1992, Rudolf M[athias] Schuster,The Hepaticae and Anthocerotae of North America: East of the Hundredth Meridian, volume V, Chicago, Ill.:Field Museum of Natural History,→ISBN, page 4:
The Jubulaceae have a leaf whose lobule, usually transformed into a water-sac, is normally very narrowly attached to the stem and to the dorsal lobe; indeed someFrullania taxa reproduce vegetatively by dropping the dorsal lobes, but not the lobules, andNeohattoria hascaducous lobules but persistent lobes.