| Mountain goat | |
|---|---|
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| Mountain goat in theCascades | |
Scientific classification | |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Mammalia |
| Order: | Artiodactyla |
| Family: | Bovidae |
| Subfamily: | Caprinae |
| Genus: | Oreamnos |
| Species: | O. americanus |
| Binomial name | |
| Oreamnos americanus (Blainville, 1816) | |
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| Synonyms | |
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Themountain goat (Oreamnos americanus), also known as theRocky Mountain goat, is acloven-footed mammal that isendemic to the remote and rugged mountainous areas of westernNorth America. Asubalpine to trulyalpine species, it is asure-footed climber commonly seen on sheer rock faces, near-vertical cliffs and icy passages. Mountain goats generally avoid venturing down into lower elevations—except during seasonal food shortages or during particularly bad weather—as the extreme elevation which they inhabit is their primary defense against predators such asblack andbrown bears,pumas andwolves.
Despite its vernacular name and both genera being in the same subfamily (Caprinae), the mountain goat is not a member ofCapra, the genus that includes all truegoats (such as thewild goat (Capra aegagrus), from which the domestic goat is derived); rather, it is more closely allied with the otherbovids known as "goat-antelopes", including the Europeanchamois (Rupicapra), thegorals (Naemorhedus), thetakins (Budorcas) and theserows (Capricornis), of Japan and eastern South Asia.
The mountain goat is an even-toedungulate of the orderArtiodactyla and the familyBovidae (along withantelopes,gazelles, andcattle). It belongs to the subfamilyCaprinae, along withtrue goats,wild sheep, thechamois, themuskox and other species. Thetakins of theHimalayan region, while not asister lineage of the mountain goat, are nonetheless very closely related and almostcoeval to the mountain goat; theyevolved in parallel from an ancestral goat. Other members of this group are thebharal, the true goats, and theHimalayan tahr. The sheep lineage is also very closely related, while the muskox lineage is somewhat more distant. The mountain goats probably diverged from their relatives in the lateTortonian, some 7.5 to 8 million years ago.[citation needed]
Given that all major caprine lineages emerged in theLate Miocene and contain at least one but usually several species from the eastern Himalayan region, their most likely place of origin is between today'sTibet andMongolia or nearby. The mountain goat's ancestors thus probably crossed theBering Strait after they split from their relatives, presumably before theWisconsinian glaciation. NoPliocene mountain goats have been identified yet; the knownfossil record is fairly recent, entirely from North America, and barely differs from the living animals. In thePleistocene era, the small prehistoric mountain goatOreamnos harringtoni lived in the southernRocky Mountains.Ancient DNA studies suggest that this was thesister species of the living mountain goat, not its ancestor; consequently, the living species would also date back to the Pleistocene at least.[2][3]
The mountain goat is theonly living species in the genusOreamnos. The nameOreamnos is derived from the Greek termὄροςóros (stemore-) meaning "mountain" (or, alternatively,oreas "mountain nymph") and the wordἀμνόςamnós meaning "lamb".
Both male and female mountain goats have beards, short tails, and long black horns, 15–28 cm (5.9–11 in) in length, which contain yearly growth rings. They are protected from the elements by their woolly greyish white double coats. The fine, dense wool of their undercoats is covered by an outer layer of longer, hollow hairs. Mountain goats molt in spring by rubbing against rocks and trees, with the adult billies shedding their extra wool first and the pregnant nannies shedding last. Their coats help them to withstand winter temperatures as low as −46 °C (−51 °F) and winds of up to 160 kilometres per hour (99 mph).
A male goat stands about 1 m (3.3 ft) at the shoulder to the waist and can weigh considerably more than the female (around 30% more in some cases). Male goats also have longer horns and longer beards than females. The head-and-body length can range from 120–179 cm (47–70 in), with a small tail adding 10–20 cm (3.9–7.9 in).[4][5][6]
The mountain goat's feet are well-suited for climbing steep, rocky slopes with pitches exceeding 60°, with inner pads that provide traction andcloven hooves that can spread apart. The tips of their feet have sharpdewclaws that keep them from slipping. They have powerful shoulder and neck muscles that help propel them up steep slopes.[7] Based on a field recording in the Rocky Mountains of Canada of a mountain goat climbing a 45-degree slope, researchers were able to measure the goat's whole body movement as it climbed. Researchers observed that when the goat propelled itself forward, it extended its back legs and the front legs were tucked close up to its chest during its first phase. During the second phase, the goat raised its back legs near to its chest, while the front leg's humerus stayed locked in a persistent location relative to the goat's chest, therefore allowing the elbow to be detained in close proximity to the whole body's center of balance. Extension of the elbow and carpal joints resulted in a vertical translation of the center of mass up the mountain slope.[8]
The mountain goat inhabits theRocky Mountains andCascade Range and other mountain regions of theWestern Cordillera of North America, fromWashington,Idaho andMontana throughBritish Columbia andAlberta, into the southernYukon and southeasternAlaska. British Columbia contains half of the world's population of mountain goats.[9] Its northernmost range is said to be along the northern fringe of theChugach Mountains in south-central Alaska. Introduced populations can also be found in such areas asIdaho,Wyoming,Utah,Nevada,Oregon,Colorado,South Dakota, and theOlympic Peninsula of Washington.
