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Shingopana

From Wikipedia, the free encyclopedia
Extinct genus of reptiles

Shingopana
Temporal range: lateCampanian-earlyMaastrichtian
~74.3–71.4 Ma[1]
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Dinosauria
Clade:Saurischia
Clade:Sauropodomorpha
Clade:Sauropoda
Clade:Macronaria
Clade:Titanosauria
Clade:Lithostrotia
Clade:Aeolosaurini
Genus:Shingopana
Gorscaket al.2017
Type species
Shingopana songwensis
Gorscaket al. 2017

Shingopana (meaning "wide neck" inSwahili) is agenus oftitanosauriansauropod from theUpper Cretaceous (lateCampanian-earlyMaastrichtian[2] age)Galula Formation ofTanzania. It is known from only thetype species,S. songwensis.[3] Gorscak & O'Connor's phylogenetic testing suggestShingopana is more closely related to the South American titanosaur family ofAeolosaurini than any of the titanosaurs found so far in North & South Africa.[3]

Discovery and naming

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Part of theholotype, TZ-07, was discovered in 2002 by scientists affiliated with the Rukwa Rift Basin Project, which was run by Patrick O'Connor and Nancy Stevens.[4] The rest of the skeleton was excavated during the following years. The speciesShingopana songwensis was officially named in 2017.[3]

Description

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Shingopanawas a quadrupedalAeolosaurin sauropod that would have reached up to 8 metres (26 ft) long when fully grown, smaller than the average sauropod.[3]

Skeleton

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The holotype was damaged by insect bore holes shortly after the animal died.[3]

Shingopana is known from a partial jaw, represented by the angular bone.Shingopana is also known from four cervical vertebrae; with two of these vertebrae having preserved cervical ribs and another isolated cervical rib.Shingopana instead had remnants of a bulbous expansion on the incompletely preserved cervical vertebrae, which probably helped to strengthen its neck.[3]

Four ribs have been preserved with the holotype, but none are complete. The ribs had flanged edges, but their function is currently unknown.[3]

An almost complete humerus and a partial pubis were also present in the holotype.

Palaeoecology

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The holotype was discovered in theUpper CretaceousGalula Formation of theRukwa Rift Basin in Tanzania. It would have coexisted with the sauropodsRukwatitan[5] andMnyamawamtuka,[6]themesoeucrocodilesPakasuchus[7] andRukwasuchus,[8] the mammalGalulatherium,[5] an unnamednotosuchian, an unnamedturtle, an unnamedtheropod[5] and two types oflungfish (Lupaceradotus and an unnamed genus).

See also

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References

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  1. ^Ogg, J.G. (2012), "Geomagnetic Polarity Time Scale",The Geologic Time Scale, Elsevier, pp. 85–113,doi:10.1016/b978-0-444-59425-9.00005-6,ISBN 978-0-444-59425-9,S2CID 127887632{{citation}}: CS1 maint: work parameter with ISBN (link)
  2. ^Widlansky, Sarah J.; Clyde, William C.; O'Connor, Patrick M.; Roberts, Eric M.; Stevens, Nancy J. (2018-03-01)."Paleomagnetism of the Cretaceous Galula Formation and implications for vertebrate evolution".Journal of African Earth Sciences.139:403–420.Bibcode:2018JAfES.139..403W.doi:10.1016/j.jafrearsci.2017.11.029.ISSN 1464-343X.
  3. ^abcdefgGorscak, E.; O'Connor, P. M.; Roberts, E. M.; Stevens, N. J. (2017)."The second titanosaurian (Dinosauria: Sauropoda) from the middle Cretaceous Galula Formation, southwestern Tanzania, with remarks on African titanosaurian diversity".Journal of Vertebrate Paleontology.361 (4):35–55.Bibcode:2017JVPal..37E3250G.doi:10.1080/02724634.2017.1343250.S2CID 90885040.
  4. ^Stevens, N.J., Gottfried, M.D., Roberts, E.M., Kapilima, S., Ngasala, S. and O'Connor, P.M. (2007).Paleontological exploration in Africa: A view from the Rukwa Rift Basin of Tanzania. Pp. 159–180 in Fleagle, J.G. and Gilbert, C.C. (eds.). Elwyn Simons: a search for origins. Springer, 460 pp.ISBN 978-0-387-73895-6
  5. ^abcO'Connor, Patrick M.; Gottfried, Michael D.; Stevens, Nancy J.; Roberts, Eric M.; Ngasala, Sifa; Kapilima, Saidi; Chami, Remigius (March 2006). "A new vertebrate fauna from the Cretaceous Red Sandstone Group, Rukwa Rift Basin, Southwestern Tanzania".Journal of African Earth Sciences.44 (3):277–288.Bibcode:2006JAfES..44..277O.doi:10.1016/j.jafrearsci.2005.11.022.ISSN 1464-343X.
  6. ^Gorscak, Eric; O'Connor, Patrick M. (2019)."A new African titanosaurian sauropod dinosaur from the middle Cretaceous Galula Formation (Mtuka Member), Rukwa Rift Basin, southwestern Tanzania".PLoS ONE.2 (14) e0211412.Bibcode:2019PLoSO..1411412G.doi:10.1371/journal.pone.0211412.PMC 6374010.PMID 30759122.
  7. ^O'Connor, Patrick M.; Sertich, Joseph J. W.; Stevens, Nancy J.; Roberts, Eric M.; Gottfried, Michael D.; Hieronymus, Tobin L.; Jinnah, Zubair A.; Ridgely, Ryan; Ngasala, Sifa E. (August 2010). "The evolution of mammal-like crocodyliforms in the Cretaceous Period of Gondwana".Nature.466 (7307):748–751.Bibcode:2010Natur.466..748O.doi:10.1038/nature09061.ISSN 0028-0836.PMID 20686573.S2CID 4405106.
  8. ^Sertich, Joseph J. W.; O'Connor, Patrick M. (2014-04-16). "A new crocodyliform from the middle Cretaceous Galula Formation, southwestern Tanzania".Journal of Vertebrate Paleontology.34 (3):576–596.Bibcode:2014JVPal..34..576S.doi:10.1080/02724634.2013.819808.ISSN 0272-4634.S2CID 16644660.
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Riojasauridae
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Topics in sauropodomorph research
Shingopana
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