowe & Kovarik, 2016*
N. factorio Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. ferrugineus (Kraepelin, 1898)*
N. fryntai Kovarik, Elmi & Frydlova, 2023*
N. gubanensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. haeckeli Kovarik, 2019*
N. kloppersi Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. kutcheri Lowe & Kovarik, 2016*
N. maidensis Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. montanus Kovarik, Lowe, Awale, Elmi & Hurre, 2018*
N. opatovae Kovarik, Elmi & Stahlavsky, 2024*
N. osoli Kovarik, Elmi & Stahlavsky, 2024*
N. solegladi Kovarik, 2019*
N. sudanensis Lourenço, 2005*
N. verae Kovarik, Elmi & Stahlavsky, 2024*
| Neogrosphus Lourenço, 1995 (3)
N. andrafiabe Lourenço, Wilme & Waeber 2015*
N. blanci Lourenço, 1996
N. griveaudi (Vachon, 1969) | Neoprotobuthus Lourenço, 2000* (1)
N. intermedius Lourenço, 2000* | Odontobuthus Vachon, 1950 (12)
O. atherii (Amir & Kamaluddin, 2008)*
O. baluchicus Barahoei, Prendini, Navidpour, Tahir, Aliabadian & Siahsarvie, 2021*
O. bidentatus Lourenço & Pezier, 2002*
O. brevidigitus Lowe, 2010*
O. chabaharensis Barahoei, Prendini, Navidpour, Tahir, Aliabadian & Siahsarvie, 2021*
O. doriae (Thorell, 1876)
O. kermanus Barahoei, Prendini, Navidpour, Tahir, Aliabadian & Siahsarvie, 2021*
O. liaqatii (Amir & Kamaluddin, 2008)*
O. odonturus (Pocock, 1897)
O. persicus Barahoei & Shahi, 2024*
O. tavighiae Navidpour, Soleglad, Fet & Kovarik, 2013*
O. tirgari Mirshamsi, Azghadi, Navidpour, Aliabadian & Kovarik, 2013*
| Odonturus Karsch, 1879 (1)
O. dentatus Karsch, 1879 | Olivierus Farzanpay, 1987 (18)
O. brutus (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. caucasicus (Nordmann, 1840)
O. elenae (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. extremus (Werner, 1936)
O. fuscus (Birula, 1897)*
O. gorelovi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. hainanensis (Birula, 1904)
O. intermedius (Birula, 1897)*
O. kaznakovi (Birula, 1904)*
O. kreuzbergi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. longichelus (Sun & Zhu, 2010)*
O. martensii (Karsch, 1879)
O. mikhailovi Fet, Kovarik, Gantenbein & Graham, 2021*
O. mischi (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. nenilini (Fet, Kovarik, Gantenbein, Kaiser, Stewart & Graham, 2018)*
O. parthorum (Pocock, 1889)*
O. przewalskii (Birula, 1897)
O. voldemari Fet, Kovarik, Gantenbein & Graham, 2021*
| Orthochiroides Kovarik, 1998 (4)
O. insularis (Pocock, 1899)*
O. socotrensis Kovarik, 2004*
O. somalilandus Kovarik & Lowe, 2022*O. vachoni Kovarik, 1998
| Orthochirus Karsch, 1891 (58)
O. afar Kovarik & Lowe, 2016*O. afghanus Kovarik, 2004*
O. arabicus Ythier & Lourenço, 2023*
O. arenicola Lourenço & Ythier, 2021* [nomen dubium]
O. atarensis Lourenço & Leguin, 2011*
O. bastawadei Zambre, Mirza, Sanap, Upadhye & Javed, 2011*
O. bicolor (Pocock, 1897)
O. birulai Kovarik, Fet & Yagmur, 2020*
O. carinatus Navidpour, Kovarik, Soleglad & Fet, 2019*
O. cloudsleythompsoni Lourenço & Leguin, 2011*
