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List of subgroups of the order Coleoptera

From Wikipedia, the free encyclopedia

This articleclassifies thesubgroups of the order Coleoptera (beetles) down to the level offamilies, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020,^[1] with common names from bugguide.net.^[2]^[3]^[4]

OrderColeoptera
- Suborder †Protocoleoptera
  - Superfamily †TshekardocoleoideaRohdendorf, 1944
    - Family †TshekardocoleidaeRohdendorf, 1944
    - Family †LabradorocoleidaePonomarenko, 1969
    - Family †OborocoleidaeKukalová, 1969
  - Superfamily †PermocupedoideaMartynov, 1933
    - Family †PermocupedidaeMartynov, 1933
    - Family †TaldycupedidaeRohdendorf, 1961
  - Superfamily †PermosynoideaTillyard, 1924
    - Family †AdemosynidaePonomarenko, 1968
    - Family †PermosynidaeTillyard, 1924
- SuborderArchostemata
  - SuperfamilyCupedoideaLaporte, 1836
    - FamilyCrowsoniellidaeIablokoff-Khnzorian, 1983
    - FamilyCupedidaeLaporte, 1836
    - FamilyMicromalthidaeBarber, 1913
      Micromalthus debilis
    - FamilyOmmatidaeSharp and Muir, 1912
    - FamilyJurodidaePonomarenko, 1985
    - Family †TriadocupedidaePonomarenko, 1966
    - Family †MagnocoleidaeHong, 1998
    - Family †ObrieniidaeZherikhin and Gratshev, 1994
  - Superfamily †AsiocoleoideaRohdendorf, 1961
    - Family †AsiocoleidaeRohdendorf, 1961
    - Family †TricoleidaePonomarenko, 1969
  - Superfamily †RhombocoleoideaRohdendorf, 1961
    - Family †RhombocoleidaeRohdendorf, 1961
  - Superfamily †SchizocoleoideaRohdendorf, 1961 (formerly SchizophoroideaPonomarenko, 1968)
    - Family †PhoroschizidaeBouchard and Bousquet, 2020 (formerly SchizophoridaePonomarenko, 1968)
    - Family †CatiniidaePonomarenko, 1968
    - Family †SchizocoleidaeRohdendorf, 1961
- SuborderMyxophaga
  - SuperfamilyLepiceroideaHinton, 1936 (1882)
    - FamilyLepiceridaeHinton, 1936 (1882)
  - SuperfamilySphaeriusoideaErichson, 1845
    - FamilyTorridincolidaeSteffan, 1964
    - FamilyHydroscaphidaeLeConte, 1874
    - FamilySphaeriusidaeErichson, 1845
- SuborderAdephaga
  - - Family †TritarsusidaeHong, 2002 (formerly TritarsidaeHong, 2002)
    - FamilyGyrinidaeLatreille, 1810 (whirligig beetles)
    - FamilyTrachypachidaeThomson, 1857 (false ground beetles)
    - FamilyRhysodidaeLaporte, 1840
    - FamilyCarabidaeLatreille, 1802 (ground beetles)
      Harpalus, in theCarabidae
    - FamilyHaliplidaeAubé, 1836 (crawling water beetles)
    - Family †TriaplidaePonomarenko, 1977
    - Family †ColymbotethidaePonomarenko, 1994
    - Family †ParahygrobiidaePonomarenko, 1977
    - Family †CoptoclavidaePonomarenko, 1961
    - Family †LiadytidaePonomarenko, 1977
    - FamilyMeruidaeSpangler and Steiner, 2005
    - FamilyNoteridaeThomson, 1860 (burrowing water beetles)
    - FamilyAmphizoidaeLeConte, 1853
    - FamilyAspidytidaeRibera, Beutel, Balke and Vogler, 2002
    - FamilyHygrobiidaeRégimbart, 1879 (1837)
    - FamilyDytiscidaeLeach, 1815 (predaceous diving beetles)
      Dytiscus latissimus, in theDytiscidae
- SuborderPolyphaga
  - InfraorderStaphyliniformia
    - SuperfamilyHisteroideaGyllenhal, 1808^[5]
      - FamilyHisteridaeGyllenhal, 1808
      - FamilySphaeritidaeShuckard, 1839
      - FamilySynteliidaeLewis, 1882
    - SuperfamilyHydrophiloideaLatreille, 1802^[6]
      - FamilyEpimetopidaeZaitzev, 1908
      - FamilyGeorissidaeLaporte, 1840
      - FamilyHelophoridaeLeach, 1815
      - FamilyHydrochidaeThomson, 1859
      - FamilyHydrophilidaeLatreille, 1802 (water scavenger beetles)
      - FamilySpercheidaeErichson, 1837
    - SuperfamilyStaphylinoideaLatreille, 1802
      - FamilyHydraenidaeMulsant, 1844
      - FamilyPtiliidaeErichson, 1845
      - FamilyAgyrtidaeThomson, 1859
      - FamilyLeiodidaeFleming, 1821
      - FamilySilphidaeLatreille, 1806
      - FamilyStaphylinidaeLatreille, 1802 (rove beetles)
        Devil's coach horse beetle,Ocypus olens, arove beetle
  - InfraorderScarabaeiformia
    - SuperfamilyScarabaeoideaLatreille, 1802
      - FamilyPleocomidaeLeConte, 1861 (rain beetles)
      - FamilyBolboceratidaeMulsant, 1842
