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Gobiazhdarcho

From Wikipedia, the free encyclopedia
Genus of azhdarchid pterosaurs

Gobiazhdarcho
Third cervical vertebra of the holotype
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Order:Pterosauria
Suborder:Pterodactyloidea
Clade:Azhdarchoidea
Family:Azhdarchidae
Clade:Quetzalcoatlini
Genus:Gobiazhdarcho
Pêgas, Zhou & Kobayashi,2025
Species:
G. tsogtbaatari
Binomial name
Gobiazhdarcho tsogtbaatari
Pêgas, Zhou & Kobayashi, 2025

Gobiazhdarcho (lit.'Gobi Desertazhdar') is anextinct genus ofazhdarchid pterosaurs known from theLate CretaceousBayanshiree Formation of Mongolia. The genus contains asingle species,Gobiazhdarcho tsogtbaatari, known from a three cervical (neck) vertebrae. It was closely related toQuetzalcoatlus and coexisted withTsogtopteryx, another azhdarchid more closely related toHatzegopteryx.

Discovery and naming

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Mongolian Cretaceous fossil localities;Gobiazhdarcho is known from the Burkhant locality in Area D (Bayanshiree Formation)

In 1995, a joint paleontological expedition between the Hayashibara Museum of Natural Sciences and theMongolian Paleontological Center conducted fieldwork in theGobi Desert of Mongolia. At the 'Burkhant' locality of theBayanshiree Formation inDornogovi Province, workers collected three associated cervical (neck) vertebral elements. These specimens,accessioned as MPC−Nd 100/302, include theatlantoaxis (fused first and second cervicals), the third cervical, and theposterior (back) part of the sixth cervical (identified as the fourth by Averianov in 2014).[1][2] In 2009, Mahito Watabe and colleagues described this specimen, in addition to another isolated pterosaur cervical vertebra from the 'Bayshin Tsav' locality of the same formation. The authors refrained from naming either specimen, but discussed their anatomy and likelyphylogenetic affinities in depth.[3]

In 2025, R. V. Pêgas, Xuanyu Zhou, and Yoshitsugu KobayashidescribedGobiazhdarcho tsogtbaatari as a new genus and species of pterosaurs based on the 'Burkhant azhdarchid' fossil remains. Thegeneric name,Gobiazhdarcho, combines "Gobi", a reference to the discovery of the specimen in the Gobi Desert, with "azhdarcho", derived fromazhdar, a dragonlike creature inPersian myth. Thespecific name,tsogtbaatari, honors Mongolian paleontologistKhishigjav Tsogtbaatar [ja]. In the same publication, the authors also named the Bayshin Tsav specimen as another new azhdarchid,Tsogtopteryx.[1]

Prior to the naming ofGobiazhdarcho andTsogtopteryx, the only pterosaur named from Mongolia was thedsungaripteridNoripterus parvus,[4] known from theTsagan-Tsab Formation.[1] Several fragmentary indeterminate pterosaur remains are also known from the country, including a possibleanurognathid,anhanguerid,[5] andtapejaroid,[6] in addition to a possible azhdarchid long bone found in the stomach of adromaeosaurid dinosaur and the very fragmentary remains of a giant azhdarchid.[7][8]

Description

[edit]
Atlantoaxis (left) and fourth cervical vertebra (right) of the holotype

Based on the size of the holotype vertebrae (third cervical 9.06 centimetres (3.57 in) long), thewingspan ofGobiazhdarcho was estimated at 3–3.5 metres (9.8–11.5 ft), which is comparable in size toEurazhdarcho,Wellnhopterus, andZhejiangopterus. The specimen likely belongs to a late subadult individual that was nearly skeletally mature (fully grown). This was determined based on the lack of a grained texture often seen in immature pterosaur bones and the full fusion of the atlas and axis—both characteristics of mature pterosaurs—combined with the lack of ribs fused to the vertebra, an indication of immaturity. In comparison, the coevalTsogtopteryx was fully grown but much smaller, with a 1.6–1.9 metres (5.2–6.2 ft) wingspan.[1]

Classification

[edit]

To determine the relationships and affinities ofGobiazhdarcho, Pêgas and colleagues scored it in a comprehensive pterosaur-focusedphylogenetic matrix modified from Zhou et al. (2025),[9] deriving from Pêgas (2024)[4] and other earlier publications, with modifications based on newer literature. Their analyses recoveredGobiazhdarcho as thesister taxon toNipponopterus in an early-branching clade within the cladeQuetzalcoatlini, which includes azhdarchids more closely related toQuetzalcoatlus thanHatzegopteryx. The results are displayed in thecladogram below:[1]

