Mean annual temperature is between 4 and 12 °C (39 and 54 °F).
However, required annual precipitation depends on factors such as distribution of rain over the year, temperatures over the year and fog presence, and definitions in other regions of the world differ considerably. For example, Australian definitions areecological-structural rather thanclimatic:
Closedcanopy of trees excludes at least 69% of the sky.
Forest is composed mainly of tree species which do not require fire for regeneration, but withseedlings able to regenerate under shade and in natural openings.[3]
Australian definitions would exclude some temperate rainforests of western North America that areCoast Douglas-fir dominant, such as parts of theKlamath Mountains in southern Oregon and northern California, thePuget Lowlands of western Washington and theGeorgia Depression in British Columbia,[4][5] as their dominant tree species, the Coast Douglas-fir, requires stand-destroying disturbance to initiate a new cohort of seedlings.[6] The North American definition would in turn exclude a part of temperate rainforests under definitions used elsewhere.[7]
Forforests,canopy refers to the upper layer orhabitat zone, formed by maturetree crowns and including other biological organisms (epiphytes,lianas, arboreal animals, etc.). The canopy level is the third level of the temperate rainforest. The trees forming the canopy,conifers, can stand as tall as 100 metres or more. A variety of species survive in the canopy. The tops of these trees collect most of the rain, moisture, and photosynthesis that therainforest takes in. They form a canopy over the forest, covering about 95% of the floor during the summer.
The canopy's coverage affects theshade tolerance levels of forest floor plants. When the canopy is in full bloom, covering about 95% of the floor, plant survival decreases. Some plant species have become shade tolerant in order to survive. The treetops take in the heavy amount of rain and keep the lower levels of the forest damp.
The canopy survives throughphotosynthesis. The leaves provide energy and nutrients for the trees, which provide homes and food for the forest. Through satellite data, the radiation use efficiency (RUE) calculates the annual amount of photosynthesis that occurs in temperate rainforests. A diverse amount of photosynthesis occurs based on the location andmicroclimates of the forest.[8]
Temperate rainforest in theMount Hood Wilderness, Oregon, US. This area, on the west side of the mountain, receives close to 100 inches (2,500 mm) of rain per year.
Red spruce and Fraser fir are dominant canopy trees in high mountain areas. In higher elevation (over 1,980 metres; 6,500 feet), Fraser fir is dominant, in middle elevation (1,675 to 1,890 metres; 5,495 to 6,201 ft) red spruce and Fraser fir grow together, and in lower elevation (1,370 to 1,650 metres; 4,490 to 5,410 ft) red spruce is dominant. Yellow birch, mountain ash, and mountain maple grow in the understory. Younger spruce and fir and shrubs like raspberry, blackberry, hobblebush, southern mountain cranberries, red elderberry, minniebush, southern bush honeysuckle are understory vegetation. Below the spruce-fir forest, at around 1,200 metres (3,900 ft), are forests of American beech, yellow birch, maple birch, and oak. Skunk cabbage and ground juniper are northern species that were pushed into the areas from the north.
The mild and wet environment supports the high diversity of fungi. Over 2,000 species live in this area and scientists estimate many unidentified fungi may be there.[10]
Largeforested regions of theAdirondack and smaller parts of theCatskill mountains in New York state routinely receive over 55 inches of rain per year, classifying them as temperate rainforests.[16][17]
The Valdivian and Magellanic temperate rainforests are the only temperate rainforests in South America. Together they are the second largest in the world, after the Pacific temperate rainforests of North America. The Valdivian forests are a refuge for theAntarctic flora, and share many plant families and genera with the temperate rainforests ofNew Zealand,Tasmania, andAustralia. Fully half the species of woody plants areendemic to this ecoregion.
