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Hornbeam

From Wikipedia, the free encyclopedia
Genus of flowering plants

Hornbeam
Temporal range:49.42–0 MaYpresianRecent
European hornbeam foliage
Scientific classificationEdit this classification
Kingdom:Plantae
Clade:Tracheophytes
Clade:Angiosperms
Clade:Eudicots
Clade:Rosids
Order:Fagales
Family:Betulaceae
Subfamily:Coryloideae
Genus:Carpinus
L. (1753)
Synonyms[1]

DistegocarpusSiebold & Zucc. (1846)

Hornbeams arehardwood trees in the plantgenusCarpinus in the familyBetulaceae. Its species occur across much of thetemperate regions of theNorthern Hemisphere.

Common names

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The common English namehornbeam derives from the hardness of the woods (likened tohorn) and the Old Englishbeam, "tree" (cognate with Dutchboom and GermanBaum).

TheAmerican hornbeam is also occasionally known as blue-beech, ironwood, or musclewood, the first from the resemblance of the bark to that of theAmerican beechFagus grandifolia, the other two from the hardness of the wood and the muscled appearance of the trunk and limbs.

The botanical name for the genus,Carpinus, is the original Latin name for the European species, although someetymologists derive it from the Celtic for ayoke.[2]

Description

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European hornbeam in Germany, during May

Hornbeams are small, slow-growing,understory trees with a natural, rounded form growing 4.5–9 metres (15–30 feet) tall and wide; the exemplar species—theEuropean hornbeam—reaches a maximum height of 32 m (105 ft).[3]: 296 

Leaves aredeciduous, dark-green, alternate and simple with a coarsely-serrated margin, varying from 3 to 10 centimetres (1 to 4 inches) in length. In autumn, leaves turn various shades of yellow, orange and red. Hornbeam saplings, stressed trees, and the lower branches of mature trees may exhibitmarcescence—where leaves wither with autumn butabscission (leafdrop) is delayed until spring.[4]

The smooth, gray trunk and larger branches of a mature tree exhibit a distinctive muscle-like fluting.[5]

As with other members of thebirch family, hornbeam flowers are wind-pollinated pendulouscatkins, produced in spring. Male and female flowers are on separate catkins, but on the same tree (monoecious). Female flowers give way to distinctive clusters of winged seeds that somewhat resemble the hops-like seeds of ironwood.[6]

The fruit is a smallnut about 3–6 millimetres (1814 in) long, held in a leafybract; the bract may be either trilobed or simple oval, and is slightly asymmetrical. The asymmetry of the seedwing makes it spin as it falls, improvingwind dispersal. The shape of the wing is important in the identification of different hornbeam species. Typically, 10–30 seeds are on each seed catkin.[7][8]

Taxonomy

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Formerly some taxonomists segregated them with the generaCorylus (hazels) andOstrya (hop-hornbeams) in aseparate family,Corylaceae. Modern botanists placeCarpinus in the subfamilyCoryloideae of the familyBetulaceae.[9][10] Species ofCarpinus are often grouped into two subgeneraCarpinus subgenusCarpinus andCarpinus subgenus Distegicarpus.

Phylogentic analyses have shown thatOstrya likely evolved from aCarpinus ancestor somewhere inC. subg. Distegicarpus makingCarpinusparaphyletic. The fossil record of the genus extends back to theEarly Eocene,Ypresian of northwestern North America, with the speciesCarpinus perryae described from fossil fruits found in theKlondike Mountain Formation ofRepublic, Washington.[9]

Southern Balkan Hornbeam
Carpinus austrobalcanica, southern Balkan hornbeam, first described 2024

Species

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43 species are currently accepted.[11]

Distribution and habitat

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The 43 species occur across much of thetemperate regions of the northern hemisphere, with the greatest number of species in east Asia, particularly China. Only three species occur in Europe, only one in eastern North America, and one inMesoamerica.[1][16][17][18][19]Carpinus betulus can be found in Europe, Turkey and Ukraine.[12]

Ecology

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Hornbeam trunk

Hornbeams are used as food plants by thelarvae of someLepidoptera species, includingautumnal moth,common emerald,feathered thorn,walnut sphinx,Svensson's copper underwing, andwinter moth (recorded on European hornbeam) as well as theColeophora case-bearersC. currucipennella andC. ostryae.[20]

Uses

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Hornbeams yield a very hard timber, giving rise to the name "ironwood".[21] Driedheartwood billets are nearly white and are suitable for decorative use. For general carpentry, hornbeam is rarely used, partly due to the difficulty of working it.

