Nemonychidae Temporal range:Oxfordian-Present
Doydirhynchus austriacus
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Order:	Coleoptera
Suborder:	Polyphaga
Infraorder:	Cucujiformia
Clade:	Phytophaga
Superfamily:	Curculionoidea
Family:	Nemonychidae Bedel, 1882
Subfamilies
Cimberidinae (but see text) Nemonychinae Rhinorhynchinae
Synonyms
CimberididaeGozis, 1882[1] CimberidaeAuctt. (missp.)

Nemonychidae is a small family ofweevils, placed within theprimitive weevil group because they have straight rather than geniculate (elbowed)antennae. They are often calledpine flower weevils. As in theAnthribidae, thelabrum appears as a separate segment to theclypeus, and the maxillary palps are long and projecting. Nemonychidae have all ventrites free, while Anthribidae have ventrites 1-4 connate or partially fused. Nemonychidae lack lateral carinae on the pronotum, while these are usually present, though may be short, in Anthribidae.^[2]

Nemonychidae are historically divided into three subfamilies: Nemonychinae of the palearctic region with the single genusNemonyx and an unusual host, the angiospermDelphinium. Most species of the other two subfamilies are associated withPinales feeding on the pollen of the maleinflorescences.Cimberidinae are found in the Northern hemisphere, whileRhinorhynchinae^[3] occur largely in the Southern hemisphere, especially found onPodocarpaceae andAraucariaceae. Recent phylogenetic research indicates that the Cimberidinae are sister to all of the remainingCurculionoidea, and it has been proposed to elevate the group to family rank.^[4]

Both Nemonychidae adults and larvae feed primarily on pollen.^[5] When mature, the larvae will drop to the ground from the male cones or flowers they resided in to pupate in the soil less than five centimeters from the surface.^[6]

There exists a fairly extensive fossil record of Nemonychidae reaching from the upper Jurassic to tertiaryamber.

• †Kuschelomacer^[7]
• †subfamily Brenthorrhininae Arnoldi 1977
  • †tribe Brenthorrhinini Arnoldi 1977
    • †subtribe Brenthorrhinina Arnoldi 1977
      • †Abrenthorrhinus Legalov 2009Karabastau Formation, Kazakhstan,Callovian/Oxfordian
      • †Brenthorrhinus Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
      • †Gobibrenthorrhinus Gratshev and Legalov 2009Ulugei Formation, Mongolia,Tithonian
    • †subtribe Brenthorrhinoidina Legalov 2003
      • †Brenthorrhinoides Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
  • †tribe Distenorrhinini Arnoldi 1977
    • †Buryatnemonyx Legalov 2010 Khasurty locality, Russia,Aptian
    • †Distenorrhinus Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian,La Pedrera de Rúbies Formation,Las Hoyas, Spain,Barremian Khasurty locality, Russia, Aptian
    • †Megabrenthorrhinus Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Microbrenthorrhinus Gratshev and Zherikhin 2000 La Pedrera de Rúbies Formation, Spain, Barremian
    • †Talbragarus Oberprieler and Oberprieler 2012Talbragar Fossil bed, Australia, Tithonian
  • †tribe Eccoptarthrini Arnoldi 1977
    • †Astenorrhinus Gratshev and Zherikhin 1995 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Cratonemonyx Legalov 2014Crato Formation, Brazil, Aptian
    • †Eccoptarthrus Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Procurculio Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
• †subfamily Cretonemonychinae Gratshev and Legalov 2009
  • †tribe Cretonemonychini Gratshev and Legalov 2009
    • †Cretonemonyx Gratshev and Legalov 2009Zaza Formation, Russia, Aptian
    • †Pseudonemonyx Gratshev and Legalov 2009 Zaza Formation, Russia, Aptian
    • †Turononemonyx Legalov 2014 Kzyl-Zhar, Kazakhstan,Turonian
  • †tribe Eocaenonemonychini Legalov 2013
    • †Eocaenonemonyx Legalov 2013Green River Formation, United States,Eocene
    • †"Eugnamptus" decemsatus Scudder 1878 Green River Formation, United States, Eocene
    • †"Sitona" grandaevus Scudder 1876 Green River Formation, United States, Eocene
• †subfamilyEobelinae Arnoldi 1977
  • †tribe Eobelini Arnoldi 1977
    • †Archaeorrhynchoides Legalov 2009 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Archaeorrhynchus Martynov 1926 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Eobelus Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Martynovirhynchus Legalov 2010 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
  • †tribe Karataucarini Legalov 2009
    • †Ampliceps Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
  • †tribe Oxycorynoidini Arnoldi 1977
    • †Oxycorynoides Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/OxfordianWeald Clay, United Kingdom, BarremianGurvan-Eren Formation, Mongolia, Aptian
    • †Microprobelus Liu et al. 2006Yixian Formation, China, Aptian
    • †Khetoxycorynoides Legalov 2011Emanra Formation, Russia, Turonian
    • †Cratomacer Zherikhin and Gratshev 2004 Crato Formation, Brazil, Aptian
  • †tribe Probelini Legalov 2009
    • †Belonotaris Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
    • †Probelus Arnoldi 1977 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
  • Incertae sedis
    • †Longidorsum Zhang 1997Dalazi Formation, China, Aptian
• †subfamily Paleocartinae Legalov 2003
  • †tribe Metrioxenoidini Legalov 2009
    • †Brasilnemonyx Legalov 2009 Crato Formation, Brazil, Aptian
    • †Cretoxenoides Legalov 2010Huhtyk Formation, Mongolia,Hauterivian
    • †Khetanamonyx Legalov 2009 Emanra Formation, Russia, Turonian
    • †Megametrioxenoides Gratshev and Legalov 2009Tsagaantsav Formation, Mongolia,Valanginian
    • †Metrioxenoides Gratshev et al. 1998Lulworth Formation, United Kingdom,Berriasian
  • †tribe Paleocartini Legalov 2003
    • †subtribe Nebrenthorrhinina Legalov 2007
      • †Nebrenthorrhinus Legalov 2003 La Pedrera de Rúbies Formation, Spain, Barremian
    • †subtribe Paleocartina Legalov 2003
      • †Paleocartus Legalov 2003 Karabastau Formation, Kazakhstan, Callovian/Oxfordian
  • †tribe Selengarhynchini Gratshev and Legalov 2009
    • †Selengarhynchoides Legalov 2010Sharin-Gol Formation, Mongolia, Barremian
    • †Selengarhynchus Gratshev and Legalov 2009 Sharin-Gol Formation, Mongolia, Barremian
• Subfamily Rhinorhynchinae Voss 1922
  • Tribe Mecomacerini Kuschel 1994
    • †Burmomacer Legalov 2018Burmese amber, Myanmar,Cenomanian
    • †Burmonyx Davis and Engel 2014 Burmese amber, Myanmar, Cenomanian
    • †Guillermorhinus Clarke and Oberprieler 2018 Burmese amber, Myanmar, Cenomanian
    • †Renicimberis Legalov 2009 Yixian Formation, China, Aptian

Wikispecies has information related toNemonychidae.

• Images of Nemonychidae species in New Zealand

• Weevils
• Polyphaga families
• Nemonychidae

• Articles with short description
• Short description is different from Wikidata
• Articles with 'species' microformats