In the Valdivian region the Andean Cordillera intercepts moist westerly winds along the Pacific coast during winter and summer months; these winds cool as they ascend the mountains, creating heavy rainfall on the mountains' west-facing slopes. The northward-flowing oceanic Humboldt Current creates humid and foggy conditions near the coast. The tree line is at about 2,400 m in the northern part of the ecoregion (35°S), and descends to 1,000 m in the south of theValdivian region. In the summer the temperature can climb to 16.5 °C (61.7 °F), while during winter the temperature can drop below 7 °C (45 °F).[18]
The temperate rainforests ofSouth Africa are part of theKnysna-Amatole forests that are located along South Africa'sGarden Route between Cape Town and Port Elizabeth on the south-facing slopes of South Africa's Drakensberg Mountains facing the Indian Ocean. There are several coniferouspodocarps that grow here. This forest receives a lot of moisture as fog from the Indian Ocean, and resembles not only other temperate rainforests worldwide, but also the montane evergreenAfromontane forests that occur at higher elevations in southern and eastern Africa. A fine example of this forest is in South Africa'sTsitsikamma National Park.
Juniper montane rainforest, in Mistérios Negros,Terceira Island, Azores
The rainforests of theAzores (also known ascloud forests, due to the constant cloud coverage caused byorographic lift) are found in the more humid, montane areas that transition from the lower altitudelaurissilva. They are generally found at altitudes ranging from 600 to 1,000 m (2,000 to 3,300 ft), and receive 2,000 to 6,000 mm (79 to 236 in) of average annual rainfall.[19][20][21]
Despite being located in the temperate zone, the Azores rainforest is similar in many ways to the cloud forest environments of the tropics and subtropics.[22][19][23] These pluvial montane forests hold the highest biodiversity and degree of endemism of the whole archipelago.[19] They are dominated by dense formations of endemicjuniper,laurel,holly andtree heaths[19][20] with several species ofepiphytic ferns[24] and an abundance of mosses and rainforest lichens (such asErioderma).[25]
The climate in the rainforest is mild and cool, averaging 12 °C (54 °F)[19] with a narrow diurnal temperature range and temperatures that only drop below freezing in exceptional years.[26]
Since human settlement in the 15th century, these rainforests, which once covered most of the high altitudes of the archipelago, have gradually been reduced to relics and are now found almost exclusively on three of the nine islands (Flores,Pico andTerceira). Their main threat is the expansion of cattle grazing pastures.[20]
Temperate rainforest occurs in fragments across the north and west of Europe in countries such as southernNorway (seeScandinavian coastal conifer forests) and northernSpain. Other temperate rainforest regions include areas of south eastern Europe such as mountains on the east coast of theAdriatic Sea, surrounding North WesternBulgaria along with theBlack Sea.
In England, they occur in theLake District (Borrowdale Woods) and steep sided riverine and estuarine valleys in Devon and Cornwall and theMicroclimate disused slate & granite quarries in these counties. This includes theFowey valley in Cornwall and the valley of theriver Dart which flows offDartmoor and has rainfall in excess of 2 metres per year.[28]
Derrycunnihy Wood, located in theKillarney National Park, is the best example of the ancient damp-climate oceanic forest that covered an estimated 80 percent of Ireland prior to the arrival of humans in 7,000 BCE.
The Colchian rainforests are found around both the southeast and west corners of the Black Sea starting inBulgaria all the way toTurkey andGeorgia and are part of theEuxine-Colchic deciduous forests ecoregion, together with the drier Euxine forests further west. The Colchian rainforests are mixed, with deciduousblack alder(Alnus glutinosa), hornbeam (Carpinus betulus andC. orientalis),Oriental beech (Fagus orientalis), andsweet chestnut(Castanea sativa) together with evergreenNordmann fir (Abies nordmanniana, the tallest tree in Europe at 78 m),Caucasian spruce (Picea orientalis) andScots pine (Pinus sylvestris).The refugium is the largest throughout the Western Asian/ near Eastern region.[38][39][40] The area has multiple representatives ofdisjunctrelict groups of plants with the closest relatives in Eastern Asia, southern Europe, and even North America.[41][42][43] Over 70 species of forest snails of the region are endemic.[44] Some relict species of vertebrates areCaucasian parsley frog,Caucasian salamander,Robert's snow vole andCaucasian grouse; they are almost entirely endemic groups of animals such as lizards of genusDarevskia. In general,species composition of this refugium is quite distinct and differs from that of the other Western Eurasian refugia.[40] Genetic data suggest that the Colchis temperate rainforest, during theIce Age, was fragmented into smaller parts; in particular, evolutionary lineages of theCaucasian Salamander from the central and south-western Colchis remained isolated from one another during the entire Ice Age.[45]
The area was declared anatural park (a level of protection lower thannational park) in 1997. It is one of six natural parks in Galicia. TheEuropean Union has recognised the park as aSite of Community Importance. There are a number of species of ferns. Invertebrate species include theKerry slug and it is an important site for amphibians.
