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Gilmoremys

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Extinct genus of turtles

Gilmoremys Temporal range:Upper Cretaceous,Maastrichtian PreꞒ Ꞓ O S D C P T J K Pg N
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	Trionychidae
Subfamily:	†Plastomeninae
Genus:	†Gilmoremys Joyce & Lyson,2011
Species
†G. lancensis (Gilmore, 1928) [originallyAspideretes lancensis] (type) †G. gettyspherensis Joyce, Lyson, and Sertich, 2018

Gilmoremys is anextinct genus ofsoftshell turtle which lived during the lateCretaceous (Maastrichtian stage) ofNorth Dakota,Montana andWyoming, United States.

Remains

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Gilmoremys is known from fiveskulls, amandible and an incomplete postcranialskeleton. Theholotype ofG. lancensis,USNM 6727, consists of a nearly completecarapace and an isolated hyoplastral fragment, was first assigned to the speciesAspideretes lancensis. Many additional specimens were later discovered includingcranial remains, and the material was assigned to its own genus,Gilmoremys. It was found from theHell Creek Formation and from theLance Formation (only the holotype). It was first named by Walter G. Joyce and Tyler R. Lyson in2011 and thetype species isGilmoremys lancensis. The generic name honors Dr.Charles W. Gilmore.^[1] A second species,G. gettyspherensis, is known from the late CampanianFruitland Formation of New Mexico.^[2]

Morphology

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Juveniles of this species have narrow skulls, a narrow processus trochlearis oticum, a deep and narrow median palatal groove, low accessory ridges, and a secondary palate fully formed by the maxilla, skeletally mature individuals have notably broad skulls, a broad processus trochlearis oticum, a shallow but broad median palatal groove, high accessory ridges, and a substantial contribution of the vomer to the secondary palate. Two large trionychid skulls and isolated shell pieces have been discovered in deposits of theHell Creek Formation exposed inCarter County, Montana, southeastern Montana. The skulls are noticeably larger than those previously discovered and differ in their overall form by being significantly broader. Detailed analysis reveals that the new skulls and shell fragments correspond with changes in overall form representing an ontogenetic shift towards the adult morphology of this species.^[3]

Phylogeny

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Cladogram after Joyce & Lyson, 2011:^[1]

Gilmoremys

Hutchemys

	H. rememdium

	H. arctochelys

	H. tetanetron

	H. sterea

References

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^^a ^bWalter G. Joyce and Tyler R. Lyson (2011). "New Material ofGilmoremys lancensis nov. comb. (Testudines: Trionychidae) from the Hell Creek Formation and the Diagnosis of Plastomenid Turtles".Journal of Paleontology.85 (3):442–459.doi:10.1666/10-127.1.S2CID 129918006.
^Walter G. Joyce; Tyler R. Lyson; Joseph J.W. Sertich (2018). "A new species of trionychid turtle from the Upper Cretaceous (Campanian) Fruitland Formation of New Mexico, USA".Journal of Paleontology.92 (6):1107–1114.doi:10.1017/jpa.2018.30.S2CID 134161813.
^Joyce, Walter; Lyson, Tyler; Williams, Scott (2016). "New Cranial Material of Gilmoremys lancensis (Testudines, Trionychidae) from the Hell Creek Formation of Southeastern Montana, U.S.A.".Journal of Vertebrate Paleontology.36 (6): e1225748.doi:10.1080/02724634.2016.1225748.S2CID 133358179.

Trionychidae family

Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Subclass:Anapsida
Order:Testudines
Superfamily:Trionychia

Genera

Species of theTrionychidae family

Amyda

Apalone

†Axestemys

Chitra

Cyclanorbis

Cycloderma

Dogania

Malayan softshell turtle

†Gilmoremys

†Gilmoremys gettyspherensis
†Gilmoremys lancensis

†Hutchemys

†Khunnuchelys

Lissemys

Nilssonia

†Palaeoamyda

†Palaeoamyda messeliana

Palea

Wattle-necked softshell turtle

Pelochelys

Pelodiscus

Rafetus

Trionyx

Phylogenetic arrangement of turtles based onturtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. †=extinct.

Testudines

Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Diapsida
Order:Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based onTurtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See alsoList of Testudines families

Taxon identifiers
Gilmoremys	Wikidata:Q5562540 Wikispecies:Gilmoremys GBIF:8537838 IRMNG:1473148 Open Tree of Life:6156170 Paleobiology Database:212673

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