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Appendage

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(Redirected fromAppendages)
External body part or natural prolongation, that protrudes from an organism's body
For I/O appendages in OS/360 and successors, seeExecute Channel Program § Appendages. For the EP by Circa Survive, seeAppendage (EP). For the horror film, seeAppendage (film).
Look upappendage in Wiktionary, the free dictionary.
A beetle leg

Anappendage (oroutgrowth) is an external body part or natural prolongation that protrudes from anorganism's body such as an arm or a leg. Protrusions from single-celledbacteria andarchaea are known as cell-surface appendages or surface appendages. In many kinds ofeukaryotic cell the protrusions are known asmembrane protrusions or cell appendages (examples includemicrovilli andcilia).

Types in animals

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Inarthropods, an appendage refers to any of thehomologous body parts that may extend from abody segment, includingantennae,mouthparts (includingmandibles,maxillae and maxillipeds),gills,locomotorlegs (pereiopods forwalking, andpleopods forswimming),sexual organs (gonopods), and parts of thetail (uropods). Typically, each body segment carries one pair of appendages. An appendage which is modified to assist in feeding is known as amaxilliped orgnathopod.[citation needed]

Inannelids lateral protrusions from the body are calledparapodia.

Inechinoderms an appendage called apedicellaria is found. The end of the pedicellaria consists of valves that give a jaw-like appearance and is thought to be used to clear the external body surface. Echinoderms also possess podia known astube feet. Tube feet form part of thewater vascular system and are used for locomotion, food and waste transportation, and respiration.

Allcephalopods, amollusc class, have flexible appendages known ascephalopod limbs. They may have further extensions assuckers.

Invertebrates, an appendage can refer to a locomotor part such as atail,fins on afish,limbs (arms,legs,flippers orwings) on atetrapod; exposedsex organ; defensive parts such ashorns andantlers; orsensory organs such asauricles,proboscis (trunk andsnout) andbarbels.[citation needed]

Appendages may becomeuniramous, as ininsects andcentipedes, where each appendage comprises a single series of segments, or it may bebiramous, as in manycrustaceans, where each appendage branches into two sections.Triramous (branching into three) appendages are also possible.[1]

Allarthropod appendages are variations of the same basic structure (homologous), and which structure is produced is controlled by "homeobox" genes. Changes to these genes have allowed scientists to produce animals (chieflyDrosophila melanogaster) with modified appendages, such as legs instead of antennae.[2]

Types in prokaryotes

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A number ofcell-surface appendages are found inprokaryotesbacteria andarchaea, and includearchaella,flagella,pili,fimbriae, andprosthecae also calledstalks.

Archaea

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A number of cell-surface appendages may be present on different archaea. Two types of appendage are species-specific;cannulae are specific toPyrodictium species, andhami are specific toAltiarchaeum.[3] Other various types of surface structure includepili,archaella (archaeal flagella), structures calledbindisomes that bind sugars, and posttranslationally modified archaellins and pilins.[4][5]

Archaella are the similar structures to bacterial flagella, serving the same function in motility, particularly swimming, but with a different composition and action. Pili are used for attachment to surfaces, possible communication between cells enabling cell-to-cell contact allowing genetic transfer, and the formation ofbiofilms.[4] Atype IV pili model is used in the assembly of several cell surface structures. The bindisome is made up of sugar binding proteins to facilitate sugar uptake. So far studies are limited toS. solfataricus.[4] Appendage fibres described asIho670 fibres are unique toIgnicoccus hospitalis.[4]

Bacteria

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Bacterial cell-surface appendages includeflagella,pili, short attachment pili known asfimbriae, and on some speciescurli fibres.Some bacteria also have stalks known asprosthecae. Other appendages arebacterial nanowires.

Types in eukaryotic cells

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Cell appendages aremembrane protrusions that extend from thecell membrane, examples aremicrovilli andcilia.

Types in plants

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Aleaf is the main appendage of a plant stem.Prosthechea is a genus oforchids named for the prostheca appendage on the back of thecolumn. Hair like structures known astrichomes are found on many types of plants.

See also

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References

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  1. ^Morris, Simon Conway (1979)."The Burgess Shale (Middle Cambrian) Fauna".Annual Review of Ecology and Systematics.10:327–349.doi:10.1146/annurev.es.10.110179.001551.JSTOR 2096795. Retrieved17 November 2020.
  2. ^Emerald, B. Starling; Cohen, Stephen (March 2004). "Spatial and temporal regulation of the homeotic selector gene Antennapedia is required for the establishment of leg identity in Drosophila".Developmental Biology.267 (2):462–472.doi:10.1016/j.ydbio.2003.12.006.PMID 15013806.
  3. ^van Wolferen, M; Pulschen, AA; Baum, B; Gribaldo, S; Albers, SV (November 2022)."The cell biology of archaea".Nature Microbiology.7 (11):1744–1755.doi:10.1038/s41564-022-01215-8.PMC 7613921.PMID 36253512.
  4. ^abcdJarrell, KF; Ding, Y; Nair, DB; Siu, S (24 January 2013)."Surface appendages of archaea: structure, function, genetics and assembly".Life.3 (1):86–117.Bibcode:2013Life....3...86J.doi:10.3390/life3010086.PMC 4187195.PMID 25371333.
  5. ^Lassak, K; Ghosh, A; Albers, SV (November 2012)."Diversity, assembly and regulation of archaeal type IV pili-like and non-type-IV pili-like surface structures".Research in Microbiology.163 (9–10):630–44.doi:10.1016/j.resmic.2012.10.024.PMID 23146836.S2CID 37802794.

See also

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