O. danielleae (Lourenço & Vachon, 1997)*
O. farzanpayi (Vachon & Farzanpay, 1987)*
O. feti Kovarik, 2004*
O. flavescens (Pocock, 1897)
O. fomichevi Kovarik, Yagmur, Fet & Hussen, 2019*
O. formozovi Kovarik, Fet & Yagmur, 2020*
O. fuscipes (Pocock, 1900)
O. gantenbeini Kovarik, Yagmur, Fet & Hussen, 2019*
O. glabrifrons (Kraepelin, 1903)*
O. gromovi Kovarik, 2004*
O. grosseri Kovarik, Fet & Yagmur, 2020*
O. gruberi Kovarik & Fet, 2006*
O. hormozganensis Kovarik & Navidpour, 2020*
O. heratensis Kovarik, 2004*
O. innesi Simon, 1910
O. iranus Kovarik, 2004*
O. iraqus Kovarik, 2004*
O. jalalabadensis Kovarik, 2004*
O. katerinae Kovarik & Just, 2022*O. kermanensis Kovarik & Navidpour, 2020*
O. kovariki Yagmur & Khalili, 2022*
O. krishnai Tikader & Bastawade, 1983
O. kryzhanovskyi Kovarik, Fet & Yagmur, 2020*
O. kucerai Kovarik & Navidpour, 2020*
O. masihipouri Kovarik & Navidpour, 2020*
O. maroccanus Lourenço & Leguin, 2011*
O. melanurus (Kessler, 1874)*
O. mesopotamicus Birula, 1918*
O. milloti Lourenço, 2021*
O. minor Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. monodi (Lourenço & Vachon, 1997)*
O. navidpouri Kovarik, Yagmur, Fet & Hussen, 2019*
O. negebensis Shulov & Amitai, 1960
O. nordmanni Kovarik, Fet & Yagmur, 2020*
O. olivaceus (Karsch, 1881)
O. pallidus (Pocock, 1897)
O. persa (Birula, 1900)*
O. samrchelsis Kovarik, 2004*
O. scrobiculosus (Grube, 1873)
O. sejnai Kovarik, Fet & Yagmur, 2020*
O. semnanensis Kovarik & Navidpour, 2020*
O. soufiensis Lourenco & Sadine, 2021*
O. stockwelli (Lourenço & Vachon, 1995)*
O. tassili Lourenço & Leguin, 2011*
O. tibesti Lourenço, Duhem & Cloudsley-Thompson, 2012*
O. varius Kovarik, 2004*
O. vignolii Kovarik & Navidpour, 2020*
O. zagrosensis Kovarik, 2004*
| Pantobuthus Lourenço & Duhem, 2009* (1)
P. complicatus Lourenço & Duhem, 2009* | Parabuthus Pocock, 1890 (43)
P. abyssinicus Pocock, 1901*
P. brevimanus (Thorell, 1876)
P. calvus Purcell, 1898
P. capensis (Ehrenberg, 1831)
P. cimrmani Kovarik, 2004*
P. distridor Lamoral, 1980
P. dorisae Kovarik, Lowe, Elmi & Stahlavsky, 2024*
P. erigavoensis Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. eritreaensis Kovarik, 2003*
P. evae Kovarik, Lowe, Elmi & Stahlavsky, 2024*
P. glabrimanus Prendini & Esposito, 2010*
P. gracilis Lamoral, 1979
P. granimanus Pocock, 1895
P. granulatus (Ehrenberg, 1831)
P. hamar Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. heterurus Pocock, 1899
P. hunteri Pocock, 1895
P. kabateki Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. kajibu Kovarik, Lowe, Pliskova & Stahlavsky, 2016*
P. kalaharicus Lamoral, 1977
P. kraepelini Werner, 1902
P. kuanyamarum Monard, 1937
P. laevifrons (Simon, 1888)
P. liosoma (Ehrenberg, 1828)
P. maximus Werner, 1913
P. mazuchi Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. mossambicensis (Peters, 1861)
P. muelleri Prendini, 2000*
P. namibensis Lamoral, 1979
P. nanus Lamoral, 1979
P. pallidus Pocock, 1895
P. planicauda (Pocock, 1889)
