Tambach Formation
Stratigraphic range:Early Permian,Sakmarian PreꞒ Ꞓ O S D C P T J K Pg N ↓
Traco quarry at Bromacker
Type	Geological formation
Underlies	Eisenach Formation
Overlies	Rotterode Formation
Lithology
Primary	Conglomerate,sandstone,mudstone
Location
Coordinates	50.8097° N, 10.6189° E
Region	Thuringia
Country	Germany
Type section
Named for	Tambach-Dietharz village

TheTambach Formation is anEarly Permian-agegeologic formation in centralGermany. It consists of red to brown-coloredsedimentary rocks (red beds) such asconglomerate,sandstone, andmudstone, and is the oldest portion of theUpper Rotliegend within theThuringian Forest Basin.^[1][2]

The overall geology records a history with three distinct stages of sedimentation within a mountainous environment. First, tectonic activity forms a basin (the Tambach Basin) dominated by high-energydebris flows,sheetfloods, andbraided rivers. These incise underlyingrhyolitic rock, depositing a coarseconglomerate known as the Bielstein Conglomerate. Second, calmer conditions allow the basin to widen, and the conglomerate is marginalized by finer sediments which were previously only common at the center of the basin, such as the characteristic Tambach Sandstone. These finer sediments were deposited through repeated sequences of flooding, followed by calm water, followed by exposure to air. The overall climate would have been similar to moderntropical savannas, with hot and dry periods broken up by heavy rainfall, likely multiple times in a year. The third stage involves a return of tectonic conditions, this time inducing wide deposits of polymictic (heterogenous) conglomerate known as the Finsterbergen conglomerate.^[1][3][2]

The Tambach Formation also includes one of the most important Permian fossil localities in Europe: the Bromacker locality. This former sandstone quarry and surrounding sites preserves several differentfacies types, with different fossil components. Thick sandstone channel fills in the lower section at Bromacker are overlain with mudstone fromephemeral lakes. A diverse assemblage of trace fossils such as footprints are imprinted onto the mudstone drapings. The upper section of Bromacker containssiltstone deposited through sheetfloods, in which well-preserved articulated skeletons of terrestrialtetrapods have been discovered. These include early amphibians likeRotaryus andTambaroter, and early reptiles likeEudibamus andThuringothyris. Unlike most fossil-preserving Permian red beds, aquaticvertebrates are absent at Bromacker while carnivoroussynapsids (likeDimetrodon) are rare and herbivorousdiadectids are abundant. Plant and arthropod fossils have also been found in shales at Bromacker.^[1][4]

Starting in the 1840s, five-toed footprints had been described from various sandstone quarries and roadcuts in theGotha district of Germany. The stratigraphy of rocks near Tambach-Dietharz was mapped out in 1876,^[5] and named as the "Tambacher Schichten" (Tambach strata) in 1895, although at the time it was believed to include several additional rock layers (now termed theElgersberg andEisenach Formations) which have since then been separated from it.^[2] in 1887 the first fossilized footprints from Bromacker were discovered by a local fossil collector namedHeinrich Friedrich Schäfer. The find was donated to the Ducal Museum in the city ofGotha, after which it was independently reported on by various German paleontologists who had received photographic evidence.^[6]Wilhelm Pabst, the curator of the Ducal Museum's natural history department, collected and described 140 sandstone slabs from the Tambach Formation from 1890 until his death in 1908. The collection was rediscovered in the 1950s, and subsequently restudied byGDR paleontologists such asHermann Schmidt,Arno Hermann Müller,^[7] andHartmut Haubold.^[8]

Fossilized tetrapod bones were discovered in the upper beds of Bromacker byThomas Martens in 1974, prompting further attention from Gotha paleontologists. These includeHarald Lutzner, who formally delineated the Tambach Formation as a sequence including two conglomerate layers separated by a sandstone layer.^[2] In the coming years, research contacts were made with Western paleontologists such asJürgen Boy (University of Mainz) andDavid Berman (Carnegie Museum of Natural History). This allowed excavation to ramp up and the Tambach Formation to achieve worldwide fame. A 1993 German-American joint expedition recovered articulated tetrapod fossils, and trace fossil collecting was resumed after more than 80 years thanks to the excavation of a new sandstone quarry at Bromacker in 1995. The first Tambach body fossils outside of Bromacker were discovered in 2008, in a construction site in downtown Tambach-Dietharz. In 2010, the exhibits comprising Gotha's Museum of Nature began the process of being moved from the Ducal Museum (which was being converted to an art museum) to theFriedenstein Castle. Due to funding issues, collecting from Bromacker has been limited and the Tambach collection is being archived at the historic Perthesforum complex prior to the construction of a new Permian exhibit.^[8]