      - FamilyGeotrupidaeLatreille, 1802 (earth-boring scarab beetles)
      - FamilyBelohinidaePaulian, 1959
      - FamilyPassalidaeLeach, 1815 (bess beetles)
      - FamilyTrogidaeMacLeay, 1819 (hide beetles)
      - FamilyGlaresidaePrudhomme de Borre, 1886
      - FamilyDiphyllostomatidaeHolloway, 1972
      - FamilyLucanidaeLatreille, 1804 (stag beetles)
        Lucanus cervus, astag beetle
      - FamilyOchodaeidaeStreubel, 1846 (sand-loving scarab beetles)
      - FamilyHybosoridaeErichson, 1847
      - FamilyGlaphyridaeMacLeay, 1819 (bumble bee scarab beetles)
      - FamilyScarabaeidaeLatreille, 1802 (scarab beetles)
      - Family †CoprinisphaeridaeGenise, 2004 (ichnotaxon)
      - Family †PallichnidaeGenise, 2004 (ichnotaxon)
  - InfraorderElateriformia
    - SuperfamilyScirtoideaFleming, 1821
      - FamilyDecliniidaeNikitsky, Lawrence, Kirejtshuk and Gratshev, 1994
      - FamilyEucinetidaeLacordaire, 1857
      - FamilyClambidaeFischer von Waldheim, 1821
      - FamilyScirtidaeFleming, 1821
      - Family †ElodophthalmidaeKirejtshuk and Azar, 2008
      - Family †MesocinetidaeKirejtshuk and Ponomarenko, 2010
    - SuperfamilyDascilloideaGuérin-Méneville, 1843 (1834)
      - FamilyDascillidaeGuérin-Méneville, 1843 (1834)
      - FamilyRhipiceridaeLatreille, 1834
    - SuperfamilyBuprestoideaLeach, 1815
      - FamilySchizopodidaeLeConte, 1859
      - FamilyBuprestidaeLeach, 1815
        Buprestidae
    - SuperfamilyByrrhoideaLatreille, 1804
      - FamilyByrrhidaeLatreille, 1804
      - FamilyProtelmidaeJeannel, 1950
      - FamilyElmidaeCurtis, 1830
      - FamilyDryopidaeBillberg, 1820 (1817)
      - FamilyLutrochidaeKasap and Crowson, 1975
      - FamilyLimnichidaeErichson, 1846
      - FamilyHeteroceridaeMacLeay, 1825
      - FamilyPsephenidaeLacordaire, 1854
      - FamilyCneoglossidaeChampion, 1897
      - FamilyPtilodactylidaeLaporte, 1836
      - FamilyPodabrocephalidaePic, 1930
      - FamilyChelonariidaeBlanchard, 1845
      - FamilyEulichadidaeCrowson, 1973
      - FamilyCallirhipidaeEmden, 1924
    - SuperfamilyElateroideaLeach, 1815
      - FamilyArtematopodidaeLacordaire, 1857
      - FamilyBrachypsectridaeHorn, 1881
      - FamilyCantharidaeImhoff, 1856 (1815) (soldier beetles)
      - FamilyCerophytidaeLatreille, 1834
      - FamilyElateridaeLeach, 1815 (click beetles; includes former Drilidae, Omalisidae, and Plastoceridae)
        Agrypnus murinus, aclick beetle
      - FamilyEucnemidaeEschscholtz, 1829
      - FamilyLampyridaeRafinesque, 1815 (fireflies)
      - FamilyLycidaeLaporte, 1836
      - FamilyOmethidaeLeConte, 1861 (includes former Telegeusidae)
      - FamilyPhengodidaeLeConte, 1861
      - FamilyRhagophthalmidaeOlivier, 1907
      - FamilySinopyrophoridaeBi, 2018
      - FamilyThroscidaeLaporte, 1840 nomen protectum
      - Family †BerendtimiridaeWinkler, 1987
      - Family †PraelateriidaeDolin, 1973
    - SuperfamilyRhinorhipoideaLawrence, 1988
      - FamilyRhinorhipidaeLawrence, 1988
  - InfraorderBostrichiformia
    - SuperfamilyDerodontoideaLeConte, 1861
      - FamilyDerodontidaeLeConte, 1861
      - FamilyNosodendridaeErichson, 1846
      - FamilyJacobsoniidaeHeller, 1926
    - SuperfamilyBostrichoideaLatreille, 1802
      - FamilyDermestidaeLatreille, 1804 (carpet beetles)
        Anthrenus verbasci, in theDermestidae
      - FamilyEndecatomidaeLeConte, 1861
      - FamilyBostrichidaeLatreille, 1802
      - FamilyPtinidaeLatreille, 1802
  - InfraorderCucujiformia
    - SuperfamilyLymexyloideaFleming, 1821
      - FamilyLymexylidaeFleming, 1821
    - SuperfamilyCleroideaLatreille, 1802
      - FamilyPhloiophilidaeKiesenwetter, 1863
      - FamilyTrogossitidaeLatreille, 1802
      - FamilyChaetosomatidaeCrowson, 1952
      - FamilyMetaxinidaeKolibáč, 2004
      - FamilyThanerocleridaeChapin, 1924
      - FamilyCleridaeLatreille, 1802 (checkered beetles)
      - FamilyAcanthocnemidaeCrowson, 1964
      - FamilyPhycosecidaeCrowson, 1952
      - FamilyPrionoceridaeLacordaire, 1857
      - FamilyMauroniscidaeMajer, 1995
      - FamilyMelyridaeLeach, 1815 (soft-winged flower beetles)
