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Flora of Turkey

From Wikipedia, the free encyclopedia
(Redirected fromFlora and vegetation of Turkey)
Plants that grow in the Eurasian country
This article is about plants and fungi. For an overview of the biodiversity seeWildlife of Turkey, and for details of animals seeFauna of Turkey.
Verbascum wiedemannianum: this showy Mullein is typical of the central Anatolian steppe. Like most of the TurkishVerbascum-species it is endemic to Anatolia.

Theflora of Turkey consists of almost 10,000 species of plants, as well as a number offungi andalgae. Around 32% of Turkey's plants are found only in the country.[1] One reason for the high proportion ofendemics is thatAnatolia is both mountainous and quite fragmented.[not verified in body] The country is divided into three main floristic areas: the Mediterranean, Euro-Siberian, and Irano-Tranian area.[2] The flora of the European part of Turkey is similar to that of adjoiningGreece. The ecoregions here includeBalkan mixed forests dominated byoaks,[3] andAegean and Western Turkey sclerophyllous and mixed forests where some of the main species are oaks,strawberry tree,Greek strawberry tree,Spanish broom andlaurel.[4] The country is at a meeting point of threephytogeographical regionsMediterranean,Euro-Siberian, andIrano-Turonian.[5][6] The region played a key role in the early cultivation ofwheat, othercereals, and varioushorticultural crops.[1]

The Euro-Siberian area is a mountainous part of western Turkey. Here the flora transitions from the Mediterranean vegetation type to the Anatolian plateau. The dominant vegetation cover here is forests of oak and pine, especiallyAnatolian black pine andTurkish pine.[7] Further east is theAnatolian plateau, a largely treeless area of plains and river basins at an average altitude of 1,000 m (3,300 ft). This area is characterised by hot dry summers and cold winters. Saltsteppes and lakes are found here, as well salt-freegrassland areas,marshes and freshwater systems. Immediately around the largeLake Tuz and other saline areas,saltmarsh plants grow, and beyond this is a sharp divide, with the flora being dominated by members of thefamiliesChenopodiaceae andPlumbaginaceae.[8]

The mountainous eastern half of the country is separated floristically from the rest of the country by theAnatolian diagonal, a floral break that crosses the country from the eastern end of the Black Sea to the northeastern corner of the Mediterranean Sea. Many species found to the east of this break are not found to the west and vice versa, and about four hundred species are only found along this divide.[9] The natural vegetation in eastern Turkey is theEastern Anatolian deciduous forests; in these oaks such asBrant's oak,Lebanon oak,Aleppo oak andMount Thabor's oak predominate in open woodland withScots pine,burnet rose,dog-rose,oriental plane,alder,sweet chestnut,maple, Caucasianhoneysuckle (Lonicera caucasica) andcommon juniper.[10]

Most European species are found in Turkey.[11] The most important reasons for the high plantbiodiversity are believed to be the relatively high proportion of endemics, together with the highvariety of soils andclimate of Turkey.

In Anatolia thePleistocene glaciations only covered the highest peaks, so there are many species with small ranges. In other words: Anatolia as a whole is a big “massif de refuge”, showing all degrees of past and recent speciation.

Naturally much of the vegetation would be steppe and forest,[12] however people have cleared much forest and their animals have changed the vegetation by grazing.[13]

Diversity and endemism

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Vascular plants

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Colchicum figlalii (Ö. Varol) Parolly & Eren: This punctual endemic of Sandras Dağ, aserpentine mountain nearMuğla, was described as new to science in 1995.