The wood is used to construct carving boards, tool handles, handplane soles, coach wheels, piano actions, shoe lasts, and other products where a very tough, hard wood is required.

The wood can also be used as gear pegs in simple machines, including traditionalwindmills.[21] It is sometimescoppiced to provide hardwood poles. It is also used in parquet flooring and for making chess pieces.

References

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  1. ^ab"Carpinus L., Sp. Pl.: 998 (1753)".World Checklist of Selected Plant Families.Royal Botanic Gardens, Kew.[permanent dead link]
  2. ^Gledhill D. 1985.The Names of Plants. Cambridge University PressISBN 0521366755
  3. ^Stace, C. A. (2010).New Flora of the British Isles (Third ed.). Cambridge, UK: Cambridge University Press.ISBN 9780521707725.
  4. ^Cutler, Tony (2020).The world encyclopedia of trees: a reference and identification guide to 1300 of the world's most significant trees. London?: Lorenz Books.ISBN 978-0754834755.
  5. ^"Carpinus caroliniana - Plant Finder".www.missouribotanicalgarden.org.
  6. ^Furlow, John J. (1990)."The Genera of Betulaceae in the Southeastern United States".Journal of the Arnold Arboretum.71 (1):1–67.doi:10.5962/bhl.part.24925.JSTOR 43782230.
  7. ^"Hornbeam, American (Musclewood) | Nebraska Forest Service".nfs.unl.edu.
  8. ^The complete encyclopedia of trees of the world : the ultimate reference and identification guide to more than 1300 of the most spectacular, best-loved and unusual trees around the globe, with 3000 specially commissioned illustrations, maps and photographs. Creative Media Partners, LLC. 10 September 2021.ISBN 978-1015282605.
  9. ^abPigg, K.B.; Manchester, S.R.; Wehr, W.C. (2003). "Corylus,Carpinus, andPalaeocarpinus (Betulaceae) from the Middle Eocene Klondike Mountain and Allenby Formations of Northwestern North America".International Journal of Plant Sciences.164 (5):807–822.doi:10.1086/376816.S2CID 19802370.
  10. ^Forest, F.; Savolainen, V.; Chase, M. W.; Lupia, R.; Bruneau, A.; Crane, P. R. (2005). "Teasing apart molecular- versus fossil-based error estimates when dating phylogenetic trees: a case study in the birch family (Betulaceae)".Systematic Botany.30 (1):118–133.doi:10.1600/0363644053661850.S2CID 86080433.
  11. ^"Carpinus L."Plants of the World Online.Royal Botanic Gardens, Kew. Retrieved25 September 2024.
  12. ^abCarpinus betulus. 2.nd Ed., The Royal Horticultural Society, Dorling Kindersley Ltd, London, pp. 234, 235.
  13. ^"Carpinus fangiana". Rogers Trees and Shrubs. Archived fromthe original on 15 July 2011.
  14. ^Dai, J.; Sun, B.; Xie, S.; Lin, Z.g; Wu, J.; Dao, K. (2013). "A new species ofCarpinus (Betulaceae) from the Pliocene of Yunnan Province, China".Plant Systematics and Evolution.299 (3):643–658.Bibcode:2013PSyEv.299..643D.doi:10.1007/s00606-012-0750-1.S2CID 16941126.
  15. ^English Names for Korean Native Plants(PDF). Pocheon:Korea National Arboretum. 2015. p. 400.ISBN 978-89-97450-98-5. Archived fromthe original(PDF) on 25 May 2017. Retrieved8 December 2016 – viaKorea Forest Service.
  16. ^Flora of China, Vol. 4 Page 289,鹅耳枥属 e er li shu,Carpinus Linnaeus, Sp. Pl. 2: 998. 1753.
  17. ^Flora of North America, Vol. 3, Hornbeam,Carpinus Linnaeus, Sp. Pl. 2: 998. 1753; Gen. Pl. ed. 5, 432, 1754.
  18. ^"Carpinus betulus L."Altervista Flora Italiana.
  19. ^"2013 BONAP North American Plant Atlas. TaxonMaps". Biota of North America Program.
  20. ^Miscellaneous Publication.U.S. Department of Agriculture. 1972. p. 297.
  21. ^abArchived atGhostarchive and theWayback Machine:Eichhorn, Markus; Haran, Brady (1 December 2011)."The Hornbeam's Heartbeat".test-tube.org.uk.University of Nottingham. Retrieved30 December 2012.

External links

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Wikimedia Commons has media related toCarpinus.
Look uphornbeam in Wiktionary, the free dictionary.
Wikisource has the text of the1911Encyclopædia Britannica article "Hornbeam".
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