Azhdarchidae

Paleoenvironment

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Speculativelife restoration ofGobiazhdarcho (larger; red) andTsogtopteryx (smaller; blue) in a Bayanshiree environment with theornithopod dinosaurGobihadros

Gobiazhdarcho is known from the 'Burkhant' locality of theBayanshiree Formation. Examinations of themagnetostratigraphy of the formation confirm that it lies entirely within theCretaceous Long Normal, which lasted only until the end of theSantonian stage.CalciteU–Pb measurements estimate the age of the Bayanshiree Formation from 95.9 ± 6.0 million to 89.6 ± 4.0 million years ago, in theAlbian throughSantonian ages.[12][13] Considering all available evidence, there are likely two distinct levels of the formation—an 'upper' and 'lower'—partially based on faunal differences: a lower part lasting from theCenomanian to lateTuronian ages and an upper part lasting the late Turonian to Santonian ages in the lateCretaceous period.[14] The Burkhant is part of the upper Bayanshiree.[1]

Fluvial,lacustrine andcaliche-based sedimentation indicates a lessersemi-arid climate, with the presence of wet environments composed of largemeanders andlakes. Large-scale cross-stratification in many of thesandstone layers at the Baynshire and Burkhant localities seems to indicate largemeandering rivers, and these large water bodies may have drained the eastern part of theGobi Desert.[15][12]

A vastfaunal diversity is known in the formation, comprising dinosaur and non-dinosaur genera. The largedromaeosauridAchillobator is also recognized from the Burkhant locality.[16][1] Theropod dinosaurs from other localities include thetherizinosaursDuonychus,Erlikosaurus,Enigmosaurus, andSegnosaurus,[17] thetyrannosauroidKhankhuuluu,[18] and theornithomimosaurGarudimimus.[19] Herbivorous dinosaurs are represented by theankylosaursTalarurus andTsagantegia,[20] the smallmarginocephaliansAmtocephale (apachycephalosaur) andGraciliceratops (aceratopsian),[21][22] thehadrosauroidGobihadros,[23] and thesauropodErketu.[24] Other fauna includesemiaquatic reptiles likecrocodylomorphs andnanhsiungchelyid turtles, and various fish.[14][25] Numerous fossilizedfruits have been recovered from the Bor Guvé and Khara Khutul localities.[24]

