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Hadrianus (turtle)

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Extinct genus of tortoise

Hadrianus Temporal range:55.4–33.9 Ma PreꞒ Ꞓ O S D C P T J K Pg N Eocene^[1]

Hadrianus majusculus
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Clade:	Pantestudinoidea
Superfamily:	Testudinoidea
(unranked):	†Pantestudinidae
Genus:	†Hadrianus Cope,1872^[2]^[1]
Species
H. corsoni H. majusculus

Hadrianus is anextinct genus oftortoise belonging to theTestudinidae^[1] found in theUnited States, theYolomécatl Formation ofMexico, theAlai Beds ofKyrgyzstan andSpain^{[citation needed]} and believed to be the oldest true tortoise known.^[3] The genus is thought to be closely related to the genusManouria.^[3] The genus may have evolved in the subtropics of Asia and subsequently migrated to North America and Europe.^[3] Evangelos Vlachos (2018) reassessed the North American species attributed to the genus, and determined only two as accepted namelyH. corsoni &H. majusculus. The remaining species were identified as either junior synonyms, moved to other genera or considerednomen dubium due to incomplete fossils.^[4]

Taxonomy

[edit]

The species 'Testudo'sharanensis is hypothesized to actually be a species ofHadrianus.^[5]

Hadrianus corsoni(Leidy, 1871)
- SynonymsEmys carteriLeidy, 1871;Testudo hadrianaCope, 1871;Hadrianus quadratusCope, 1871;Hadrianus octonariaCope, 1871;Hadrianus tumidusHay 1908;Hadrianus robustusGilmore, 1915;Hadrianus utahensisGilmore, 1915;Geochelone gilmoreiAuffenberg, 1974
Hadrianus majusculusHay, 1904

Nomen dubium

[edit]

"Hadrianus allabiatus"Cope, 1871

Moved species

[edit]

"Hadrianus schucherti"Hay, 1899 moved toCymatholcus schucherti(Hay, 1899)

References

[edit]

^^a ^b ^c"Fossilworks: Hadrianus".fossilworks.org. Retrieved17 December 2021.
^E. D. Cope. 1872. Second account of new Vertebrata from the Bridger Eocene. Proceedings of the American Philosophical Society (separate) 1-3
^^a ^b ^cEhret, Dana Joseph 2004 "Skeletochronology as a method of aging OligoceneGopherus laticuneus andStylemys nebrascensis, usingGopherus polyphemus as a modern analog" Thesis, University of Florida.
^Vlachos, E (2018)."A Review of the Fossil Record of North American Turtles of the Clade Pan-Testudinoidea".Bulletin of the Peabody Museum of Natural History.59 (1): 3.doi:10.3374/014.059.0101.hdl:11336/117896.ISSN 0079-032X.S2CID 214641639.
^Chroust, Milan; Szczygielski, Tomasz; Luján, Àngel H. (2025-10-03)."Manouria morla sp. nov., the Ancient One: an Early Miocene large tortoise from the Swamps of Ahníkov, Czechia".Swiss Journal of Palaeontology.144 (1): 63.doi:10.1186/s13358-025-00400-6.ISSN 1664-2384.

Taxon identifiers
Hadrianus	Wikidata:Q5638062 Wikispecies:Hadrianus GBIF:4818723 IRMNG:1367012 Open Tree of Life:4132403 Paleobiology Database:58498

Tortoise family

Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Subclass:Anapsida
Order:Testudines
Suborder:Cryptodira
Superfamily:Testudinoidea
Family:Testudinidae

Genera

Species of thetortoise family

Agrionemys

Russian tortoise

Angulate tortoise

†Hadrianus

†Hesperotestudo percrassa

Pancake tortoise

Leopard tortoise

†Stylemys

Testudo

Phylogenetic arrangement of turtles based onturtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. Key: †=extinct.

Testudines

Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Diapsida
Order:Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based onTurtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See alsoList of Testudines families

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