    - SuperfamilyCucujoideaLatreille, 1802
      - Family †ParandrexidaeKirejtshuk, 1994
      - Family †SinisilvanidaeHong, 2002
      - FamilyBoganiidaeSen Gupta and Crowson, 1966
      - FamilyByturidaeGistel, 1848
      - FamilyHelotidaeChapuis, 1876
      - FamilyProtocucujidaeCrowson, 1954
      - FamilySphindidaeJacquelin du Val, 1860
      - FamilyBiphyllidaeLeConte, 1861
      - FamilyErotylidaeLatreille, 1802
      - FamilyMonotomidaeLaporte, 1840
      - FamilyHobartiidaeSen Gupta and Crowson, 1966
      - FamilyCryptophagidaeKirby, 1826
      - FamilyAgapythidaeSen Gupta and Crowson, 1969
      - FamilyPriasilphidaeCrowson, 1973
      - FamilyPhloeostichidaeReitter, 1911
      - FamilySilvanidaeKirby, 1837
      - FamilyCucujidaeLatreille, 1802
      - FamilyMyraboliidaeLawrence and Britton, 1991
      - FamilyCavognathidaeSen Gupta and Crowson, 1966
      - FamilyLamingtoniidaeSen Gupta and Crowson, 1969
      - FamilyPassandridaeBlanchard, 1845
      - FamilyPhalacridaeLeach, 1815
      - FamilyPropalticidaeCrowson, 1952
      - FamilyLaemophloeidaeGanglbauer, 1899
      - FamilyTasmosalpingidaeLawrence and Britton, 1991
      - FamilyCyclaxyridaeGimmel, Leschen and Ślipiński, 2009
      - FamilyKateretidaeKirby, 1837
      - FamilyNitidulidaeLatreille, 1802
      - FamilySmicripidaeHorn, 1880
      - FamilyBothrideridaeErichson, 1845
      - FamilyCerylonidaeBillberg, 1820
      - FamilyAlexiidaeImhoff, 1856
      - FamilyDiscolomatidaeHorn, 1878
      - FamilyEndomychidaeLeach, 1815
      - FamilyCoccinellidaeLatreille, 1807 (ladybirds or lady beetles)
        Asian multicolored lady beetle,Harmonia axyridis
      - FamilyCorylophidaeLeConte, 1852
      - FamilyAkalyptoischiidaeLord, Hartley, Lawrence, McHugh, Whiting and Miller, 2010
      - FamilyLatridiidaeErichson, 1842
    - SuperfamilyTenebrionoideaLatreille, 1802
      Tenebrionoidea -Tenebrio molitor
      - FamilyMycetophagidaeLeach, 1815
      - FamilyArcheocrypticidaeKaszab, 1964
      - FamilyPterogeniidaeCrowson, 1953
      - FamilyCiidaeLeach, 1819
      - FamilyTetratomidaeBillberg, 1820
      - FamilyMelandryidaeLeach, 1815
      - FamilyMordellidaeLatreille, 1802 (tumbling flower beetles)
      - FamilyRipiphoridaeLaporte, 1840
      - FamilyZopheridaeSolier, 1834
      - FamilyUlodidaePascoe, 1869
      - FamilyPromecheilidaeLacordaire, 1859
      - FamilyChalcodryidaeWatt, 1974
      - FamilyTrachelostenidaeLacordaire, 1859
      - FamilyTenebrionidaeLatreille, 1802 (darkling beetles)
        Heliotautus ruficollis, adarkling beetle
      - FamilyProstomidaeThomson, 1859
      - FamilySynchroidaeLacordaire, 1859
      - FamilyStenotrachelidaeThomson, 1859
      - FamilyOedemeridaeLatreille, 1810
      - FamilyMeloidaeGyllenhal, 1810 (blister beetles)
        Lytta magister, in theMeloidae
      - FamilyMycteridaePerty, 1840 (palm beetles and flower beetles)
      - FamilyBoridaeThomson, 1859
      - FamilyTrictenotomidaeBlanchard, 1845
      - FamilyPythidaeSolier, 1834
      - FamilyPyrochroidaeLatreille, 1806
      - FamilySalpingidaeLeach, 1815
      - FamilyAnthicidaeLatreille, 1819
      - FamilyAderidaeCsiki, 1909
      - FamilyScraptiidaeGistel, 1848
    - CladePhytophaga
      - SuperfamilyChrysomeloideaLatreille, 1802
        FamilyOxypeltidaeLacordaire, 1868
        FamilyVesperidaeMulsant, 1839
        FamilyDisteniidaeThomson, 1861
        FamilyCerambycidaeLatreille, 1802 (longhorn beetles)
        FamilyMegalopodidaeLatreille, 1802
        FamilyOrsodacnidaeThomson, 1859
        FamilyChrysomelidaeLatreille, 1802 (leaf beetles)
        Colorado potato beetle,Leptinotarsa decemlineata, aleaf beetle
      - SuperfamilyCurculionoideaLatreille, 1802
        FamilyNemonychidaeBedel, 1882
        FamilyAnthribidaeBillberg, 1820
        Family †UlyanidaeZherikhin, 1993
        FamilyBelidaeSchönherr, 1826
        FamilyCaridaeThompson, 1992
        FamilyAttelabidaeBillberg, 1820
        FamilyBrentidaeBillberg, 1820
        FamilyBrachyceridaeBillberg, 1820
        FamilyCurculionidaeLatreille, 1802 (snout beetles, weevils, and bark beetles)^[7]