The Vinatovača rainforest, alternatively spelledVintovača, is the only rainforest in Serbia.[46] It has been left undisturbed for centuries due to strict conservation laws starting in the 17th century.
Vinatovača is situated in the centralKučaj mountains in theUpper Resava region, at an altitude between 640 m (2,100 ft) and 800 m (2,600 ft). It is isolated and hard to reach which helped its preservation. It is believed that trees have not been cut in Vinatovača since about 1650. Being under strict protection means not only that the trees that die of old age are not being cleared or removed, but even picking herbs or mushrooms is forbidden. It is considered as an example of what central and eastern Serbia's natural look is. Beech trees are up to 45 m (148 ft) tall and some specimens are estimated to be over 300 years old.[46]
TheCaspian Hyrcanian mixed forests ecoregion in northernIran contains a jungle in the form of a rainforest which stretches from the east in theKhorasan province to the west in theArdabil Province, covering the other provinces ofGilan,Mazandaran, andGolestan. The Elburz orAlborz mountain range is the highest mountain range in theMiddle East which captures the moisture of theCaspian Sea to its north and formssubtropical and temperate rainforests in the northern part ofIran. The Iranians call this forest and regionShomal which meansnorth inPersian. This forest was known for most of the history for being home to the now extinctCaspian Tiger.
Baemsagol valley ofJirisan, which is the southern end of Baekdu Mountain Range.
The forests that cover the mountains and valleys of theBaekdu Mountain Range – fromMt. Baekdu, in the north, toMt. Jiri, in the southwest, forming the spine of theKorean Peninsula – and thesouthern coast and islands of the peninsula – includingJeju Island – feature a wide variety of conifers and broadleaf trees. Much of these forests are protected in mountain and marine national forests, such as inHallyeohaesang National Park, which encompasses 150.14 km2 (57.97 sq mi) of mountainous forests spread out over 69 uninhabited islands and 30 inhabited islands in Korea's South Sea that provide a home to 1,142 plant species, including major species such as red pine, black pine, common camellia, serrata oak, and cork oak, as well as rare species such as nadopungnan (Phalaenopsis japonica), daeheongnan (Cymbidium macrorhizon) and the Korean winter hazel (Corylopsis coreana). Major animals species, such as otters, small-eared cats, and badgers also call Hallyeohaesang National Park home, and overall there are 25 mammal species, 115 bird species, 16 reptile species, 1,566 insect species, and 24 freshwater fish species found among the forested, mountains islands.[49]
Seoraksan National Park covers 398.539 km2 (153.877 sq mi) of mountainous forests near the eastern coast of the Korean Peninsula, and is aUNESCO designated Biosphere Preservation District. Over 2,000 animal species live in Seoraksan, including the Korean goral, musk deer, and there are also more than 1,400 rare plant species, such as the edelweiss.[50]
Some of the best preserved examples of forest are found inKirishima-Yaku National Park on the Island ofYakushima off of Kyūshū in a very wet climate (the annual rainfall is 4,000 to 10,000 mm depending on altitude). Because of relatively infertile soils on granite, Yakushima's forests in higher elevations are dominated by a giantconifer species, Japanese Cedar (Cryptomeria japonica), rather than deciduous forests typical of the mainland.[48][53] Other areas includeMount Kirishima nearKagoshima in southern Kyūshū. On Southern Honshū, there is a forest with theNachi Falls located inYoshino-Kumano National Park. This particular area of Honshū has been described as one of the rainiest spots in Japan.
