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Orodromeus

From Wikipedia, the free encyclopedia
Extinct genus of dinosaurs

Orodromeus
Temporal range:Late Cretaceous,76.7 Ma
Orodromeus makelai
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Clade:Dinosauria
Clade:Ornithischia
Family:Thescelosauridae
Subfamily:Orodrominae
Genus:Orodromeus
Horner &Weishampel,1988
Species:
O. makelai
Binomial name
Orodromeus makelai
Horner & Weishampel, 1988
Reconstruction of theOrodromeus skeleton

Orodromeus (meaning "Mountain Runner") is agenus ofherbivorousorodrominethescelosauriddinosaur from theLate Cretaceous ofNorth America. Only one species is known, thetype speciesOrodromeus makelai.

Discovery and naming

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Size of known individuals ofOrodromeus (left) compared to its relativesThescelosaurus (right) andParksosaurus (center), as well as a human

The remains ofOrodromeus were discovered byRobert Makela during the excavation inTeton County,Montana, of theEgg Mountain brooding colony of a much larger relative,Maiasaura. Thetype species,Orodromeus makelai andOrodromeus niedae, were named and shortly described byJack Horner andDavid B. Weishampel in1988. The generic name is derived from Greek ὄρος,oros, "mountain", in reference to the Egg Mountain site, and δρομεύς,dromeus, "runner", referring to the cursorial habits of the animal. Thespecific name honoured the late Makela.[1]

Theholotype specimen, MOR 294, was found in a layer of theTwo Medicine Formation, dating from theCampanian stage, about 75million years ago. It consists of a partial skeleton with skull. Theparatypes are MOR 246, aclutch of nineteeneggs, some withembryos; PP 22412, a set ofhindlimbs; MOR 331, a partial skeleton; MOR 248, a skeleton with skull; and MOR 403, a braincase.[1] A full publisheddescription is still lacking, though an unpublished thesis onOrodromeus exists.[2] However, MOR 246 and other eggs from Egg Mountain are now considered to belong to atroodontid[3] which may beStenonychosaurus.[4]

Description

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Orodromeus was a small fastbipedal herbivore that probably coexisted with dinosaurs such asDaspletosaurus andEiniosaurus. Its length was estimated by Horner & Weishampel at 2.5 metres.[1]

Skull and partial vertebral column ofOrodromeus MOR 294

Orodromeus is distinguished by apalpebral that is at its back attached to thepostorbital; a boss on thejugal; a non-fused wrist; and triangularmaxillary anddentary teeth with a superficial flatocclusion.[1]

Phylogeny

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Reconstructed pelvis ofOrodromeus.
Model nest and hatchlings ofOrodromeus makelai

Orodromeus was by Horner & Weishampel assigned to theHypsilophodontidae, as the youngest known member.[1] Today these are seen as an unnatural,paraphyletic, group, andOrodromeus is simply considered to be a basal member of theEuornithopoda.Brownet al. (2013[5]) put it in the familyThescelosauridae and named a new subfamily (Orodrominae) after it.

Palaeobiology

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Because of the advanced development of the bones and teeth of the embryos, Horner concluded that the young ofOrodromeus wereprecocial.

It has been speculated that this animal may haveburrowed much like its relativeOryctodromeus, based upon the packing of their bones in situations where they typically would have been scattered.[6]

Mallon et al. (2013) examined herbivorecoexistence on the island continent ofLaramidia, during the Late Cretaceous. It was concluded that smallornithischians likeOrodromeus were generally restricted to feeding on vegetation at, or below the height of 1 meter.[7]

References

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  1. ^abcdeHorner, J. and Weishampel, D., 1988, "A comparative embryological study of two ornithischian dinosaurs",Nature (London),332(No. 6161): 256-257 (1988)
  2. ^Scheetz, R.D., 1999,Osteology ofOrodromeus makelai and the phylogeny of basal ornithopod dinosaurs D. Ph. Thesis in Biology, Montana State University, Bozeman, 189 pp
  3. ^Varricchio, D.J.; Jackson, F.; Borkowski, J.J.; Horner, J.R. (1997). "Nest and egg clutches of the dinosaurTroodon formosus and the evolution of avian reproductive traits".Nature.385 (6613):247–250.Bibcode:1997Natur.385..247V.doi:10.1038/385247a0.S2CID 4313286.
  4. ^van der Reest, A. J.; Currie, P. J. (2017). "Troodontids (Theropoda) from the Dinosaur Park Formation, Alberta, with a description of a unique new taxon: implications for deinonychosaur diversity in North America".Canadian Journal of Earth Sciences.54 (9):919–935.Bibcode:2017CaJES..54..919V.doi:10.1139/cjes-2017-0031.hdl:1807/78296.
  5. ^Brown; et al. (2013). "New data on the diversity and abundance of small-bodied ornithopods (Dinosauria, Ornithischia) from the Belly River Group (Campanian) of Alberta".Journal of Vertebrate Paleontology.33 (3):495–520.Bibcode:2013JVPal..33..495B.doi:10.1080/02724634.2013.746229.S2CID 129160518.
  6. ^Varricchio, David J.; Martin, Anthony J.; Katsura, Yoshihiro (2007)."First trace and body fossil evidence of a burrowing, denning dinosaur".Proceedings of the Royal Society B: Biological Sciences.274 (1616):1361–1368.doi:10.1098/rspb.2006.0443.PMC 2176205.PMID 17374596.
  7. ^Mallon, Jordan C; David C Evans; Michael J Ryan; Jason S Anderson (2013)."Feeding height stratification among the herbivorous dinosaurs from the Dinosaur Park Formation (upper Campanian) of Alberta, Canada".BMC Ecology.13 (1): 14.Bibcode:2013BMCE...13...14M.doi:10.1186/1472-6785-13-14.PMC 3637170.PMID 23557203.
Avemetatarsalia
Ornithischia
    • see below↓
Heterodontosauridae
Thyreophora
Ankylosauria
Stegosauria
Thescelosauridae
Ornithopoda
Marginocephalia
Ceratopsia
Pachycephalosauria
Laquintasaura venezuelae

Heterodontosaurus tuckiScutellosaurus lawleriKulindadromeus zabaikalicus

Thescelosaurus neglectus
See also
Nomina dubia
Incertae sedis
Other taxa
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Orodromeus
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