Xotodon sp. skull at the Museum für Naturkunde, Berlin
Toxodontidae is anextinctfamily ofnotoungulate mammals, known from theOligocene to theHolocene (11,000 BP) of South America,[1] with one genus,Mixotoxodon, also known from thePleistocene of Central America and southern North America (as far north as Texas).[2] Member of the family were medium to large-sized,[3] ranging from around 350–400 kilograms (770–880 lb) inNesodon to 1,000–1,200 kilograms (2,200–2,600 lb) inToxodon,[4] and had medium to high-crowned dentition, which in derived members of the group evolved into ever-growing cheek teeth.[5] Isotopic analyses have led to the conclusion that Pleistocene members of the family were flexible mixed feeders (bothbrowsing andgrazing).[6][7]
The endemic notoungulate andlitoptern ungulates of South America have been shown by studies of collagen and mitochondrial DNA sequences to be asister group to theperissodactyls.[8][9][10]
In 2014, a study identifying a new species of toxodontid resolved the families' phylogenetic relations. The below cladogram was found by the study:[11]
^E. Lundelius, et al. 2013. The first occurrence of a toxodont (Mammalia, Notoungulata) in the United States.Journal of Vertebrate Paleontology, Vol 33, No 1, pp. 229–232DOI:10.1080/02724634.2012.711405
^MacFadden, Bruce J. (September 2005). "Diet and habitat of toxodont megaherbivores (Mammalia, Notoungulata) from the late Quaternary of South and Central America".Quaternary Research.64 (2):113–124.Bibcode:2005QuRes..64..113M.doi:10.1016/j.yqres.2005.05.003.
McKenna, Malcolm C., and Bell, Susan K. 1997.Classification of Mammals Above the Species Level. Columbia University Press, New York, 631 pp. ISBN0-231-11013-8