Stipules (Botany)

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In botany, a stipule is an outgrowth typically borne on both sides (sometimes on just one side) of the base of a leafstalk (the petiole). They are primarily found among dicots and rare among monocots. Stipules are considered part of the anatomy of the leaf of a typical flowering plant, although in many species they may be inconspicuous —or sometimes entirely absent, and the leaf is then termed exstipulate. At the other end of the scale are species like Artocarpus elasticus where the stipules can be up to eight inches (twenty cm) in length. (In some older botanical writing, the term "stipule" was used more generally to refer to any small leaves or leaf-parts, notably prophylls.) The word stipule was coined by Linnaeus from Latin stipula, straw, stalk.