References

[edit]
  1. ^abcdefgPêgas, R. V.; Zhou, Xuanyu; Kobayashi, Yoshitsugu (2025)."Azhdarchid pterosaur diversity in the Bayanshiree Formation, Upper Cretaceous of the Gobi Desert, Mongolia".PeerJ.13 e19711.doi:10.7717/peerj.19711.PMC 12447946.
  2. ^Averianov, Alexander (2025-08-11)."Review of taxonomy, geographic distribution, and paleoenvironments of Azhdarchidae (Pterosauria)".ZooKeys.432:1–107.doi:10.3897/zookeys.432.7913.ISSN 1313-2970.PMC 4141157.PMID 25152671.
  3. ^Watabe, Mahito; Tsuihiji, Takanobu; Suzuki, Shigeru; Tsogtbaatar, Khishigjav (2009)."The First Discovery of Pterosaurs from the Upper Cretaceous of Mongolia"(PDF).Acta Palaeontologica Polonica.54 (2):231–242.doi:10.4202/app.2006.0068.ISSN 0567-7920.
  4. ^abPêgas, Rodrigo V. (2024-06-10)."A taxonomic note on the tapejarid pterosaurs from the Pterosaur Graveyard site (Caiuá Group, ?Early Cretaceous of Southern Brazil): evidence for the presence of two species".Historical Biology:1–22.doi:10.1080/08912963.2024.2355664.ISSN 0891-2963.
  5. ^Bakhurina, Natalia N.; Unwin, David M. (1995). "A survey of pterosaurs from the Jurassic and Cretaceous of the former soviet union and Mongolia".Historical Biology.10 (3):197–245.doi:10.1080/10292389509380522.ISSN 0891-2963.
  6. ^Andres, Brian; Norell, Mark A. (2005-03-24). "The First Record of a Pterosaur from the Early Cretaceous Strata of Öösh (Övörkhangai; Mongolia)".American Museum Novitates.3472:1–6.doi:10.1206/0003-0082(2005)472<0001:TFROAP>2.0.CO;2.ISSN 0003-0082.
  7. ^Hone, David; Tsuihiji, Takanobu; Watabe, Mahito; Tsogtbaatr, Khishigjaw (2012-05-01). "Pterosaurs as a food source for small dromaeosaurs".Palaeogeography, Palaeoclimatology, Palaeoecology.331–332:27–30.doi:10.1016/j.palaeo.2012.02.021.
  8. ^Tsuihiji, Takanobu; Andres, Brian; O'connor, Patrick M.; Watabe, Mahito; Tsogtbaatar, Khishigjav; Mainbayar, Buuvei (September 2017). "Gigantic pterosaurian remains from the Upper Cretaceous of Mongolia".Journal of Vertebrate Paleontology.37 (5) e1361431.doi:10.1080/02724634.2017.1361431.ISSN 0272-4634.
  9. ^Zhou, Xuanyu; Ikegami, Naoki; Pêgas, Rodrigo V.; Yoshinaga, Toru; Sato, Takahiro; Mukunoki, Toshifumi; Otani, Jun; Kobayashi, Yoshitsugu (March 2025). "Reassessment of an azhdarchid pterosaur specimen from the Mifune Group, Upper Cretaceous of Japan".Cretaceous Research.167 106046.Bibcode:2025CrRes.16706046Z.doi:10.1016/j.cretres.2024.106046.ISSN 0195-6671.
  10. ^Vremir, Mátyás;Witton, Mark;Naish, Darren; Dyke, Gareth;Brusatte, Stephen L.;Norell, Mark; Totoianu, Radu (March 17, 2015). "A Medium-Sized Robust-Necked Azhdarchid Pterosaur (Pterodactyloidea: Azhdarchidae) from the Maastrichtian of Pui (Haţeg Basin, Transylvania, Romania)".American Museum Novitates (3827):1–16.doi:10.1206/3827.1.ISSN 0003-0082.
  11. ^Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018)."Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary".PLOS Biology.16 (3) e2001663.doi:10.1371/journal.pbio.2001663.PMC 5849296.PMID 29534059.
  12. ^abHicks, J. F.; Brinkman, D. L.; Nichols, D. J.; Watabe, M. (1999)."Paleomagnetic and palynologic analyses of Albian to Santonian strata at Bayn Shireh, Burkhant, and Khuren Dukh, eastern Gobi Desert, Mongolia".Cretaceous Research.20 (6):829–850.Bibcode:1999CrRes..20..829H.doi:10.1006/cres.1999.0188.Archived from the original on 2022-05-06. Retrieved2020-01-26.
  13. ^Kurumada, Y.; Aoki, S.; Aoki, K.; Kato, D.; Saneyoshi, M.; Tsogtbaatar, K.; Windley, B. F.; Ishigaki, S. (2020)."Calcite U–Pb age of the Cretaceous vertebrate-bearing Bayn Shire Formation in the Eastern Gobi Desert of Mongolia: usefulness of caliche for age determination".Terra Nova.32 (4):246–252.Bibcode:2020TeNov..32..246K.doi:10.1111/ter.12456.
  14. ^abAverianov, Alexander;Sues, Hans-Dieter (April 2012)."Correlation of Late Cretaceous continental vertebrate assemblages in Middle and Central Asia".Journal of Stratigraphy.36 (2):462–485.hdl:10088/19377.
  15. ^Samoilov, V. S.; Benjamini, C. (1996). "Geochemical features of dinosaur remains from the Gobi Desert, South Mongolia".PALAIOS.11 (6):519–531.Bibcode:1996Palai..11..519S.doi:10.2307/3515188.JSTOR 3515188.
  16. ^Perle, A.; Norell, M. A.; Clark, J. (1999)."A new maniraptoran Theropod –Achillobator giganticus (Dromaeosauridae) – from the Upper Cretaceous of Burkhant, Mongolia".Contributions from the Geology and Mineralogy Chair, National Museum of Mongolia (101):1–105.OCLC 69865262.
  17. ^Kobayashi, Yoshitsugu; Zelenitsky, Darla K.; Fiorillo, Anthony R.; Chinzorig, Tsogtbaatar (2025-03-25)."Didactyl therizinosaur with a preserved keratinous claw from the Late Cretaceous of Mongolia".iScience.28 (4).Bibcode:2025iSci...28k2141K.doi:10.1016/j.isci.2025.112141.ISSN 2589-0042.PMC 12131255.
  18. ^Voris, Jared T.; Zelenitsky, Darla K.; Kobayashi, Yoshitsugu; Modesto, Sean P.; Therrien, François; Tsutsumi, Hiroki; Chinzorig, Tsogtbaatar; Tsogtbaatar, Khishigjav (2025-06-11). "A new Mongolian tyrannosauroid and the evolution of Eutyrannosauria".Nature.doi:10.1038/s41586-025-08964-6.
  19. ^Kobayashi, Y.; Barsbold, R. (2005). "Reexamination of a primitive ornithomimosaur,Garudimimus brevipes Barsbold, 1981 (Dinosauria: Theropoda), from the Late Cretaceous of Mongolia".Canadian Journal of Earth Sciences.42 (9):1501–1521.Bibcode:2005CaJES..42.1501K.doi:10.1139/e05-044.hdl:2115/14579.
  20. ^Park, J. (2020). "Additional skulls ofTalarurus plicatospineus (Dinosauria: Ankylosauridae) and implications for paleobiogeography and paleoecology of armored dinosaurs".Cretaceous Research.108 e104340.Bibcode:2020CrRes.10804340P.doi:10.1016/j.cretres.2019.104340.S2CID 212423361.
  21. ^Sereno, P. C. (2000)."The fossil record, systematics and evolution of pachycephalosaurs and ceratopsians from Asia"(PDF).The Age of Dinosaurs in Russia and Mongolia. Cambridge University Press. pp. 489–491.Archived(PDF) from the original on 2019-12-25. Retrieved2020-01-26.
  22. ^Watabe, M.; Tsogtbaatar, K.; Sullivan, R. M. (2011)."A new pachycephalosaurid from the Baynshire Formation (Cenomanian-late Santonian), Gobi Desert, Mongolia"(PDF).Fossil Record 3. New Mexico Museum of Natural History and Science, Bulletin.53:489–497.Archived(PDF) from the original on 2015-09-24. Retrieved2020-01-26.
  23. ^Tsogtbaatar, K.; Weishampel, D. B.; Evans, D. C.; Watabe, M. (2019)."A new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous Baynshire Formation of the Gobi Desert (Mongolia)".PLOS ONE.14 (4) e0208480.Bibcode:2019PLoSO..1408480T.doi:10.1371/journal.pone.0208480.PMC 6469754.PMID 30995236.
  24. ^abKsepka, D. T.; Norell, M. A. (2006)."Erketu ellisoni, a long-necked sauropod from Bor Guvé (Dornogov Aimag, Mongolia)"(PDF).American Museum Novitates (3508):1–16.doi:10.1206/0003-0082(2006)3508[1:EEALSF]2.0.CO;2.S2CID 86032547.Archived(PDF) from the original on 2021-08-31. Retrieved2020-01-26.
  25. ^Turner, A. H. (2015)."A Review ofShamosuchus andParalligator (Crocodyliformes, Neosuchia) from the Cretaceous of Asia".PLOS One.10 (2) e0118116.Bibcode:2015PLoSO..1018116T.doi:10.1371/journal.pone.0118116.PMC 4340866.PMID 25714338.
Avemetatarsalia
Pterosauria
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Preondactylia
Caviramidae?
Austriadraconidae
Raeticodactylidae
Eudimorphodontidae
Dimorphodontidae
Campylognathoididae
Rhamphorhynchidae
Scaphognathidae?
Pterodactylomorpha
    • see below↓
Campylognathoides liasicus