References

[edit]

^Bouchard, P.; Bousquet, Y. (2020)."Additions and corrections to "Family-group names in Coleoptera (Insecta)"".ZooKeys (922):65–139.doi:10.3897/zookeys.922.46367.PMC 7113323.PMID 32256157.
^"Coleoptera information page".BugGuide.net. Retrieved2018-08-27.
^Lawrence, J.F.; Newton, A.F. Jr. (1995). "Families and subfamilies of Coleoptera (with selected genera, notes, references, and data on family-group names)". In Pakaluk, J.; Slipinski, S.A. (eds.).Biology, phylogeny, and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson. Warszawa: Muzeum i Instytut Zoologii PAN. pp. 779–1006.
^Bouchard, Patrice; Bousquet, Yves; Davies, Anthony E.; Alonso-Zarazaga, Miguel A.; et al. (2011)."Family-group names in Coleoptera (Insecta)".ZooKeys (88). Pensoft Publishers:1–972.doi:10.3897/zookeys.88.807.ISSN 1313-2989.PMC 3088472.PMID 21594053.
^Mckenna, Duane D.; Farrell, Brian D.; Caterino, Michael S.; Farnum, Charles W.; Hawks, David C.; Maddison, David R.; Seago, Ainsley E.; Short, Andrew E. Z.; Newton, Alfred F.; Thayer, Margaret K. (2015)."Phylogeny and evolution of Staphyliniformia and Scarabaeiformia: forest litter as a stepping stone for diversification of nonphytophagous beetles".Systematic Entomology.40 (1):35–60.doi:10.1111/syen.12093.ISSN 1365-3113.S2CID 83304675.
^Short, Andrew Edward Z. (2018)."Systematics of aquatic beetles (Coleoptera): current state and future directions".Systematic Entomology.43 (1):1–18.doi:10.1111/syen.12270.ISSN 1365-3113.S2CID 90159079.
^Chamorro, Maria Lourdes; Medeiros, Bruno A. S. de; Farrell, Brian D. (2021)."First phylogenetic analysis of Dryophthorinae (Coleoptera, Curculionidae) based on structural alignment of ribosomal DNA reveals Cenozoic diversification".Ecology and Evolution.11 (5):1984–1998.doi:10.1002/ece3.7131.ISSN 2045-7758.PMC 7920784.PMID 33717436.