A third of Turkish plant species are endemic to Turkey:[14] one reason there are so many is because the surface of Anatolia is both mountainous and quite fragmented. In fact, the Anatolian mountains resemble archipelagos like the famousGalapagos Islands. SinceDarwin we know that geographic isolation between islands or separated mountains is an important means ofspeciation, leading to high spatial diversity. For Anatolia this assumption is confirmed by concentrations of endemism on highly isolated and relatively oldmassifs such asUludağ and Ilgaz Dağ, whereas very young volcanic cones such asErciyes Dağ and Hassan Dağ are surprisingly poor in endemics.[citation needed]

Gypsum hills south ofSivas: gypsum and serpentine areas are exceptionally rich in endemic species

For a visitor from Central Europe, climatic diversity within Turkey is quite astonishing. All climatic zones present in Europe can be found in Turkey on a somewhat smaller scale. TheBlack Sea coast is humid all year round, with the highest rainfall betweenRize andHopa. South of thePontic Range there is much less rain so Central Anatolia is dry; also it is cold in the winter. Approaching the southern and western coasts, the climate turns more and more Mediterranean, with mild but very rainy winters and dry, hot summers. This simple scheme is complicated a lot by the mountainous surface of Anatolia. On the high mountains, harsh climatic conditions persist all the year round and, as of 2019[update], there areglaciers in Turkey, for example onMount Ararat.[citation needed]

Anatolia's diversity ofsoils is astonishingly high. Saline soils are quite common in the driest parts of central Anatolia; additionally, the Aras valley betweenKağızman andArmenia is full of impressive salt outlets, some pouring directly out of the mountains and thus resembling snow patches from a distance. South ofSivas and around Gürün there are extensivegypsum hills with a very special flora. A further lot of endemics have been described from the extensiveserpentine areas in South-West Anatolia, especially Sandras Dağ (Cicekbaba D.) nearKöyceğiz.[citation needed]

TheAnatolian diagonal is an ecological dividing line that runs slant-wise across central and eastern Turkey from the northeastern corner of theMediterranean Sea to the southeastern part of theBlack Sea. Many species of plants that exist west of the diagonal are not present to the east, while others found to the east are not in the west. Of 550 species analysed, 135 were found to be "eastern" and 228 "western".[15] Besides the Anatolian diagonal forming a barrier to floralbiodiversity, about four hundred species of plant are endemic to the diagonal itself.[16]

Flora ofAkdoğan Mountains,Eastern Anatolia Region. Quçan region is completelybrown soil. Other places are semi-brown and consist of different types of soil.
See also:Category:Flora of European Turkey
Heavily pastured thorn-cushion vegetation, consisting mainly ofAstragalus angustifolius.— Melendiz Daği (Niğde), c. 2000 m s.l.
Species-numbers of the most important genera in Turkey

With almost 400 species the genusAstragalus (milk-vetch, goat's-thorn;Fabaceae) has by far the most species of the Turkish flora; as historically humans have dramatically expanded its favored treeless, dry and heavily grazed habitats. But not as many asCentral Asia: the formerUSSR has twice as many. The plasticity of this genus is astonishingly high. Depending on environmental conditions a big variety of life forms evolved, ranging from tiny annuals to small woody and thorny bushes. Speciation seems to be in plain progress inAstragalus. Nearly all of its different sections consists of clusters of closely related species whose determination is one of the hardest tasks in a closer study of the Anatolian flora. One of the most successful growth forms of Turkish Astragali is the thorn cushion, which is very characteristic of the dry mountains of inner Anatolia. Such thorn cushions were not exclusively invented by many Astragali. Really striking examples ofconvergent evolution are the impressive thorn cushions ofOnobrychis cornuta, also belonging to the Fabaceae. But there are a lot of thorn cushions also inAcantholimon (Plumbaginaceae). Even someAsteraceae (in Turkey e.g.Centaurea urvillei, C. iberica) andCaryophyllaceae (e.g.Minuartia juniperina) evolved in that direction. Second in importance comesVerbascum (Scrophulariaceae) and third isCentaurea (Asteraceae). ForVerbascum Turkey evidently is the centre of distribution. Of approximately 360 species worldwide no less than 232 are to be found in Turkey, almost 80% of them being Anatolian endemics! MostVerbascum species are protected against water loss and hungry cattle by a dense cover of tree-shaped micro hairs.Centaurea species rarely have woolly hairs, but in defence against heavy grazing developed thornyphyllaries, or evolved to have no visible stem or a very short one.[citation needed]