Scaphognathus crassirostris

Dorygnathus banthensis
Darwinoptera
Wukongopteridae
Anurognathidae
Pterodactyloidea
Lophocratia
    • see below↓
Jeholopterus ninchengensisKryptodrakon progenitor
Germanodactylidae
Gallodactylidae
Aurorazhdarchia
Aurorazhdarchidae
Ctenochasmatidae
Eupterodactyloidea
Ornithocheiroidea
    • see below↓
Pterodactylus antiquusPlataleorhynchus streptorophorodon
Dsungaripteridae
Thalassodromidae?
Tapejaridae
Dsungaripteromorpha?
Chaoyangopteridae
Azhdarchiformes
Alanqidae?
Azhdarchidae
Pteranodontoidea
    • see below↓
Bakonydraco galaczi

Tupandactylus imperator

Quetzalcoatlus
Pteranodontia
Pteranodontidae
Nyctosauromorpha
Aponyctosauria
Nyctosauridae
Lonchodectidae
Istiodactyliformes
Mimodactylidae
Istiodactylidae
Boreopteridae
Ornithocheiridae
Targaryendraconia?
Cimoliopteridae
Targaryendraconidae
Hamipteridae?
Anhangueridae
Pteranodon longiceps

Nyctosaurus gracilis

Ludodactylus sibbicki
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