ExtantColeoptera families

Kingdom:Animalia
Phylum:Arthropoda
Class:Insecta
Subclass:Pterygota
Infraclass:Neoptera
Superorder:Holometabola

SuborderArchostemata
Crowsoniellidae(Crowsoniella relicta) Cupedidae(reticulated beetles) Jurodidae(Sikhotealinia zhiltzovae) Micromalthidae Ommatidae

SuborderAdephaga

Extant families	Amphizoidae(trout-stream beetles) Aspidytidae Carabidae(ground beetles) Cicindelidae(tiger beetles) Dytiscidae(predaceous diving beetles) Gyrinidae(whirligig beetles) Haliplidae(crawling water beetles) Hygrobiidae Meruidae(Meru phyllisae) Noteridae(burrowing water beetles) Trachypachidae(false ground beetles)

SuborderMyxophaga
Hydroscaphidae(skiff beetles) Lepiceridae Sphaeriusidae Torridincolidae

SuborderPolyphaga

Bostrichiformia

Bostrichoidea	Bostrichidae(auger beetles) Dermestidae(skin beetles) Endecatomidae Jacobsoniidae(Jacobson's beetles) Nosodendridae(wounded-tree beetles) Ptiniidae(furniture beetles, death watch beetles, spider beetles)
Derodontoidea	Derodontidae(tooth-necked fungus beetles)

Cucujiformia

Chrysomeloidea	Cerambycidae(longhorn beetles) Chrysomelidae(leaf beetles) Disteniidae Megalopodidae Orsodacnidae Oxypeltidae Vesperidae
Cleroidea	Acanthocnemidae(Acanthocnemus nigricans) Biphyllidae(false skin beetles) Byturidae(fruitworm beetles) Chaetosomatidae Cleridae(checkered beetles) Lophocateridae Mauroniscidae Melyridae(soft-wing flower beetles) Metaxinidae(Metaxina ornata) Peltidae Phloiophilidae(Phloiophilus edwardsi) Phycosecidae Prionoceridae Protopeltidae Rentoniidae Rhadalidae Thanerocleridae Thymalidae Trogossitidae(bark-gnawing beetles)
Coccinelloidea	Akalyptoischiidae Alexiidae Anamorphidae Bothrideridae(dry bark beetles) Cerylonidae(minute bark beetles) Coccinellidae(lady beetles, or God's cows) Corylophidae(minute fungus beetles) Discolomatidae Endomychidae(handsome fungus beetles) Eupsilobiidae Euxestidae(well polished beetles) Latridiidae(minute brown scavenger beetles) Murmidiidae Mycetaeidae Teredidae
Cucujoidea	Agapythidae(Agapytho foveicollis) Boganiidae Cavognathidae Cryptophagidae(silken fungus beetles) Cucujidae(flat bark beetles) Cybocephalidae Cyclaxyridae Erotylidae(pleasing fungus beetles) Helotidae Hobartiidae Kateretidae(short-winged flower beetles) Laemophloeidae(lined flat bark beetles) Lamingtoniidae(Lamingtonium binnaberrense) Monotomidae(root-eating beetles) Myraboliidae Nitidulidae(sap beetles) Passandridae(parasitic flat bark beetles) Phalacridae(shining flower beetles) Phloeostichidae Priasilphidae Propalticidae Protocucujidae Silvanidae(silvanid flat bark beetles) Smicripidae(palmetto beetles) Sphindidae(dry-fungus beetles) Tasmosalpingidae
Curculionoidea (weevils)	Anthribidae(fungus weevils) Attelabidae(leaf-rolling weevils) Belidae(primitive weevils) Brentidae(straight snout weevils, New York weevil) Caridae Curculionidae(true weevils, bark beetles, ambrosia beetles) Nemonychidae(pine flower weevils)
Lymexyloidea	Lymexylidae(ship-timber beetles)
Tenebrionoidea	Aderidae(ant-like leaf beetles) Anthicidae(ant-like flower beetles) Archeocrypticidae(cryptic fungus beetles) Boridae(conifer bark beetles) Chalcodryidae Ciidae(minute tree-fungus beetles) Melandryidae(false darkling beetles) Meloidae(blister beetles) Mordellidae(tumbling flower beetles) Mycetophagidae(hairy fungus beetles) Mycteridae(palm and flower beetles) Oedemeridae(false blister beetle) Perimylopidae, or Promecheilidae Prostomidae(jugular-horned beetles) Pterogeniidae Pyrochroidae(fire-coloured beetles) Pythidae(dead log bark beetles) Ripiphoridae(wedge-shaped beetles) Salpingidae(narrow-waisted bark beetles) Scraptiidae(false flower beetles) Stenotrachelidae(false longhorn beetles) Synchroidae(synchroa bark beetles) Tenebrionidae(darkling beetles) Tetratomidae(polypore fungus beetles) Trictenotomidae Ulodidae Zopheridae(ironclad beetles, cylindrical bark beetles)

