Mary Eileen McDonnell was born April 28, 1952,[1][2][3] inWilkes-Barre, Pennsylvania, one of six children born to Eileen (née Mundy) and John "Jack" McDonnell, a computer consultant.[4] She is of Irish descent[5] and was raisedRoman Catholic.[6] As a child, McDonnell relocated with her family toIthaca, New York, where she spent the remainder of her upbringing.[6]
After more than 21 years of theater and television work, McDonnell made her film breakthrough in 1990 as the daughter of American settlers raised bySioux Indians named Stands With A Fist inKevin Costner'sDances With Wolves. Portraying the adopted daughter ofGraham Greene's character Kicking Bird, McDonnell, then 37, was actually two months older than Greene and less than two years younger thanTantoo Cardinal, who played Black Shawl, her adoptive mother. She was nominated for anAcademy Award for Best Supporting Actress for the role.[11]
On television, McDonnell had her first regular part in 1980 on the soap operaAs the World Turns. She starred in 1984 on the short-lived CBS medical comedyE/R. Coincidentally, she guest-starred in 2001 on the NBC medical series of the same nameER. She was nominated for anEmmy Award for her role on the show as Eleanor Carter. She played Dr. Virginia Dixon, a surgeon withAsperger syndrome for three episodes ofGrey's Anatomy in 2008 and 2009.[13]
In 2003, McDonnell starred in the miniseriesBattlestar Galactica asLaura Roslin. The miniseries led to theweekly series, with McDonnell's reprising her Laura Roslin role. The series ended in March 2009. McDonnell received worldwide recognition for her performance in the show, part of which was shown when she was invited to the United Nations for a retrospective and discussion withEdward James Olmos (Admiral Adama).
In 2011, she appeared in the role of Kate Roberts, the mother ofEmma Roberts's character inScream 4.[15]
From 2009 to 2012, McDonnell had a recurring role inThe Closer as Capt. Sharon Raydor, a police captain in theForce Investigation Division, who often butts heads withKyra Sedgwick's Golden Globe and Emmy award-winning character.[16] McDonnell received an Emmy nomination for Outstanding Guest Actress in a Drama Series in 2011 for the role.[17]
AfterThe Closer wrapped up its final season 2012, McDonnell's character continued as the lead in the spin-off,Major Crimes, which debuted August 13, 2012 and ended in January 2018.[18]