Mountain goats are the largest mammals found in their high-altitudehabitats, which can exceed elevations of 13,000 ft (4,000 m). They sometimes descend to sea level in coastal areas although they are primarily analpine andsubalpine species. The animals usually stay above thetree line throughout the year but they will migrate seasonally to higher or lower elevations within that range. Winter migrations to low-elevationmineral licks often take them several kilometers through forested areas.[10]
Daily movements by individual mountain goats are primarily confined to areas on the same mountain face,drainage basin, or alpine opening. Daily movements reflect an individual's needs for foraging, resting,thermoregulation and security from predators or disturbance. Seasonal movements primarily reflect nutritional needs (such as movements to and frommineral licks/salt lick), reproductive needs (in other words, movement of pre-parturient females to "kidding" areas; movement to rutting areas), and climatic influences (including movement to areas in response to foraging conditions). In general, seasonal movements are likely to exhibit a strong elevational component, whereby lower, forested elevations are used during the spring-summer (security cover effects) to access lower elevation mineral licks, and during winter (thermal cover effects) to access forage. The farthest movements are expected to be by dispersing mountain goats. Such movements are likely to involve mountain goats crossing forested valleys as they move between mountain blocks.
Mountain goats are herbivores and spend most of their time grazing. Their diets includegrasses,herbs,sedges,ferns,mosses,lichens, and twigs and leaves from the low-growing shrubs andconifers of their high-altitude habitat.
In captivity, the mountain goat's diet can also include grain,alfalfa, fruits, vegetables andgrass.
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In the wild, mountain goats usually live 12 to 15 years, with their lifespans limited by the wearing down of their teeth. In zoos, however, they can live for 16-20 years.
Mountain goats reach sexual maturity at about 30 months.[11] Nannies in a herd undergo synchronizedestrus in late October through early December, at which time females and males participate in a mating ritual. Mature billies stare at nannies for long periods, dig rutting pits, and fight each other in showy (though occasionally dangerous) scuffles. Nannies often ignore young billies, who try to participate but are discounted in favor of older partners. Both females and males usually mate with multiple individuals during breeding season, although some billies try to keep other males away from certain nannies. After the breeding season is over, females and males move away from each other. Nannies form loose-knit nursery groups of up to 50 animals. The adult billies leave, often alone or with two or three other billies.
Kids are born in the spring (late May or early June) after a six-month gestation period. Nannies give birth, usually to a single offspring, after moving to an isolated ledge;post partum, they lick the kid dry and ingest theplacenta. Kids weigh a little over 3 kg (6.6 lb) at birth and begin to run and climb (or attempt to do so) within hours. Although lactation is mostly finished at one month, kids follow their mothers closely for the first year of life (or until the nanny gives birth again, if this does not occur the next breeding season); nannies protect their young by leading them out of danger, standing over them when faced by predators, and positioning themselves below their kids on steep slopes to stop freefalls.
Nannies can be very competitive and protective of their space and food sources. They fight with one another for dominance in conflicts that can ultimately include all the nannies in the herd. In these battles, nannies circle each other with their heads lowered, displaying their horns. These conflicts can occasionally lead to injury or death, but are usually harmless. To avoid fighting, an animal may show a posture of nonaggression by stretching low to the ground.
In regions below the tree line, nannies use their fighting abilities to protect themselves and their offspring from predators. Predators, includingwolves,wolverines,lynxes, andbears, attack goats of most ages given the opportunity. Thecougar, or mountain lion, is perhaps the primary predator, being powerful enough to overwhelm the largest adults and uniquely nimble enough to navigate the rocky ecosystem of the goats. Though their size protects them from most potential predators in higher altitudes, nannies must sometimes defend their young from bothbald andgolden eagles,[12] which can be a predatory threat to kids.[5] Nannies have even been observed trying to dominate the more passive, but often heavierbighorn sheep that share some of their territory. In 2021, a mountain goat gored agrizzly bear to death inYoho National Park,British Columbia.[13]
Mountain goats introduced in the 1920s into Washington'sOlympic Mountains were in time found to be a nuisance there, in particular while seeking human urine and sweat for its salt content, the park lacking naturalsalt licks, and even aggressively approaching human visitors.[14] One such goat killed a hiker in 2010.[15] Officials finally chose to eradicate them from theOlympic Peninsula, removing hundreds, mostly by capturing them and relocating them to theCascade Mountains.[16]
Although mountain goats have never beendomesticated andcommercialized for theirwool,pre-Columbianindigenous peoples of the Pacific Northwest Coast did incorporate their wool into theirweaving by collecting springmoulted wool left by wild goats.[17]
Avalanches can constitute 23-65% of Alaskan mountain goat mortality depending on the region.[18]
