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Youngina

From Wikipedia, the free encyclopedia
(Redirected fromYounginia)
Extinct genus of reptiles

Youngina
Temporal range:Late Permian,259.9–254.17 Ma
Speculativelife restoration
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Diapsida
Clade:Neodiapsida
Family:Younginidae
Genus:Youngina
Broom 1914
Type species
Youngina capensis

Youngina (named afterJohn Young (1823–1900)) is anextinctgenus ofdiapsidreptile from theLate PermianBeaufort Group (Tropidostoma-Dicynodon zones) of theKaroo Red Beds ofSouth Africa.[1] This, and a few related forms, make up thefamily Younginidae, within the orderEosuchia (proposed by Broom in 1914). Eosuchia, having become awastebasket taxon for many probably distantly-related primitive diapsid reptiles ranging from the LateCarboniferous to theEocene,Romer proposed that it be replaced byYounginiformes (that included Younginidae and the Tangasauridae, ranging from thePermian to theTriassic).

Taxonomy

[edit]
Reconstructed skull

Youngina is known from several specimens. Many of these were attributed to as separate genera and species (such asYoungoides andYoungopsis), but it was later realized that they were not distinct fromY. capensis.[2][3] Theholotype specimen ofYoungina, discovered byBroom himself,[2] was described briefly in1914.[4] The "Youngoides romeri" specimen was first attributed toYoungina,[5] but later given its eponymous and separate designation in a later paper.[6]Acanthotoposaurus[7] is also ajunior synonym ofYoungina.[8]

Description

[edit]
Size compared to a human hand

Youngina was a relatively small reptile, with a skull length of 5 centimetres (2.0 in) and a total body length of 30 centimetres (12 in).[9] Thebraincase anatomy was redescribed in 2010.[10]Youngina shows a mosaic of features found in more primitive diapsids and more derived taxa such asarchosauromorphs andlepidosauromorphs suggesting a non-orthogenetic evolution of these characters. Though the palatobasal articulation is open,[10] it was probably immobile, similar to the skull of thetuatara,[11] contrary to some earlier claims made about the metakinetic mobility of basicranial joints inYoungina and other early diapsid reptiles.[12]

Phylogeny

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Youngina was once thought to be closely related toAcerosodontosaurus, and more distantly totangasaurids (Kenyasaurus,Hovasaurus,Thadeosaurus, andTangasaurus), but the monophyly of younginiforms has not been demonstrated in published analyses of diapsid reptiles, and it is likely this group is paraphyletic.Acerosodontosaurus is probably closer to other formeryounginiforms, rather than being closely related toYoungina.[13] Below is acladogram from the analysis of Reiszet al. (2011) showing thephylogenetic position ofYoungina among early diapsids:[14]

References

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  1. ^Smith, R.; Evans, S. (1996)."New Material ofYoungina: Evidence of Juvenile Aggregation in Permian Diapsid Reptiles"(PDF).Palaeontology.39 (2):289–303.Archived(PDF) from the original on 2019-07-30.
  2. ^abGow, C. E. (1975). "The morphology and relationships ofYoungina capensis Broom andProlacerta broomi Parrington".Palaeontologia Africana.18:89–131.hdl:10539/16290.
  3. ^Evans, S. E. (1987). "The braincase ofYoungina capensis (Reptilia: Diapsida; Permian)".Neues Jahrb. Geol. Paläontol. Monats.1987:193–203.
  4. ^Broom, R. (1914)."A new thecodont reptile".Proceedings of the Zoological Society of London. B.84 (4):1072–1077.doi:10.1111/j.1469-7998.1914.tb07729.x.
  5. ^Olson, E. C. (1936). "Notes on the Skull of Youngina Capensis Broom".Journal of Geology.44 (4):523–533.Bibcode:1936JG.....44..523O.doi:10.1086/624447.JSTOR 30068180.S2CID 140142731.
  6. ^Olson, E. C.; Broom, R. (1937). "New genera and species of tetrapods form the Karroo beds of South Africa".Journal of Paleontology.11 (7):613–619.JSTOR 1298390.
  7. ^Evans, S. E.; Van der Heever, J. A. (1987)."A new reptile (Reptilia: Diapsida) from the Upper Permian Daptocephalus zone of South Africa".South African Journal of Science.83 (11):724–730.
  8. ^Reisz, R. R.; Modesto, S. P.; Scott, D. (2000)."Acanthotoposaurus bremneri and the origin of the Triassic archosauromorphreptile fauna of South Africa".South African Journal of Science.96 (8):443–445.
  9. ^Hunt, Annabel K.; Ford, David P.; Fernandez, Vincent; Choiniere, Jonah N.; Benson, Roger B. J. (September 2023)."A description of the palate and mandible of Youngina capensis (Sauropsida, Diapsida) based on synchrotron tomography, and the phylogenetic implications".Papers in Palaeontology.9 (5).doi:10.1002/spp2.1521.ISSN 2056-2799.
  10. ^abGardner, Nicholas M.; Holliday, Casey M.; O'Keefe, F. Robin (2010)."The Braincase ofYoungina capensis (Reptilia, Diapsida): New Insights from High-Resolution CT Scanning of the Holotype".Palaeontologia Electronica.13 (3): 19A.
  11. ^Johnston, P. (2010). "The constrictor dorsalis musculature and basipterygoid articulation inSphenodon".Journal of Morphology.271 (3):280–292.doi:10.1002/jmor.10797.PMID 19806656.S2CID 32965233.
  12. ^Evans, S. E. (1980)."The skull of a new eosuchian reptile from the Lower Jurassic of South Wales".Zoological Journal of the Linnean Society.70 (3):203–264.doi:10.1111/j.1096-3642.1980.tb00852.x.
  13. ^Bickelmann; Müller; Reisz (2009). "The enigmatic diapsidAcerosodontosaurus piveteaui (Reptilia: Neodiapsida) from the Upper Permian of Madagascar and the paraphyly of "younginiform" reptiles".Canadian Journal of Earth Sciences.46 (9):651–661.Bibcode:2009CaJES..46..651S.doi:10.1139/E09-038.
  14. ^Reisz, Robert R.; Modesto, Sean P.; Scott, Diane M. (2011)."A new Early Permian reptile and its significance in early diapsid evolution".Proceedings of the Royal Society B.278 (1725):3731–3737.doi:10.1098/rspb.2011.0439.PMC 3203498.PMID 21525061.

External links

[edit]
Sauropsida(Reptiliasensu lato)
Tetrapodomorpha
Reptiliomorpha
Sauropsida
    • see below↓
Acleistorhinidae
Bolosauridae
Mesosauria
Millerettidae
Procolophonia
Diapsida
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Mesosaurus tenuidensMilleretta rubidgei
Tangasauridae
Weigeltisauridae
Drepanosauromorpha
Kuehneosauridae
Choristodera
Ichthyosauromorpha
Sauropterygia
Thalattosauria
Sauria
(Crown-Reptilia)
Lepidosauromorpha
Archelosauria
Archosauromorpha
Pantestudines
Hovasaurus boulei

Weigeltisaurus jaekeli

Megalancosaurus preonensis
See also
Possible sauropsids
Other taxa
Youngina
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