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Hoof

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Tip of a toe of an ungulate mammal, strengthened by a thick and horny keratin covering
"Cow toe" redirects here. For prostration in East Asian cultures, seekowtow.
The feet of theroe deer (Capreolus capreolus) have cloven hooves with prominent dewclaws.

Thehoof (pl.:hooves) is the tip of atoe of anungulatemammal, which is covered and strengthened with a thick and hornykeratin covering.[1]Artiodactyls are even-toed ungulates, species whose feet have an even number of digits; the ruminants with two digits are the most numerous, e.g.giraffe,deer,bison,cattle,goats,gazelles,pigs, andsheep.[2] The feet ofperissodactyl mammals have an odd number of toes, e.g. thehorse, therhinoceros, and thetapir.[3] Although hooves are limb structures primarily found inplacental mammals, hadrosaurs such asEdmontosaurus possessed hoofed forelimbs. ThemarsupialChaeropus also had hooves.[4]

Description

[edit]
Sagittal section of a wildhorse hoof.
Pink: soft tissues;
light gray: bone;
cyan: tendons;
red:corium;
yellow: digital cushion;
dark gray: frog;
orange: sole;
brown: walls

The hoof surrounds the distal end of thesecond phalanx, thedistal phalanx, and thenavicular bone.[5] The hoof consists of thehoof wall, the bars of the hoof, the sole andfrog and soft tissue shock absorption structures.[5] The weight of the animal is normally borne by both the sole and the edge of the hoof wall. Hooves perform many functions, including supporting the weight of the animal, dissipating the energy impact as the hooves strike the ground or surface, protecting the tissues and bone within the hoof capsule, and providing traction for the animal. Numerous factors can affect hoof structure and health, including genetics, hoof conformation, environmental influences, and athletic performance of the animal. The ideal hoof has a parallel hoof-pastern axis, a thick hoof wall, adequate sole depth, a solid heel base and growth rings of equal size under the coronary band.[5]

There are four layers within the exterior wall of the hoof. From the outside, a hoof is made up of the stratum externum, the stratum medium, the stratum internum and the dermis parietis. The stratum externum and the stratum medium are difficult to distinguish, the stratum externum is thin and the stratum medium is what makes up the bulk of the hoof wall.[6] Inside the hoof wall is alaminar junction, a soft tissue structure that allows the hoof to withstand the demands of force transmission it undergoes.[7] This tissue structure binds the inner surface of the hoof wall, the dermis parietis and the outer surface of the third phalanx.[7]

Mosteven-toed ungulates (such assheep,goats,deer,cattle,bison andpigs) have two main hooves on each foot, together called acloven hoof.[Note 1] Most of these cloven-hooved animals also have two smaller hooves calleddewclaws a little further up the leg – these are not normally used for walking, but in some species with larger dewclaws (such as deer and pigs) they may touch the ground when running or jumping, or if the ground is soft. In the mountain goat, the dewclaw serves to provide extra traction when descending rocky slopes as well as additional drag on loose or slippery surfaces made of ice, dirt, or snow.[8] Other cloven-hooved animals (such asgiraffes andpronghorns) have no dewclaws.

In some so-called "cloven-hooved" animals, such ascamels, the "hoof" is not properly a hoof – it is not a hard or rubbery sole with a hard wall formed by a thick nail – instead it is a soft toe with little more than a nail merely having an appearance of a hoof.

Someodd-toed ungulates (equids) have one hoof on each foot; others have (or had) three distinct hooved or heavily nailed toes, or one hoof and two dewclaws. Thetapir is a special case, having three toes on each hind foot and four toes on each front foot.

Management

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Anoxshoe is being nailed on the hooves of a bull used fordraft atChinawal, India, to prevent them from wearing out too much.

Hooves grow continuously. In nature, wild animals are capable of wearing down the hoof as it continuously grows, but captive domesticated species often must undergo specific hoof care for a healthy, functional hoof. Proper care improves biomechanical efficiency and prevents lameness.[5] If not worn down enough by use, such as in the dairy industry, hooves may need to be trimmed. However, too much wear can result in damage of the hooves, and for this reason,horseshoes andoxshoes are used by animals that routinely walk on hard surfaces and carry heavy weight.[9]

Horses

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Within the equine world, the expression, "no foot, no horse" emphasizes the importance of hoof health.[10] Hoof care is important in theequine industry.[5] Problems that can arise with poorhorse hoof care include hoof cracks, thrush, abscesses andlaminitis.[11]

Cattle

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Trimming the hoof of a cow with anangle grinder

Acow hoof is cloven, or divided, into two approximately equal parts, usually called claws.[12] Approximately 95% of lameness in dairy cattle occurs in the feet.[12] Lameness in dairy cows can reducemilk production and fertility, and cause reproductive problems and suffering. For dairy farm profitability, lameness, behind only infertility andmastitis, is the third most important cow health issue.[13]

Hoof trimmers trim and care for bovine hooves, usuallydairy cows. Hooves can be trimmed with a sharp knife while the cow is restrained and positioned with ropes. Professional hoof-trimming tend to useangle grinders and some type ofhoof trimming crush to make the process quicker and less physically demanding on the hoof trimmer. A hoof trimmer using modern machinery may trim the hooves of more than 10,000 cows per year.[14][15] The trimmer shapes the hooves to provide the optimal weight-bearing surface. A freshly trimmed hoof may be treated withcopper sulfate pentahydrate to preventfoot rot.

