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Tugulusaurus

From Wikipedia, the free encyclopedia
Extinct genus of dinosaurs

Tugulusaurus
Temporal range:Early Cretaceous,Barremian–Albian
A hypothetical skeletal reconstruction ofTugulusaurus with known material in white
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Dinosauria
Clade:Saurischia
Clade:Theropoda
Clade:Alvarezsauria
Superfamily:Alvarezsauroidea
Genus:Tugulusaurus
Dong, 1973
Species:
T. faciles
Binomial name
Tugulusaurus faciles
Dong, 1973

Tugulusaurus (meaning "Tugulu lizard") is agenus ofcoelurosauriantheropoddinosaur, belonging to theAlvarezsauroidea, that is known from the EarlyCretaceousTugulu Group in theUrhe area ofChina. It was one of the first members of Alvarezsauria ever discovered. Thetype and only known species isT. faciles.

Discovery

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From 1964 dinosaur fossils were excavated in theJunggar Basin ofXinjiang. In 1973 a number of these were described by paleontologistDong Zhiming, among them the bones of a small theropod which he namedTugulusaurus faciles. The generic name refers to the Tugulu Group. Thespecific name is derived fromLatinfacilis, here with the meaning of "easily moving", referring to the agility of the animal as indicated by its "delicate bones".[1]

Theholotype,IVPP V4025, was found in layers of theLianmuqin Formation dating from the Barremian–Albian. It consists of a partial skeleton including four partial tailvertebrae, much of the left leg and part of the right, the first fingers of both hands, and a rib. Thefemur has a length of about 215 millimetres (8.5 in). The left firstmetacarpal is very short: 26 millimetres (1.0 in). The skeleton represents the only remains of the species that have ever been discovered.[2]

Description

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Tugulusaurus was an early alvarezsaur, likely measuring at 1.9–3 metres (6.2–9.8 ft) in length. It probably had a relatively long neck and tail, as well as a streamlined head with a beak-like snout, and large, hooked claws. It was an agile animal, likely having long legs, which would aid in reaching high speeds, in order to chase prey and retreat from potential threats. Being an alvarezsauroid, it likely used its long claws to dig and tear into wood or dirt in order to find food.[1]

Classification

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Tugulusaurus was originally classified by Dong in 1973 as a member of theOrnithomimidae, within theCoelurosauria.[3] In the years that followed, the genus was often considered anomen dubium.[4] However, in 2005Oliver Rauhut andXu Xing concluded that it is a valid genus ofbasal coelurosaurian of unknown affinities.[2] In their cladistic analysis of the newly described taxaBannykus andXiyunykus, Xu et al. (2018) recoveredTugulusaurus as a member ofAlvarezsauria.[5]

References

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  1. ^abDong, Z.-M. 1973. [Dinosaurs from Wuerho]. Memoirs of the Institute of Vertebrate Paleontology and Paleoanthropology, Academia Sinica11: 45–52 [Chinese]
  2. ^abRauhut, Oliver W.M.; Xu, Xing (2005). "The small theropod dinosaursTugulusaurus andPhaedrolosaurus from the Early Cretaceous of Xinjiang, China".Journal of Vertebrate Paleontology.25 (1):107–118.doi:10.1671/0272-4634(2005)025[0107:TSTDTA]2.0.CO;2.
  3. ^Dong Zhiming (1992).Dinosaurian Faunas of China. China Ocean Press, Beijing.ISBN 978-3-540-52084-9.
  4. ^Norman, D. B. (1990). "Problematic Theropoda: Coelurosaurs". 280-305 in Weishampel, D. B., Dodson, P., & Osmólska, H. (eds.)The Dinosauria. Berkeley: University of California Press, Berkeley 1990, xvi-733
  5. ^Xing Xu; Jonah Choiniere; Qingwei Tan; Roger B.J. Benson; James Clark; Corwin Sullivan; Qi Zhao; Fenglu Han; Qingyu Ma; Yiming He; Shuo Wang; Hai Xing; Lin Tan (2018). "Two Early Cretaceous fossils document transitional stages in alvarezsaurian dinosaur evolution". Current Biology. Online edition.doi:10.1016/j.cub.2018.07.057.
Avemetatarsalia
Theropoda
Maniraptora
    • see below↓
Alvarezsauridae
Parvicursorinae
Ceratonykini
Mononykini
Therizinosauria
Therizinosauroidea
Therizinosauridae
Pennaraptora
Oviraptorosauria
Paraves
    • see below↓
Patagonykus puertai

Mononykus olecranus

Therizinosaurus cheloniformis
Scansoriopterygidae?
Anchiornithidae
Archaeopterygidae
Dromaeosauridae
Troodontidae
Jeholornithiformes
Omnivoropterygidae?
Confuciusornithidae
Jinguofortisidae
Ornithothoraces
Enantiornithes
Euornithes
    • see below↓
Ambopteryx longibrachium

Archaeopteryx lithographica

Confuciusornis sp.
Schizoouridae
Patagopterygiformes
Ambiortiformes
Hongshanornithidae
Songlingornithidae
Yanornithidae
Gansuidae?
Ichthyornithes
Hesperornithes
Hesperornithidae
Cimolopterygidae
Aves / Neornithes
    • see below↓
Patagopteryx deferrariisiIchthyornis dispar
Palaeognathae
Neognathae
Galloanserae
Anserimorphae
Pangalliformes
Incertae sedis
Dromornithidae
Gastornithiformes
Pelagornithidae
Asteriornis maastrichtensisDromornis stirtoni
Tugulusaurus faciles
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