The Tambach Formation mainly lies within a basin currently occupied by theThuringian Forest, and it is stratigraphically younger than theRotterode Formation and older than theEisenach Formation. It is the oldest part of the Thuringian Forest Basin's UpperRotliegend succession, a name referring to a sequence of purely sedimentary rocks in the Lower Permian of Germany. The sediments of the Tambach Formation were deposited in a small Permiangraben (termed the Tambach Basin), which was oriented in a northeast to southwest direction and incised into the igneous and sedimentary rocks of the Rotterode Formation. The Tambach Basin would have about 250 square km during the Permian, though modern outcrops only occupy about 50 square km, not counting the northeastern portion of the basin which has had its deposits erased by later geological processes.^[1][2]

Traditionally, the Tambach Formation is considered to be divided into three discrete layers: a lower and an upperconglomerate layer (Bielstein and Finsterbergen Konglomerate, respectively), separated by a somewhat narrower layer of finer sediments such assandstone, the Tambach-Sandstein (Tambach sandstone) member.^[9][3][2] However, the borders between these layers are often imprecise, and some geologists have offered a more complex depositional picture with three stratigraphic stages defined by changes in basin-wide sedimentology, rather than specific rock types.^[1]

The lowest stage (stage I) experienced a period of high tectonic activity (part of theVariscan orogeny) to the southeast, the Oberhof uplift. The formation of the basin among this tectonic backdrop initially led to powerfuldebris flows andsheet floods, and then activebraided rivers flowing along its edge with lower-energy rivers,floodplains, and lakes at its center. The coarse (cobble/boulder-scale) andrhyolitic Bielstein conglomerate was deposited among the high-energy marginal environments while early portions of the Tambach sandstone were deposited further away from the basin's edge. The most prominent flow direction in the well-preserved eastern part of the basin is northwest, towards the center of the basin.^[1][2][10]

Decreasing tectonic activity in the middle stage (stage II) leads to increasederosion, lowering the relief along the edge of the basin. As a result, the braided rivers at the edge slowed down, lowering the size of the clasts to cobble/pebble conglomerate. The center of the basin graded into small streams and marshes, depositing sandstone,siltstone,shale and mudstone in fossiliferousred beds. Although the Tambach Basin may have been hydrologically isolated during this period, with its waterways draining internally,^[1] some paleontologists instead consider its waters to flow into another basin in the northeast, which was not preserved.^[2] The last stage (stage III) experienced a return of tectonic activity (the Ruhla crystalline uplift) to the northwest, although the relief was still fair flat. Ruhla-sourcedalluvial fans and braidplains became more common, gradually allowing mineral-rich and polymictic pebble-sizedconglomerate to build up and expand into the center of the basin, forming the Finsterbergen conglomerate.^[1][2][10]

The most famous and fossiliferous locality within the Tambach Formation is the Bromacker locality, a cluster of small abandoned quarries near the town ofTambach-Dietharz. Strata exposed to the surface at Bromacker corresponds to the center of the Tambach Basin, during the time of the upper portion of stage I and the lower portion of stage II. Stage I sediments at Bromacker are termed the "lower beds"^[1] or "Bromacker sandstone"^[3] and stage II sediments are the "upper beds"^[1] or "Bromacker horizon".^[3]

The lower beds are dominated by thick sheets of fine-grained sandstone, often withcross-bedding indicating that thepaleocurrent was oriented towards the northeast. These sandstone layers are typically blanketed by homogeneousmudstone, which sometimes preservedmudcracks, plant fossils,invertebrate burrows, andtetrapod footprints. The thick sandstone layers (and their mudstone drapings) are not continuous, interrupted by a succession of finer and darkermicaceous sediments such asshale,siltstone, and (rarely) very fine sandstone. This reconstructs a sequence of repeated flooding events, involving strong, straight rivers eroding channels through the siltyfloodplain at the center of the basin, leaving behind sandy channel fills (sandstone) and fine-grainedoverbank deposits (shale and other sediments). The mudstone drapings can be explained as precipitate from extensiveephemeral lakes that evaporated in the weeks following the floods, after which they became mudflats. Many of the mudcrack fragments were ripped up by the next flood, being incorporated into the subsequent sandstone sheets asintraclasts.^[1]

The narrower upper beds also represent alternating flooding and still water conditions, albeit with finerclasts and lower topography. The sandstone channel fills are mostly replaced by homogeneous layers of red siltstone, which were prone to breaking along sharp edges. This siltstone contained mudcrack fragments,calcite-encased roots, and well-preserved partial or articulated skeletons of terrestrial tetrapods. These sediments indicate that reduced relief in stage II of the Tambach Formation had madesheetfloods the dominant erosional force, rather than individual channels. The lowest sheetflood deposit was particularly rich indiadectid fossils. The fine sediment successions were also altered, becoming dominated by finely laminated (and only occasionally micaceous) shale in whichconchostracans andarthropod remains were fossilized. These indicate a transition to more permanent lakes and broad floodplain conditions at the center of the Tambach Basin, rather than the ephemeral fluvial environment of the lower beds.^[1]