Non-vascular plants

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There are over 700 species of moss.[17]

Fungi

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There are over 12,000 varieties of mushroom in Turkey,[18] some of which are edible.[19]

Algae

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There are over 2000 taxa of freshwater algae.[20]

Vegetation types

[edit]
This section is an excerpt fromForests in Turkey.[edit]
Autumn inYedigöller National Park

Forests cover almost a third ofTurkey. They are almost allstate-owned, and vary fromtemperate rainforest in the north-east tomaquis in the south and west.Pine,fir,oak andbeech are common.

After theglaciers retreated over ten thousand years ago, woods grew to cover most of the land which is now Turkey; however over thousands of years, many of the trees have been cut down.The country is slowly reforesting, which is beneficial forits wildlife and toabsorb carbon to help limitclimate change.

As of the mid-2020s, the main product of Turkey's forests is wood, and they are also important for recreation. Almost half the forest is badlydegraded, and the woodlands are threatened bydrought,wildfire,mining, andpests and diseases.
Biomes of Turkey[needs update]Zagros Mountain Forests & East Anatolian Steppe Bioregion[21] Black Sea, Caucasus-Anatolian Mixed Forests & Steppe Bioregion[22] Aegean Sea & East Mediterranean Mixed Forests Bioregion[23] Balkan Mixed Forests Ecoregion[24]
This section is an excerpt fromSoil in Turkey.[edit]
Soil profile with oxidation inManisa Province, Turkey

The main types ofsoil inTurkey arecalcisols,cambisols andleptosols, andfluvisols.[25]

By the mid-20th centuryerosion had reduced the amount ofarable land,[26] butthe government is combatingdesertification and erosion in various ways.[27] However, soil erosion is forecast to increase withclimate change, with about 30% occurring onagricultural land.[28] Degraded soil could be improved.[29]: 11 Soil surveys have been done at least since the 1950s,[30][31] and theMinistry of Agriculture has published soil maps.[32][33][34][35]

TheTurkish Foundation for Combating Soil Erosion is anon-governmental organization as is the Soil Science Society of Turkey.[36] A 2016 study said soil had been degraded and that there was great potential tosequester carbon.[37] There is a soil database (Toprak Bilge Sistem) but it is not public.[38] Increasingsoil organic carbon (SOC) in agricultural soils is important, and in 2017 total SOC down to 0.7 m was estimated at 9.23Pg.[39] Another estimate is slightly under 3000 tonnes/km2.[40] SOC is being measured and mapped.[41] Accumulation ofsoil organic matter depends partly on cultivation but can be hindered by aridity.[42]

Steppe grassland is mostly inCentral andsoutheast Anatolia. Above 2000m in theBlack Sea Region there isAlpine grassland.[13]

The distribution of plants uses theWorld Geographical Scheme for Recording Plant Distributions (WGSRPD). SeeList of codes used in the World Geographical Scheme for Recording Plant Distributions for its coding system. Turkey is divided into two botanical areas:

Pinus nigra forms extensive stands in the central Taurus Mts.— between Akseki and Bademli, 1360 m s.l.

The Pontic mountain range along the north Anatolian coast is a more or less continuous barrier against humid air from the Black Sea, causing high precipitation on the northern slopes of thePontus all year. Climatic conditions on the northern coast therefore resemble those in central Europe and so does the vegetation. A limited Mediterranean influence is noticeable only on a very narrow coastal strip, but almost completely missing in the northeast. In the lower forest zone oftenHornbeam (Carpinus betulus) prevails, frequently intermingled withSweet Chestnut (Castanea sativa). Further up Oriental Beech (Fagus orientalis) and/or Nordmann Fir (Abies nordmanniana) form extensive forests. Humidity becomes extremely high in Lazistan, where the Pontic barrier culminates in the nearly 4000 m highKaçkar Mountains. East ofTrabzon therefore vegetation becomes somewhat sub-tropic, with a lot of evergreens in the forest and tea plantations everywhere on the slopes.