Eastern Himalayan broadleaf forests (Bhutan, India, Nepal)
TheRussian Far East region is the eastern-most region of both Russia and the Asian continent as a whole. The Russian federal subjects ofPrimorsky Krai andKhabarovsk Krai are located in the southeast of this region, with Primorsky Krai sharing a land border withChina andNorth Korea, and both federal subjects face the Pacific ocean to the east and share maritime borders with Japan. TheSikhote-Alin mountain range is located here and extends for about 1000 km in a northeast direction, parallel to the coast, from near the coastal city ofVladivostok.[54]
Whilst the mountain range ascends from sea level to a maximum altitude of around 1900 metres contains a variety of different habitats, they are located in region with a temperate climate. During the lastglacial maximum (or ice age), the area was not glaciated, allowing for the development of a complexecosystem containing species with origins in Siberia’s boreal forest and Manchuria’s subtropical forests.[55] Temperate rainforest covers most of the mountain slopes and the biogeographic region is known as the Primorye centre of plant diversity, a biogeographic meeting point of flora and fauna from temperate, subtropical and taiga climatic regions.[54] Historically these forests ranged from the southeastern Pacific coast of Russia, through North Korea and into northern China, however vast human development, particularly in China, has limited the forest to its current range in the Russian Far East.[56] In 2001,UNESCO recognized a 1.5 million hectare area of forest in the central part of the Sikhote-Alin mountains as aWorld Heritage Site in Russia, citing the area as one of the most unique and valuable areas of intact forest in the world[57]
Along with the neighbouringAmur region of Russia, the temperate rainforests of the Russian Far East hold the last remaining habitats for the critically endangeredSiberian tiger,Amur leopard, andManchurian sika deer. It has been estimated that there are less than 600 tigers[61] and around 90 leopards left in the wild.[62] The area also contains populations ofAsiatic black bears,Kamchatka brown bears, andMongolian grey wolves, as the Russian Far East, altogether, might probably be the only place in the world where endangered tigers, leopards, bears, and grey wolves coexist. This region also happens to be some of the last of habitat of theBlakiston’s fish owl (Bubo blakistoni); along with being the world’s largest owl, it is unique in the way that it eats fish (primarilyMasu salmon) and relies onold growth forests along river banks to hunt, nest, and breed.[63] TheSiberian grouse is similar to thespruce grouse andFranklin's grouse of North America, and can be found in the dense, remote pockets of broadleaf, coniferous and deciduous forests of Far East Russia. Common ungulates includered deer,roe deer,wild boar, Manchurian moose, andmusk deer.
InAustralia rainforests occur near the mainland east coast and inTasmania. There are warm-temperate and cool-temperate rainforests. They arebroadleafevergreen forests with the exception ofmontane rainforests of Tasmania.Eucalypt forests are not classified as rainforests although some eucalypt forest types receive high annualrainfall (to over 2000 mm in Tasmania[64]), and in the absence offire they may develop to rainforest. If these widespreadwet sclerophyll forests were considered rainforests, the total area of rainforest in Australia would be much larger.[65]
The temperate rainforests of New Zealand occur on the western shore of theSouth Island and on theNorth Island. The forests are made up of coniferouspodocarps and broadleaf evergreen trees. The podocarps are abundant at lower elevations, whilesouthern beech (Nothofagus) can be found on higher slopes and in the cooler southernmost rainforests. Ecoregions include theFiordland temperate forests andWestland temperate rainforests.
^Floyd, A. (1990)Australian Rainforests in New South Wales, Vol. 1. Surrey Beatty & Sons Pty Ltd, Chipping Norton, NSW,ISBN0949324302.