An extensive older sequence, the "Tambach-Wechsellagerung" (Tambachinterbedding) was discovered in 2004 throughborehole data. This sequence somewhat resembled the lower beds, with alternating fine micaceous deposits and thick sandstone sheets filled with abreccia of mudstone intraclasts. However, the sandstone layers had no evidence of cross-bedding, and the mud drapings responsible for most of the Tambach trace fossils were absent as well. Rare fragments of vertebrate fossils were present, along with calcite structures.^[3]

Uranium-Lead dating is not possible for the Tambach Formation, which lacks fresh volcanic rocks. The similar^[2]Elgersburg Formation to the southeast containsrhyolite dated to 274 ± 4.9 million years ago.^[11] However, it is unclear whether the strata atElgersburg are younger, older, or equivalent in age to the Tambach Formation.^[3]Biostratigraphy is more informative but still imprecise. Insect andconchostracan biostratigraphy places it into theSakmarian-ArtinskianMoravamylacris kukalovae^[12] and late ArtinskianLioestheria monticula/andreevi^[13] biozones, respectively. The only species of tetrapod known to exist in both the Tambach Formation and North American faunas isSeymouria sanjuanensis, which persisted for approximately 15 million years between theAsselian and the earlyKungurian. Since the species ofDimetrodon present at Tambach is smaller than those present in thered beds of Texas, the Tambach Formation was likely older than those formations.^[14] The Tambach Formation was placed within theSeymouran LVF (Land Vertebrate Faunachron) of Lucas (2006), a biozone which was estimated to include the Artinskian-Kungurian boundary.^[15] Combining both invertebrate and tetrapod biostratigraphy, the age of the Tambach Formation was considered to be probably Artinskian in age.^[12] In a study published in 2022, Menning and colleagues consider the age of the Tambach formation to be probably between 294 and 292 Ma, corresponding to the Sakmarian.^[16] This estimate is based primarily on the radiometric age of 295.8 ± 0.4 Ma (lateAsselian) of the Rotterode Formation whichunconformably underlies the Tambach Formation, and on the estimate that the interval of geologic time not represented between the two formations is less than 2 million years.^[16][17] In addition, comparison of the footprint assemblage of the Tambach Formation with radiometrically dated Permian footprint assemblages fromFrance andItaly also suggests a Sakmarian age.^[16][18]

The sand and silt-rich portions of the Tambach Formation were likely deposited in a warm climate with both hot, dry parts of the year and periodic heavy rainfall events. The dry times were severe enough to evaporate the Tambach basin's flood-induced ephemeral lakes within a matter of days, restricting the ability of a permanent aquatic fauna to colonize the basin. However, most plant root fossils are horizontally (rather than vertically) oriented, indicating that the climate was generally humid enough that native plants would not need to evolve deep roots or otherxerophytic adaptations. Because of this, the Tambach Formation would probably fall under the moderntropical savanna climate, despite its lack of grass. Modern climatic equivalents include the northern African savanna and theLlanos ofVenezuela andColombia.^[1] However, from the sedimentgeochemistry of the Tambach Sandstone Member, Scholze and Pint proposed a mean annual temperature of only 10.9 to 15°C (12.7°C on average).^[19] There is evidence that sub-zero temperatures may have occurred on some nights during the dry season, likely as a result of the basin's high elevation.^[20][2][19] The climate may have been drier during the conglomerate-rich periods of the Tambach Formation.^[1][2]

The ecosystem of the Tambach Formation is unusual for its lack of aquatic animals such asxenacanthid sharks,Eryops, orDiplocaulus, which are otherwise common in Early Permian red beds. This is best explained by its mountainous environment, isolated from the monsoonal lowland floodplains which deposited most of the red beds. In addition, the ephemeral nature of the Tambach Basin's lakes and rivers means that only aquatic animals adapted to such conditions, such asconchostracans, were able to flourish. The Tambach Basin did support a diverseamphibian fauna, but only terrestrially-adapted types includingdissorophoids andseymouriamorphs. Large herbivorous tetrapods such ascaseids and especiallydiadectids are the most common body fossils recovered from the formation, while carnivorous synapsids are relatively rare. This is in contrast to North American environments, where fossils of carnivores such asDimetrodon outnumber herbivore fossils. The environmental conditions of Tambach likely created a food web which was very different from that of the lowlands. The most common plants were tough, drought-adapted types such asconifers, whileseed ferns and other lowland plants were much rarer. Fibrous terrestrial plants encouraged colonization of the basin by herbivorous land animals, but the dry climate prevents the development of an aquatic food chain, inhibiting animals such as large species ofDimetrodon, which get a large portion of their food from waterways.^[1][4]

• Permian Germany
• Permian System of Europe
• Sakmarian
• Paleontology in Germany

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