South of the Pontic watershed the climate immediately gets drier. In the mountains firstAbies nordmanniana, but then soonPinus becomes dominant. In the western parts of Anatolia this is often Black Pine (Pinus nigra), in the east nearly exclusively Scots Pine (Pinus sylvestris). Penetrating further into the central parts of inner Anatolia leads to still dryer, wintercold conditions. Today the lower parts of central Anatolia are virtually treeless. Fields on deep alluvial soils alternate with steppe on the dryer hills. But it is still an open question where and to what degree this central Anatolian steppe is due to aridity or to humandeforestation.[43] Aridity is most pronounced aroundTuz Gölü south of Ankara and in the Aras-valley near the Armenian border. Between Kağizman andTuzluca this valley is so dry, that here and there pure salt deposits glitter like white snowfields on the bare slopes.

TheTaurus Mountains form the southern edge of the central Anatolian Plateau and are already very influenced by the Mediterranean, with a lot of snow in winter, but dry and warm summers. Climax forests are formed by Black Pine, Cilician Fir (Abies cilicica) and Lebanon Cedar (Cedrus libani). Unfortunately, there has been a lot of deforestation in the Taurus, most gravely affecting the stands of cedar. On the Aegean and Mediterranean coasts pronounced Mediterranean conditions prevail, with very hot and dry summers and very rainy winters.Antalya Province has considerably more total precipitation than, for example, the south of England (1071 mm versus 759 mm), but its seasonal distribution is completely different and the average temperature is of course much higher (18.3 °C versus 9.7°). But due to massive forest destruction hills and slopes in coastal West and South Anatolia are nowadays mostly covered withmaquis shrubland. Where fertile alluvial soils prevail, e.g. in the Cilician Plain aroundAdana, there is intense agriculture.

Mediterranean vegetation is resilient todrought.[44]

  • Picea orientalis with Abies nordmanniana and Fagus orientalis on the moist northern slopes of Kaçkar Dağ (Northeastern Pontus). Main component of the scrub between the trees is Rhododendron luteum, above Ayder, c.1700 m s.l.
    Picea orientalis withAbies nordmanniana andFagus orientalis on the moist northern slopes of Kaçkar Dağ (Northeastern Pontus). Main component of the scrub between the trees isRhododendron luteum, aboveAyder, c.1700 m s.l.
  • Scots Pine (Pinus sylvestris) is the dominant tree in the dry and cold areas of north-eastern Anatolia, southern slope of Kücükhacet Dağ (Ilgaz Dağ), c.1950 m s. l.
    Scots Pine (Pinus sylvestris) is the dominant tree in the dry and cold areas of north-eastern Anatolia, southern slope of Kücükhacet Dağ (Ilgaz Dağ), c.1950 m s. l.
  • Rest of the Anatolian steppe with Crambe tatarica (white), with fields in the background, Ahiboz, c.35 km south of Ankara, c.1000 m s.l.
    Rest of the Anatolian steppe withCrambe tatarica (white), with fields in the background, Ahiboz, c.35 km south of Ankara, c.1000 m s.l.
  • Semidesert vegetation in the Aras-valley near the Armenian border. Sparse plant cover consists mainly of drought- and salt-tolerant members of the Goosefoot Family (Chenopodiaceae), 35 km west of Tuzluca, 1110 m s.l.
    Semidesert vegetation in the Aras-valley near the Armenian border. Sparse plant cover consists mainly of drought- and salt-tolerant members of the Goosefoot Family (Chenopodiaceae), 35 km west of Tuzluca, 1110 m s.l.
  • In Turkey the cultivation of tea is confined to the almost subtropical part of the Black Sea coast around Rize, Ikizdere valley south of Rize, 200 m s.l.
    In Turkey the cultivation of tea is confined to the almost subtropical part of the Black Sea coast around Rize, Ikizdere valley south of Rize, 200 m s.l.
  • Orchards with hazelnuts (Corylus maxima) are very typical for the mountainous parts of the Anatolian Black Sea coast. East of Trabzon they give way to tea plantations, near Tirebolu (Giresun-Province), c. 20 m s. l.
    Orchards withhazelnuts (Corylus maxima) are very typical for the mountainous parts of the Anatolian Black Sea coast. East of Trabzon they give way to tea plantations, nearTirebolu (Giresun-Province), c. 20 m s. l.
  • Turkish women drying hazelnuts, Sacmalipinar Düzce Province.
    Turkish women drying hazelnuts, SacmalipinarDüzce Province.
  • Cotton grows best in hot, sunny regions but needs plenty of water, near Belek (Antalya-Province).
    Cotton grows best in hot, sunny regions but needs plenty of water, near Belek (Antalya-Province).