Gallery

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  • Rear foot of a giraffe (no dewclaws)
    Rear foot of agiraffe (no dewclaws)
  • Rear hooves of a horse
    Rear hooves of ahorse
  • Malayan tapir hooves: front with four toes, back with three toes
    Malayan tapir hooves: front with four toes, back with three toes
  • Cloven hooves of a pig
    Cloven hooves of apig
  • Sheep hooves
    Sheep hooves
  • American bison hooves
    Americanbison hooves
  • Broad hooves of a camel
    Broad hooves of acamel
  • Cloven hooves of a deer
    Cloven hooves of a deer
  • Reindeer hooves
    Reindeer hooves
  • Horses' hooves in snow
    Horses' hooves in snow
  • Cattle hooves
    Cattle hooves
  • Muskoxen broad hooves
    Muskoxen broad hooves
  • Padded hooves of a deer
    Padded hooves of a deer

In culture

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Hooves have historical significance in ceremonies and games. They have been used in burial ceremonies.[16]

See also

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Notes

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  1. ^The term "cloven hoof" therefore being a technical misnomer as nothing is actually "cloven".


References

[edit]
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  1. ^"the definition of hoof".Dictionary.com. Retrieved2016-11-30.
  2. ^Keller, Anna; Clauss, Marcus; Muggli, Evelyne; Nuss, Karl (2009-07-15)."Even-toed but uneven in length: the digits of artiodactyls"(PDF).Zoology.112 (4):270–278.doi:10.1016/j.zool.2008.11.001.PMID 19386479. Archived fromthe original(PDF) on 2021-03-03. Retrieved2019-12-14.
  3. ^Holbrook, Luke T. (1999-09-01)."The Phylogeny and Classification of Tapiromorph Perissodactyls (Mammalia)".Cladistics.15 (3):331–350.doi:10.1111/j.1096-0031.1999.tb00270.x.ISSN 1096-0031.PMID 34902952.S2CID 221584511.
  4. ^Sánchez-Villagra, Marcelo R. (22 December 2012)."Why are There Fewer Marsupials than Placentals? On the Relevance of Geography and Physiology to Evolutionary Patterns of Mammalian Diversity and Disparity"(PDF).Journal of Mammalian Evolution.20 (4):279–290.doi:10.1007/s10914-012-9220-3.S2CID 18789008.
  5. ^abcdeO'Grady, Stephen E. (2008). "Basic Farriery for the Performance Horse".Veterinary Clinics of North America: Equine Practice.24 (1):203–218.doi:10.1016/j.cveq.2007.12.002.PMID 18314044.
  6. ^Goulet, Catherine; Olive, Julien; Rossier, Yves; Beauchamp, Guy (2015-11-01). "RADIOGRAPHIC AND ANATOMIC CHARACTERISTICS OF DORSAL HOOF WALL LAYERS IN NONlAMINITIC HORSES".Veterinary Radiology & Ultrasound.56 (6):589–594.doi:10.1111/vru.12280.ISSN 1740-8261.PMID 26226838.
  7. ^abDouglas, Janet E.; Thomason, Jeffrey J. (2000). "Shape, Orientation and Spacing of the Primary Epidermal Laminae in the Hooves of Neonatal and Adult Horses(Equus caballus)".Cells Tissues Organs.166 (3):304–318.doi:10.1159/000016744.PMID 10765026.S2CID 36816180.
  8. ^A Beast the Color of Winter: The Mountain Goat Observed. U of Nebraska Press. 1 February 2002. p. 52.ISBN 978-0-8032-6421-2.
  9. ^"Why Do Only Some Horses Wear Shoes?".wideopenpets.com. 2016-04-21.
  10. ^Aoki, Yasuhiro (2006). "Changes in walking parameters of milking cows after hoof trimming|(Aoki, Y. et al., 2006)".Animal Science Journal.77:103–109.doi:10.1111/j.1740-0929.2006.00326.x.Lameness, behind infertility and mastitis, is the biggest cause of economic loss to a dairy farmer (Weaver, A., 2006). Many farmers and veterinarians have used a phase that distinguished that if the animal has bad hooves then it is of no use, the most common version of this phrase is used with equines, "No hoof no horse."
  11. ^"Common hoof problems : Horse : University of Minnesota Extension".www.extension.umn.edu. Archived fromthe original on 2016-12-01. Retrieved2016-11-30.
  12. ^abShearer, Jan K.; Sarel Rens Van Amstel; Adrian Gonzalez (2005).Manual of foot care in cattle. Hoard's Dairyman Books.ISBN 9780932147424.
  13. ^Weaver, A. D. (1985-01-01). "Lameness in cattle—Investigational and diagnostic check lists".British Veterinary Journal.141 (1):27–33.doi:10.1016/0007-1935(85)90123-X.PMID 3995246.
  14. ^Dolecheck, K.A.; Dwyer, R.M.; Overton, M.W.; Bewley, J.M. (September 2018)."A survey of United States dairy hoof care professionals on costs associated with treatment of foot disorders".Journal of Dairy Science.101 (9):8313–8326.doi:10.3168/jds.2018-14718. Retrieved29 May 2024.
  15. ^"HOOF TRIMMING CHUTE ANKA PRO 2.0 (380V)".CowCare. Retrieved29 May 2024.
  16. ^M. E. Robertson-Mackay (1980)."A head and hooves burial beneath a round barrow, with other Neolithic and Bronze Age sites on Hemp Knoll, near Avebury, Wiltshire". Proceedings of the Prehistoric Society.{{cite journal}}:Cite journal requires|journal= (help)
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