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Ulmus 'Exoniensis'

From Wikipedia, the free encyclopedia
Elm cultivar

Ulmus 'Exoniensis'
Exeter Elm inAmsterdam
GenusUlmus
Cultivar'Exoniensis'
OriginExeter, England

Ulmus 'Exoniensis', theExeter elm, was discovered nearExeter,England, in 1826, and propagated by the Ford & Please nursery in that city.[1][2][3][4][5] Traditionally believed to be acultivar of the Wych ElmU. glabra, its fastigiate shape when young, upward-curving tracery, smallsamarae and leaves, late leaf-flush and late leaf-fall, taken with its south-west England provenance, suggest a link with theCornish Elm, which shares these characteristics. The seed, however, is on the stalk side of the samara, a feature of wych elm and its cultivars, whereas in hybrids it would be displaced towards the notch.[6][7]

Description

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The tree initially has an upright, columnar form,[8] but later develops a large rounded crown and occasionally reaches 17 m in height. Older specimens may develop pendulous branches.[9] Exeter Elm is chiefly distinguished by its contorted leaves, < 11 cm long by 8 cm broad, rounder than the type [wych] and with morelaciniate margins,[10][11] which occasionally wrap around the branchlets and remain thus well into winter.[12] Thesamara is narrower than type wych.[13][7] 'Exoniensis' is often pollarded to produce a denser, fan-shaped crown (see main picture).

  • 'Exoniensis' in April
    'Exoniensis' in April
  • Bole of Exeter Elm
    Bole of Exeter Elm
  • Characteristic bole-burls of Exeter elm (left), Bruntsfield Links, Edinburgh
    Characteristic bole-burls of Exeter elm (left), Bruntsfield Links, Edinburgh
  • 'Exoniensis' samarae
    'Exoniensis' samarae
  • 'Exoniensis' fruiting, late April, the Brink, Betondorp, the Netherlands (2014)
    'Exoniensis' fruiting, late April, theBrink,Betondorp, the Netherlands (2014)
  • 'Exoniensis' foliage
    'Exoniensis' foliage
  • 'Exoniensis' foliage
    'Exoniensis' foliage
  • Dried 'Exoniensis' leaves
    Dried 'Exoniensis' leaves
  • 'Exoniensis' scion grafted on U. minor stock
    'Exoniensis' scion grafted onU. minor stock

Pests and diseases

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Chevalier noted (1942) thatUlmus montana fastigiata (Exeter Elm) was one of four European cultivars found by researchers in The Netherlands to have significant resistance to the earlier strain ofDutch elm disease prevalent in the 1920s and '30s, the others being'Monumentalis'Rinz,'Berardii' and'Vegeta'. The four were rated less resistant thanU. foliacea clone 23, from Spain, later cultivated asU. minor 'Christine Buisman'.[14] 'Exoniensis' possesses a moderate resistance to the more virulent strain ofDutch elm disease, and consequently often featured in the Dutch elm breeding programme in association with the Field Elm (U. minor) and Himalayan Elm (U. wallichiana).[15]

Cultivation

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Once commonly planted in the UK and parts of western Europe, notably in the Netherlands (aspluim-iep, plume-elm),[16] and in Stockholm, Sweden,[17] 'Exoniensis' is also known to have been marketed in Poland in the 19th century by the Ulrich nursery,[18]Warsaw, and remains in commerce there. TheSpäth nursery of Berlin cultivated the tree asU. montana fastigiata (U. exoniensisHort.) from the early 20th century.[19] It is possible that three trees supplied by theSpäth nursery to theRoyal Botanic Garden Edinburgh in 1902 asU. montana fastigiata[20] were Exeter Elm,[19] old specimens of which survive inEdinburgh (one each in Drummond Place Gardens and Lochend Park).[20] Among plantings of Exeter elm inTurku, Finland, are three old trees in front of the city hall, on the riverAura.[21][22]

'Exoniensis' is found in Australia at theBallarat Botanical Gardens where it is listed on the Significant Tree Register of the National Trust. AnUlmus plumosa (a synonym of 'Exoniensis' in continental Europe[23]), of "elegant and pyramidal shape" and "dark green foliage", appeared in the 1902 catalogue of the Bobbink and Atkins nursery,Rutherford, New Jersey.[24]