P. puntlandus Kovarik, Stahlavsky, Elmi & Starha, 2025*
P. quincyae Kovarik, Lowe, Elmi & Stahlavsky, 2024*
P. raudus (Simon, 1888)
P. robustus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. schlechteri Purcell, 1899
P. setiventer Prendini & Esposito, 2010*
P. somalilandus Kovarik, Lowe, Elmi & Stahlavsky, 2019*
P. starhai Kovarik, 2025*
P. stridulus Hewitt, 1913
P. transvaalicus Purcell, 1899
P. villosus (Peters, 1862)
| Pectinibuthus Fet in Orlov & Vasilyev, 1984 (1)
P. birulai Fet in Orlov & Vasilyev, 1984 | Physoctonus Mello-Leitao, 1934* (3)
P. amazonicus Lourenço, 2017*
P. debilis (C. L. Koch, 1840)*
P. striatus Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
| Picobuthus Lowe, 2010* (2)
P. dundoni Lowe, 2010*
P. wahibaensis Lowe, 2010*
| Plesiobuthus Pocock, 1900 (1)
P. paradoxus Pocock, 1900 | Polisius Fet, Capes & Sissom, 2001* (1)
P. persicus Fet, Capes & Sissom, 2001* | | Pseudolissothus Lourenço, 2001* (1)
P. pusillus Lourenço, 2001*
| Pseudolychas Kraepelin, 1911 (3)
P. ochraceus (Hirst, 1911)
P. pegleri (Purcell, 1901)
P. transvaalicus Lawrence, 1961
| | Pseudouroplectes Lourenço, 1995 (6)
P. betschi Lourenço, 1995
P. jacki Lourenço, 2021*
P. lalyae Lourenço & Ythier, 2010*
P. maculatus Lourenço & Goodman, 2006*
P. pidgeoni Lourenço & Goodman, 1999*
P. tsingy Lourenço, Wilme & Waeber, 2016*
| Razianus Farzanpay, 1987 (4)
R. birulai Tahir, Navidpour & Prendini, 2014*
R. farzanpayi Tahir, Navidpour & Prendini, 2014*
R. xinjianganus Lourenco, Sun & Zhu, 2010*
R. zarudnyi (Birula, 1903) | Reddyanus Vachon, 1972* (37)
R. aareyensis (Mirza & Sanap, 2010)*
R. acanthurus (Pocock, 1899)*
R. alessandroi (Lourenco & Ythier, 2025)*
R. assamensis (Oates, 1888)*
R. basilicus (Karsch, 1879)*
R. besucheti (Vachon, 1982)*
R. bilyi (Kovarik, 2003)*
R. brachycentrus (Pocock, 1899)*
R. ceylonensis Kovarik, Lowe, Ranawana, Hoferek, Jayarathne, Pliskova & Stahlavsky, 2016*
R. corbeti (Tikander & Batawade, 1983)*
R. deharvengi (Lourenço & Duhem, 2010)*
R. feti (Kovarik, 2013)*
R. furai Kovarik & Stahlavsky, 2019*
R. hainanensis (Lourenco, 2005)*
R. heimi (Vachon, 1976)*
R. hofereki Kovarik & Stahlavsky, 2019*
R. jayarathnei Kovarik, 2016*
R. jendeki (Kovarik, 2013)*
R. justi Kovarik, Lowe & Stahlavsky, 2020*
R. khammamensis (Kovarik, 2003)*
R. krasenskyi (Kovarik, 1998)*
R. kurkai (Kovarik, 1997)*
R. lao (Louren�o & Leguin, 2012).*
R. loebli (Vachon, 1982)*
R. kanak (Lourenco, 2023)*
R. majkusi Kovarik & Stahlavsky, 2019*
R. melanodactylus (L. Koch, 1867)*
R. navaiae (Kovarik, 1998)*
R. neradi (Kovarik, 2013)*
R. petrzelkai (Kovarik, 2003)*
R. problematicus (Kovarik, 2003)*
R. ranawanai Kovarik, 2016*
R. rigidulus (Pocock, 1897)*
R. rolciki Kovarik & Stahlavsky, 2019*
R. schwotti Kovarik & Stahlavsky, 2019*
R. vittatus (Pocock, 1900)
R. zideki (Kovarik, 1994)*
| | Rhopalurus Thorell, 1876 (3)
R. caribensis Teruel & Roncallo, 2008*
R. laticauda Thorell, 1876
R. ochoai Esposito, Yamaguti, Souza, Pinto da Rocha & Prendini, 2017*
| | Sahil Kovarik, 2024* (1)