^Reilly, Matthew & Spies, Thomas. (2015). Regional variation in stand structure and development in forests of Oregon, Washington, and inland Northern California. Ecosphere. 6. art192. 10.1890/ES14-00469.1.
^abcElias, Rui Bento; Dias, Eduardo (2008).Ecologia das Florestas de Juniperus dos Açores. Angra do Heroísmo: Cadernos de Botância nº5.ISBN978-989-630-978-7.
^Fernández-Palacios, José María; Otto, Rüdiger; Capelo, Jorge; Caujapé-Castells, Juli; de Nascimento, Lea; Duarte, Maria Cristina; Elias, Rui B.; García-Verdugo, Carlos; Menezes de Sequeira, Miguel; Médail, Frédéric; Naranjo-Cigala, Agustín; Patiño, Jairo; Price, Jonathan; Romeiras, Maria M.; Sánchez-Pinto, Lázaro; Whittaker, Robert J. (December 2024)."In defence of the entity of Macaronesia as a biogeographical region".Biological Reviews.99 (6):2060–2081.doi:10.1111/brv.13112.hdl:10451/65343.ISSN1464-7931.PMID38888215. Retrieved24 November 2024.
^Zazanashvili N, Sanadiradze G, Bukhnikashvili A, Kandaurov A, Tarkhnishvili D. (2004). "Caucasus", pp. 148–153 in Mittermaier RA, Gil PG, Hoffmann M, Pilgrim J, Brooks T, Mittermaier CG, Lamoreux J, da Fonseca GAB (eds.)Hotspots revisited, Earth's biologically richest and most endangered terrestrial ecoregions. Sierra Madre: CEMEX/Agrupacion Sierra Madre.
^van Zeist W, Bottema S. (1991).Late Quaternary vegetation of the Near East. Weisbaden: Reichert,ISBN3882265302.
^Kikvidze Z, Ohsawa M. (1999) "Adjara, East Mediterranean refuge of Tertiary vegetation", pp. 297–315 in: Ohsawa M, Wildpret W, Arco MD (eds.) Anaga Cloud Forest, a comparative study on evergreen broad-leaved forests and trees of the Canary Islands and Japan. Chiba: Chiba University Publications.
^Tarkhnishvili, David N.; Thorpe, Roger S.; Arntzen, Jan Willem (2000). "Pre-Pleistocene refugia and differentiation between populations of the Caucasian salamander (Mertensiella caucasica)".Molecular Phylogenetics and Evolution.14 (3):414–422.Bibcode:2000MolPE..14..414T.doi:10.1006/mpev.1999.0718.PMID10712846.
^abSatoo, T. (1983).Temperate broad-leaved evergreen forests of Japan. In: Ovington, J.V. (ed.)Ecosystems of the world 10: Temperate broad-leaved evergreen forests, pp. 169–189. Elsevier, Amsterdam
^Ching, K.K. (1991).Temperate deciduous forests in East Asia. In: Röhrig, E. & Ulrich, B. (eds.)Ecosystems of the world 7: Temperate deciduous forests, pp. 539–556. Elsevier, Amsterdam
^abcCushman, S. A., & Wallin, D. O. (2002). Separating the effects of environmental, spatial and disturbance factors on forest community structure in the Russian Far East. Forest Ecology and Management, 168(1), 201-215
^DellaSala D. 2011. Temperate and Boreal Rainforests of the World. Island Press
^Carroll, C., & Miquelle, D. G. (2006). Spatial viability analysis of Amur tiger Panthera tigris altaica in the Russian Far East: the role of protected areas and landscape matrix in population persistence. Journal of Applied Ecology, 43(6), 1056-1068
^Uphyrkina, O., Miquelle, D., Quigley, H., Driscoll, C., & O'Brien, S. J. (2002). Conservation genetics of the Far Eastern leopard (Panthera pardus orientalis). Journal of Heredity, 93(5), 303-311