  • Exeter elms (right), Grimsby Magistrates Court, Lincolnshire (2009)
    Exeter elms (right), Grimsby Magistrates Court, Lincolnshire (2009)
  • Three Exeter elms (bottom right) before the City Hall, Turku, Finland (2011)
    Three Exeter elms (bottom right) before the City Hall, Turku, Finland (2011)
  • Exeter elm (background), De Iepenhof [: elmcourt] cemetery, Hoofddorp, Netherlands (2016)[16]
    Exeter elm (background), De Iepenhof [: elmcourt] cemetery,Hoofddorp, Netherlands (2016)[16]
  • Exeter elm (background), De Iepenhof cemetery, Hoofddorp (2016)
    Exeter elm (background), De Iepenhof cemetery, Hoofddorp (2016)
  • 'Exoniensis' in April, the Brink, Betondorp, the Netherlands (2022)
    'Exoniensis' in April, theBrink,Betondorp, the Netherlands (2022)

In Sweden 'Exoniensis' is sometimes pruned from an early age to form a tidy cone-shaped tree called locally 'pyramidalm' (: pyramid elm - also one of Späth's names for 'Exoniensis').[25][26][17]

Notable trees

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Bean (1936) noted a large old specimen, 12 feet in girth, in the garden of the Old Vicarage,Bitton, Gloucestershire.[27] The oldest in Sweden, atGustav III's Pavilion inHagaparken, had a height in 2008 of 19 m and a bole girth of 462 cm.[17] A 180-year-old specimen inHamburg has attained a height of 28 m and a trunk diameter of 1.45 m.[28] The UKTROBI Champion tree is in Scotland, atBaxter Park,Dundee, measuring 15 m high by 103 cmd.b.h. in 2004.[29] The cultivar is represented in Éire by a tree atBirr Castle (Mount Palmer),County Offaly, with ad.b.h. of 29 cm when measured in 2002.

  • Early slide of an Exeter Elm in the Netherlands
    Early slide of an Exeter Elm in the Netherlands
  • Old unpollarded Exeter Elm, Lochend Park, Edinburgh, showing pendulous branch
    Old unpollarded Exeter Elm, Lochend Park, Edinburgh, showing pendulous branch

Varieties

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The Baudriller nursery of Angers, France, marketed a variegated Exeter elm in the 1880s,U. montana 'Fastigiata Variegata'.[30]

Hybrid cultivars

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'Clusius','Columella','Dodoens','Lobel','Plantyn','Nanguen' =Lutèce,'Wanoux' =Vada. The cultivar 'Columella' features the same rough, rounded, contorted leaves, the result of a recessive gene inherited from the Exeter Elm.

'Exoniensis' also indirectly featured in the Italian elm breeding programme as an ancestor of 'Plantyn', which was crossed with clones of theSiberian ElmUlmus pumila to produce the cultivars'Arno','Plinio', and'San Zanobi'.[31][32]

Despite being of British provenance and, at the time of writing, the only wych cultivar with any resistance to Dutch elm disease, and despite contributing to the complex hybrids above, Exeter elm did not appear in Seddon and Shreeve'sGreat British Elms (Kew, 2024).[33]

Synonymy

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Accessions

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Europe
Australasia
  • Ballarat Botanical Gardens, Australia. 1 tree, planted c.1900. Acc. no. T11321.[37][38]