S. elmi Kovarik, 2024* | Saharobuthus Lourenço & Duhem, 2009* (1)
S. elegans Lourenço & Duhem, 2009* | Sanaag Kovarik, 2024* (1)
S. maidensis (Kovarik, Lowe, Just, Awale, Elmi & Stahlavsky, 2018)*
| | Sassanidotus Farzanpay, 1987 (2)
S. gracilis (Birula, 1900)*
S. zarudnyi (Birula, 1900) | Somalibuthus Kovarik, 1998 (2)
S. demisi Kovarik, 1998
S. sabae Kovarik & Njoroge, 2021*
| Somalicharmus Kovarik, 1998 (1)
S. whitmanae Kovarik, 1998 | Spelaeolychas Kovarik, 2019* (1)
S. hosei (Pocock, 1891)
| Teruelius Lowe & Kovarik, 2019* (22)
T. ankarafantsika (Lourenço, 2003)*
T. ankarana (Lourenço & Goodman, 2003)*
T. annulatus (Fage, 1929)*
T. bemaraha (Lourenco, Wilme & Waeber, 2018)*
T. bicolor (Lourenço, 2012)*
T. bistriatus (Kraepelin, 1900)*
T. eliseanneae (Lourenço & Wilme, 2016)*
T. feti (Lourenço, 1996)*
T. flavopiceus (Kraepelin, 1900)*
T. ganzhorni (Lourenço, Wilme & Waeber, 2016)*
T. grandidieri (Kraepelin, 1900)*
T. haeckeli Lowe & Kovarik, 2022*
T. intertidalis (Lourenço, 1999)*
T. limbatus (Pocock, 1889)*
T. magalieae (Lourenço, 2014)*
T. mahafaliensis (Lourenço, Goodman & Ramilijaona, 2004)*
T. makay (Lourenço & Wilme, 2015)*
T. mavo (Lourenço & Rossi 2020)*
T. olgae (Lourenço, 2004)*
T. rossii (Lourenço, 2013)*
T. sabineae (Lourenço & Wilme, 2016)*
T. waeberi (Lourenço & Wilme, 2016)*
| Thaicharmus Kovarik, 1995 (4)
T. guptai Mirza, Sanap & Kunte, 2016*
T. indicus Kovarik, 2013*
T. lowei Kovarik, Soleglad & Fet, 2007*
T. mahunkai Kovarik, 1995 | | Tityopsis Armas, 1974 (7)
T. canizaresorum Teruel & Rodriguez-Cabrera, 2020*
T. inaequalis (Armas, 1974)
T. inexpectata (Moreno, 1940)
T. mulata Teruel & Rodriguez-Cabrera, 2020*
T. pumila Teruel & Rodriguez-Cabrera, 2020*
T. rolandoi Kovarik, Stockmann, Stahlavsky & Yong, 2024*
T. sheylae Teruel & Rodriguez-Cabrera, 2020* | Tityus C. L. Koch, 1836 (230)
T. aba Candido, Lucas, de Souza, Diaz & Lira-da-Silva, 2005*
T. achilles Laborieux, 2024*
T. abudi Armas, 1999*
T. acananensis Gonzalez-Sponga, 2009*
T. adisi Lourenço & Pezier, 2002*
T. adrianoi Lourenço, 2003*
T. ahincoi Gonzalez-Sponga, 2001*
T. alejandroi Teruel, Rivera & Santos, 2015*
T. altithronus Armas, 1999*
T. androcottoides (Karsch, 1879)
T. anduzei Gonzalez-Sponga, 1997
T. angelesae Santiago-Blay, 2009*
T. anneae Lourenço, 1997
T. anori Lourenço, Rossi & Wilme, 2019*
T. antioquensis Lourenço& Otero Patino, 1998
T. apiacas Lourenço 2002*
T. arellanoparrai Gonzalez-Sponga, 1985
T. argentinus Borelli, 1899
T. asthenes Pocock, 1893
T. atriventer Pocock, 1897
T. bahiensis (Perty, 1833)
T. barquisimetanus Gonzalez-Sponga, 1994
T. bastosi Lourenço, 1984
T. bellulus Armas, 1999*
T. birabeni Abalos, 1955
T. blanci Lourenço, 1994
T. blaseri Mello-Leitao, 1931
T. boconoensis Gonzalez-Sponga, 1981
T. bolivianus Kraepelin, 1895
T. brazilae Lourenço & Eickstedt, 1984
T. breweri Gonzalez-Sponga, 1997
T. cachipalensis Gonzalez-Sponga, 2002*
T. caesarbarrioi Gonzalez-Sponga, 2001*
T. canopensis Lourenço & Pezier, 2002*
