Columella (in plants) is an axis of sterile tissue which passes through the center of the spore-case ofmosses.[1] Infungi, it refers to a centrally vacuolated part of ahypha, bearingspores. The word finds analogous usage inmyxomycetes.[citation needed]
The term columella is also used to refer to story 1 to story 4 (S1 – S4) cells in theroot cap, located apically of thequiescent centre.[citation needed] Invascular plants, columella tissue helps the roots to perceive gravity. Within columella cells, theamyloplasts andGolgi apparatus organelles tend to fall to the bottom third of the cell.[2]