References

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  1. ^Bean, W. J. (1981).Trees and shrubs hardy in Great Britain, 7th edition. Murray, London
  2. ^Elwes, Henry John;Henry, Augustine (1913).The Trees of Great Britain & Ireland. Vol. 7. p. 1866.
  3. ^Richens, R. H. (1983).Elm. Cambridge University Press.
  4. ^White, J. & More, D. (2003).Trees of Britain and Northern Europe. Cassell's, London.
  5. ^Harvey, J. (1974).Early Nurserymen.  p.104. Phillimore & Co. Ltd. 1975.ISBN 978-0850331929
  6. ^Coleman, Max, ed.:Wych Elm (Edinburgh, 2009)
  7. ^abFruit of Exeter elm, exetertrees.uk
  8. ^Photograph of fastigiate form of young Exeter elm
  9. ^Wilkinson, Gerald,Epitaph for the Elm (London 1978), p.62
  10. ^Bean, W. J. (1936)Trees and shrubs hardy in Great Britain, 7th edition, Murray, London, vol. 2, p.617
  11. ^ab"Muséum national d'histoire naturelle, spécimen P06883092" Two clones, (left)U. exoniensis (misnamed "U. oxoniensis" in France in 19th century), (right)U. pyramidata (1863)
  12. ^"Photograph of 'Exoniensis' leaves in early summer". Archived fromthe original on 11 July 2011. Retrieved10 November 2010.
  13. ^'Exoniensis' fruit and new leaves, Arboretum Dreyen, Wagingingen, 1962; herbarium specimen WAG.1846640; bioportal.naturalis.nl
  14. ^"Les Ormes de France"(PDF).Revue de botanique appliquée et d'agriculture coloniale.22 (254): 441. 1942.[dead link]
  15. ^Heybroek, H.M. (1993)."The Dutch Elm Breeding Program". In Sticklen, Mariam B.; Sherald, James L. (eds.).Dutch Elm Disease Research. New York, USA: Springer-Verlag. pp. 16–25.ISBN 978-1-4615-6874-2. Retrieved26 October 2017.
  16. ^abExeter elm in the Netherlands, monumentaltrees.com
  17. ^abcLars Lagerstedt, Pyramid Elm inLustgarden, 2013, p.40
  18. ^Ulrich, C. (1894),Katalog Drzew i Krezewow, C. Ulrich, Rok 1893–94, Warszawa
  19. ^abKatalog(PDF). Vol. 108. Berlin, Germany: L. Späth Baumschulenweg. 1902–1903. pp. 132–133.
  20. ^abAccessions book. Royal Botanic Garden Edinburgh. 1902. pp. 45, 47.
  21. ^Exeter elms, City Hall, Turku, Finland; /turunpuut.fi/en/ulmus/ulmus-glabra-exoniensis/2189/
  22. ^Coppiced Exeter elm, Turku, Finland; https://turunpuut.fi/en/ulmus/ulmus-glabra-exoniensis/2985/
  23. ^De Vos, C. (1867).Beredeneerd woordenboek der voornaamste heesters en coniferen, in Nederland gekweekt. Groningen: J. B. Wolters. p. 135.
  24. ^Bobbink and Atkins, Rutherford. N.J. 1902. p. 51.
  25. ^Photographs of 'Pyramidalm' and unpruned 'Exoniensis' in Sweden, www.tradgardsakademin.se[1]Archived 16 June 2011 at theWayback Machine
  26. ^Lagerstedt, Lars (2014)."Märkesträd i Sverige - 10 Almar" [Notable trees in Sweden - 10 Elms](PDF).Lustgården.94: 63, 73. Retrieved15 May 2018.
  27. ^Bean, W. J. (1936)Trees and shrubs hardy in Great Britain, 7th edition, Murray, London, vol. 2, p.617
  28. ^U. glabra 'Exoniensis', the "Planten un Blomen", Hamburg: from theHandbuch der Ulmengewächse,[2]
  29. ^Johnson, O. (2011).Champion Trees of Britain & Ireland, p. 169. Kew Publishing, Kew, London.ISBN 9781842464526.
  30. ^Baudriller Établissement d'Horticulture,Catalogue général descriptif et raisonné des arbres fruitiers, forestiers & d'ornement cultivés dans l'établissement. Année. 1880, p.117
  31. ^Santini A., Fagnani A., Ferrini F. & Mittempergher L., (2002) 'San Zanobi' and 'Plinio' elm trees.[3]Archived 26 January 2004 at theWayback MachineHortScience 37(7): 1139–1141. 2002. American Society for Horticultural Science, Alexandria, VA 22314, USA.
  32. ^Santini A., Fagnani A., Ferrini F., Mittempergher L., Brunetti M., Crivellaro A., Macchioni N., Elm breeding for DED resistance, the Italian clones and their wood properties.[4]Archived 26 October 2007 at theWayback MachineInvest Agrar: Sist. Recur. For. (2004) 13 (1), 179–184. 2004
  33. ^Mark Seddon and David Shreeve,Great British Elms (Kew Gardens, 2024)
  34. ^De Vos, C. (1867).Beredeneerd woordenboek der voornaamste heesters en coniferen, in Nederland gekweekt. Groningen: J. B. Wolters. p. 136.
  35. ^"List of plants in the {elm} collection". Brighton & Hove City Council. Retrieved23 September 2016.
  36. ^Netherlands Plant Collection: Iepen,Ulmus
  37. ^Ulmus glabra 'Exoniensis', Ballarat Botanical Gardens; Victorian Heritage Database Report
  38. ^Ulmus glabra 'Exoniensis' photographs, Ballarat Botanical Gardens; vhd.heritagecouncil.vic.gov.au
Species, varieties and subspecies
Disputed species, varieties and subspecies
Hybrids
Speciescultivars
American elm
Cedar elm
Chinese elm
European white elm
Field elm
Japanese elm
Siberian elm
Winged elm
Wych elm
Hybridcultivars
Dutch elm
U. ×intermedia
Unconfirmed derivation cultivars
Fossil elms
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