T. carabobensis Gonzalez-Sponga, 1987
T. carinatoides Mello-Leitao, 1945
T. caripitensis Quiroga, deSousa & Parrilla-Alvarez, 2000*
T. carolineae Kovarik, Teruel, Cozijn & Seiter, 2013
T. carrilloi Ojanguren-Affilastro, 2021*
T. carvalhoi Mello-Leitao, 1945
T. cerroazul Lourenço, 1986
T. championi Pocock, 1898*
T. charalaensis Mello-Leitao, 1940 [nomen nudum]
T. charreyroni Vellard, 1932
T. chilensis Lourenço, 2005*
T. choco Lourenço & Florez D, 2018*
T. cisandinus Lourenço & Ythier, 2017*
T. clathratus C. L. Koch, 1844
T. columbianus (Thorell, 1876)
T. confluens Borelli, 1899
T. costatus (Karsch, 1879)
T. crassicauda Lourenço & Ythier, 2013*
T. crassimanus (Thorell, 1876)
T. cuellari Lourenço, 1994
T. culebrensis Gonzalez-Sponga, 1994
T. curupi Ojanguren-Affilastro, Adilardi, Cajade, Ramirez, Ceccarelli & Mola, 2017*
T. cylindricus (Karsch, 1879)*
T. dasyurus Pocock, 1897
T. dedoslargos Francke & Stockwell, 1987
T. demangei Lourenço, 1981
T. dillerorum Kovarik, Teruel, Lowe & Friedrich, 2015*
T. dinizi Lourenço, 1997
T. discrepans (Karsch, 1879)
T. dorae Gonzalez-Sponga, 2001*
T. duacaensis Gonzalez-Sponga, 2007*
T. dulceae Gonzalez-Sponga, 2006*
T. dupouyi Gonzalez-Sponga, 1987
T. ecuadorensis Kraepelin, 1896
T. elii Armas & Marcano Fondeur, 1992*
T. elizabethae Lourenço & Ramos, 2004*
T. elizabethebravoi Gonzalez-Sponga & Wall Gonzalez, 2007*
T. engelkei Pocock, 1902
T. estherae Santiago-Blay, 2009*
T. evandroi Mello-Leitao, 1945
T. exstinctus Lourenço, 1995
T. fasciolatus Pessoa, 1935
T. festae Borelli, 1899
T. filodendron Gonzalez-Sponga, 1981
T. florezi Lourenço, 2000*
T. footei Chamberlin, 1916
T. forcipula (Gervais, 1843)
T. fuhrmanni Kraepelin, 1914
T. funestus Hirst, 1911
T. gaffini Lourenço, 2000*
T. gasci Lourenço, 1982
T. generaltheophiloi Lourenço, 2017*
T. gonzalespongai Quiroga, deSousa, Parrilla-Alvarez & Manzanilla 2004*
T. grahami Lourenço, 2012*
T. grottoedensis Botero-Trujillo & Florez D, 2014*
T. guane Moreno-Gonzalez, Gonzalez O & Florez D, 2019*
T. guaricoensis Gonzalez-Sponga, 2004*
T. haetianus Teruel & Santos, 2018*
T. horacioi Lourenço& Leguin, 2011*
T. icarus Laborieux, 2024*
T. imei Borges, de Sousa & Manzanilla, 2006*
T. indecisus Mello-Leitao, 1934
T. insignis (Pocock, 1889)
T. intermedius Borelli, 1899
T. isabelceciliae Gonzalez-Sponga, D'Suze & Sevcik, 2001*
T. ivani Gonzalez-Sponga, 2008*
T. ivicnancor Gonzalez-Sponga, 1997
T. jaimei Miranda, Bermudez, Florez D & de Armas, 2020*
T. jeanvellardi Lourenço, 2001*
T. julianae Lourenço, 2005*
T. juliorum Santiago-Blay, 2009*
T. jussarae Lourenço, 1988
T. kaderkai Kovarik, 2005*
T. kalettai Gonzalez-Sponga, 2007*
T. karaja Lourenço, 2016*
T. kindli Kovarik & Teruel, 2014*
T. kukututee Ythier, Chevalier & Gangadin, 2020*
T. kuryi Lourenço, 1997
T. lancinii Gonzalez-Sponga, 1972
T. lokiae Lourenço, Adis & Araujo, 2005*
T. longidigitus Gonzalez-Sponga, 2008*
T. lourencoi Florez, 1996
T. lutzi Giltay, 1928
T. macrochirus Pocock, 1897
T. magnimanus Pocock, 1897