Origins and evolution

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As local endemics take a long time to evolve, we also have to compare the history of the central and north European mountains with the Anatolian ones. During each of theglacial periods the former were covered by thick shields of permanent ice, which destroyed most pre-glacial endemism and hindered neo-endemics from forming. Only less glaciated, peripheral areas, the so-called “massifs de refuge”, offered suitable conditions for the survival of local endemics during glacial periods.

In Anatolia thePleistocene glaciations only covered the highest peaks, so there are many species with small ranges. In other words: Anatolia as a whole is a big "massif de refuge", showing all degrees of past and recent speciation.[45][page needed]

Human impact

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See also:Climate change in Turkey § Ecosystems, andAgriculture in Turkey

Without humans the main vegatation types would be steppe and forest.[13]Rangeland in Central Anatolia wasovergrazed, andrangeland management to limitsoil erosion has been suggested.[46] There is a national biodiversity action plan to 2028,[47] and anIUCN SSC Turkey Plant Red List Authority.[48] Potentially there could be moreforest in Turkey.[49] Technology is being used to revegetate steep slopes to try to preventdesertification.[50]

Botanical resources

[edit]
  • "The Flora of Turkey and the East Aegean Islands", 9 volumes, 1965-1985, ed.P H Davis, followed by two supplementary volumes 10 (1988) & 11 (2000, Güner, A. & al.). All volumes are in English, mostly text.
  • "Resimli Türkiye Florası" (Illustrated Flora of Turkey), projected to be 30 volumes, ed. Adil Güner, released in PDF and print format, currently published (May 2024) vols. 1,2,3a,3b,4a,4b, primarily distributed viaANG. All volumes are in Turkish, with keys, descriptive text, illustrations and details and map of herbarium specimens used for each taxon.
  • "Check-list of additional taxa to the supplement of flora of Turkey I... X" are 10 (as of May 2024) supplemental lists of new taxa that have been found in Turkey (released as PDFs).
  • "Orchids of Europe, North Africa and the Middle East, ed 3", 2006, 640 pp, by Pierre Delforge (English, includes the entirety of Turkey).
  • "Endemism in Mainland Regions – Case Studies: Turkey", 2013, by Pils, p. 240-255 in: Endemism in Vascular Plants, Springer Verlag, ed. C Hobohm[1]
  • Turkish Journal Of Botany

AVCI. M. 2005. "Çeşitlilik Ve Endemizm Açısından Türkiye’nin Bitki Örtüsü-Diversity and endemism in Turkey's Vegetation", İstanbul Üniversitesi Edebiyat Fakültesi Coğrafya Bölümü Coğrafya Dergisi 13:27-55.

Information for this article was taken mainly from:Flowers of Turkey - a photo guide.- 448 pp.– EigenverlagGerhard Pils (2006).

Notes

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References

[edit]
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Flora of Asia
Sovereign states
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