Elateriformia

Buprestoidea	Buprestidae(jewel beetles, or metallic wood-boring beetles) Schizopodidae
Byrrhoidea	Byrrhidae(pill beetles) Callirhipidae(cedar beetles) Chelonariidae(turtle beetles) Cneoglossidae Dryopidae(long-toed water beetles) Elmidae(riffle beetles) Eulichadidae(forest stream beetles) Heteroceridae(variegated mud-loving beetles) Limnichidae(minute mud beetles) Lutrochidae(travertine beetles) Psephenidae(water-penny beetles) Ptilodactylidae
Dascilloidea	Dascillidae(soft bodied plant beetles) Rhipiceridae(cicada beetle, cicada parasite beetles)
Elateroidea	Artematopodidae(soft-bodied plant beetles) Brachypsectridae(Texas beetles) Cantharidae(soldier beetles) Cerophytidae(rare click beetles) Elateridae(click beetles) Eucnemidae(false click beetles) Jurasaidae Lampyridae(fireflies) Lycidae(net-winged beetles) Omethidae(false fireflies, long-lipped beetles) Phengodidae(glowworm beetles) Rhagophthalmidae Sinopyrophoridae Throscidae(false metallic wood-boring beetles)
Rhinorhipoidea	Rhinorhipidae(Rhinorhipus tamborinensis)
Scirtoidea	Clambidae Decliniidae(Declinia relicta) Eucinetidae(plate-thigh beetles) Scirtidae

Scarabaeiformia

Scarabaeoidea	Belohinidae(Belohina inexpectata) Bolboceratidae Diphyllostomatidae(false stag beetles) Geotrupidae(dor beetles) Glaphyridae(bumble bee scarab beetles) Glaresidae(enigmatic scarab beetles) Hybosoridae(scavenger scarab beetles) Lucanidae(stag beetles) Ochodaeidae(sand-loving scarab beetles) Passalidae(betsy beetles) Pleocomidae(rain beetles) Scarabaeidae(scarabs) Trogidae(hide beetles)

Staphyliniformia

Histeroidea	Histeridae(clown beetles) Sphaeritidae(false clown beetles) Synteliidae
Hydrophiloidea	Epimetopidae Georissidae(minute mud-loving beetles) Helophoridae Hydrochidae Hydrophilidae(water scavenger beetles) Spercheidae
Staphylinoidea	Agyrtidae(primitive carrion beetles) Hydraenidae Leiodidae(round fungus beetles) Ptiliidae(feather-winged beetles) Silphidae(carrion beetles) Staphylinidae(rove beetles)

List of subgroups of the order Coleoptera

Retrieved from "https://en.wikipedia.org/w/index.php?title=List_of_subgroups_of_the_order_Coleoptera&oldid=1316047386"

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Beetle taxonomy

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