T. maimirensis Gonzalez-Sponga, 2007*
T. mana Lourenço, 2012*
T. manakai Gonzalez-Sponga, 2004*
T. maniapurensis Gonzalez-Sponga, 2009*
T. marajoensis Lourenço & da Silva, 2007*
T. maranhensis Lourenço, de Jesus Junior & Limeira-de-Oliveira, 2006*
T. marechali Lourenço, 2013*
T. martinpaechi Lourenço, 2001*
T. matthieseni Pinto-da-Rocha & Lourenço, 2000
T. mattogrossensis Borelli, 1901
T. maturinensis Gonzalez-Sponga, 2008*
T. melanostictus Pocock, 1893
T. melici Lourenço, 2003*
T. metuendus Pocock, 1897
T. michelii Armas, 1982
T. microcystis Lutz & Mello, 1922
T. monaguensis Gonzalez-Sponga, 1974
T. mongei Lourenço, 1996
T. moralensis Moreno-Gonzalez, Pinto da Rocha & Cabra-Garcia, 2022*
T. mucusunamensis Gonzalez-Sponga, 2006*
T. munozi Lourenço, 1997
T. neblina Lourenço, 2008*
T. neglectus Mello-Leitao, 1932
T. neibae Armas, 1999*
T. nelsoni Lourenço 2005*
T. nematochirus Mello-Leitao, 1940
T. neoespartanus Gonzalez-Sponga, 1996
T. nororientalis Gonzalez-Sponga, 1996
T. obispoi Gonzalez-Sponga, 2006*
T. obscurus (Gervais, 1843)*
T. obtusus (Karsch, 1879)
T. ocelote Francke & Stockwell, 1987
T. osmanus Gonzalez-Sponga, 1996
T. oteroi Lourenço, 1998
T. ottenwalderi Armas, 1999*
T. pachyurus Pocock, 1897
T. pampanensis Gonzalez-Sponga, 2007*
T. panguana Kovarik, Teruel, Lowe & Friedrich, 2015*
T. paraguayensis Kraepelin, 1895
T. parvulus Kraepelin, 1914
T. paulistorum Lourenço & da Silva, 2006*
T. perijanensis Gonzalez-Sponga, 1994
T. pictus Pocock, 1893
T. pintodarochai Lourenço, 2005*
T. pittieri Gonzalez-Sponga, 1981
T. pococki Hirst, 1907
T. portoplatensis Armas & Marcano Fondeur, 1992
T. potameis Lourenço & Leao Giupponi, 2004*
T. prancei Lourenço, 2000*
T. proseni Abalos, 1954
T. pugilator Pocock, 1898
T. pusillus Pocock, 1893
T. quiriquirensis Gonzalez-Sponga, 2008*
T. quirogae De Sousa, Manzanilla & Parrilla-Alvarez, 2006*
T. quisqueyanus Armas, 1982*
T. ramirezi Esquivel de Verde, 1969
T. raquelae Lourenço, 1988
T. rebieri Lourenço, 1997
T. riocaurensis Gonzalez-Sponga, 1996
T. rionegrensis Lourenço, 2006*
T. riverai Teruel & Sanchez, 2009*
T. roigi Maury & Lourenço, 1987
T. rojasi Gonzalez-Sponga, 1996
T. romeroi Gonzalez-Sponga, 2008*
T. rondonorum Rojas-Runjaic & Armas, 2007*
T. rosenbergi Pocock, 1898*
T. rufofuscus Pocock, 1897
T. rupestre Lourenço 2019*
T. rugosus Schenkel, 1932
T. rusmelyae Gonzalez-Sponga, D'Suze & Sevcik, 2001*
T. sabineae Lourenço, 1994
T. sanarensis Gonzalez-Sponga, 1997
T. sanfernandoi Gonzalez-Sponga, 2008*
T. sarisarinamensis Gonzalez-Sponga, 2002*
T. sastrei Lourenço & Florez, 1990
T. schrammi Teruel & Santos, 2018*
T. serrulatus Lutz & Mello, 1922
T. shiriana Gonzalez-Sponga, 1991
T. silvestris Pocock, 1897
T. simonsi Pocock, 1900
T. smithii Pocock, 1893
T. soratensis Kraepelin, 1911
T. spelaeus Moreno-Gonzalez, Pinto-da-Rocha & Gallao, 2021*
T. stigmurus (Thorell, 1876)
T. strandi Werner, 1939
T. surmeridensis Gonzalez-Sponga, 2002*
T. surorientalis Gonzalez-Sponga, 1996
T. sylviae Lourenço 2005*
T. tamayoi Gonzalez-Sponga, 1987
T. tayrona Lourenço, 1991
T. tenuicauda Prendini, 2001*
T. timendus Pocock, 1898
T. tocantim Lourenço, 2022
T. trinitatis Pocock, 1897
T. trivittatus Kraepelin, 1898
T. tucurui Lourenço, 1988
T. uniformis Mello-Leitao, 1931
T. unus Pinto-da-Rocha & Lourenço, 2000
T. uquirensis Gonzalez-Sponga, 2001*
T. urachichensis Gonzalez-Sponga, 2007*
T. urbinai Scorza, 1952
T. uruguayensis Borelli, 1901
T. vaissadei Lourenço, 2002*
T. valerae Scorza, 1954
T. venamensis Gonzalez-Sponga, 1981
T. ventuarensis Gonzalez-Sponga, 2009*
T. wachteli Kovarik, Teruel, Lowe & Friedrich, 2015*
T. walli Gonzalez-Sponga & Wall Gonzalez, 2007*
T. yerenai Gonzalez-Sponga, 2009*
T. ythieri Lourenço, 2007*
T. zulianus Gonzalez-Sponga, 1981
| | Troglorhopalurus Lourenço, Baptista & Giupponi, 2004* (3)
T. iuiu Carvalho, Silva, Emanuela de Souza & Feirreira, 2025*
T. lacrau (Lourenço & Pinto-da-Rocha, 1997)*
T. translucidus Lourenço, Baptista & Giupponi, 2004*
| Trypanothacus Lowe, Kovarik, Stockmann & Stahlavsky, 2019* (3)
T. azraqensis Al-Saraireh, Afifeh, Aloufi, Amr & Lourenco, 2021*
T. barnesi Lowe, Kovarik, Stockmann & Stahlavsky, 2019*
T. buettikeri (Hendrixson, 2006)*
| | Uroplectes Peters, 1861 (35)
U. ansiedippenaarae Prendini, 2015*
U. carinatus (Pocock, 1890)
U. chubbi Hirst, 1911
U. ebogo Kovarik, Stahlavsky & Govorov, 2024*
U. emiliae (Werner, 1916)
U. fischeri (Karsch, 1879)
U. flavoviridis Peters, 1861
U. formosus Pocock, 1890
U. gracilior Hewitt, 1913
U. insignis Pocock, 1890
U. katangensis Prendini, 2015*
U. lineatus (C. L. Koch, 1844)
U. longimanus Werner, 1936
U. machadoi Lourenço, 2000*
U. malawicus Prendini, 2015*
U. marlothi Purcell, 1901
U. ngangelarum Monard, 1930
U. occidentalis Simon, 1876
U. olivaceus Pocock, 1896
U. otjimbinguensis (Karsch, 1879)
U. pardalis Werner, 1913
U. pardii Kovarik, 2003*
U. pictus Werner, 1913
U. pilosus (Thorell, 1876)
U. planimanus (Karsch, 1879)
U. schlechteri Purcell, 1901
U. schubotzi Kraepelin, 1929
U. silvestrii Borelli, 1913
U. teretipes Lawrence, 1966
U. triangulifer (Thorell, 1876)
U. tumidimanus Lamoral, 1979
U. variegatus (C. L. Koch, 1844)
U. vittatus (Thorell, 1876)
U. xanthogrammus Pocock, 1897
U. zambezicus Prendini, 2015*
| | Vachoniolus Levy, Amitai & Shulov, 1973 (4)
V. batinahensis Lowe, 2010*
V. gallagheri Lowe, 2010*
V. globimanus Levy, Amitai & Shulov, 1973
V. iranus Navidpour, Kovarik, Soleglad & Fet, 2008*
| Vachonus Tikader & Bastawade, 1983 (4)
V. asiyaae Amir & Kamaluddin, 2009* [nomen dubium]
V. inexpectatus Lourenço, 2015*
V. iqbali Amir & Kamaluddin, 2009* [nomen dubium]
V. rajasthanicus Tikader & Bastawade, 1983
| Xenobuthus Lowe, 2018* (3)
X. anthracinus (Pocock, 1895)*
X. arabicus (Lourenço & Qi, 2006)*
X. xanthus Lowe, 2018*
| Zabius Thorell, 1893 (3)
Z. birabeni Mello-Laeitao, 1938
Z. fuscus (Thorell, 1876)
Z. gaucho Acosta, Candido, Buckup & Brescovit, 2008*
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