| Fantails | |
|---|---|
 | |
| Grey fantail (Rhipidura albiscapa) | |
Scientific classification | |
| Domain: | Eukaryota |
| Kingdom: | Animalia |
| Phylum: | Chordata |
| Class: | Aves |
| Order: | Passeriformes |
| Family: | Rhipiduridae |
| Subfamily: | Rhipidurinae |
| Genus: | Rhipidura Vigors &Horsfield, 1827 |
| Type species | |
| Muscicapa flabellifera[1] Gmelin, JF, 1788 | |
| Species | |
Over 50, see text | |
Fantails are smallinsectivoroussongbirds of thegenusRhipidura in thefamilyRhipiduridae, native toAustralasia,Southeast Asia and theIndian subcontinent. Most of the species are about 15 to 18 cm (5.9 to 7.1 in) long, specialist aerial feeders, and named as "fantails", but the Australianwillie wagtail is a little larger, and, though still an expert hunter of insects on the wing, concentrates equally on terrestrial prey.
The truewagtails are part of the genusMotacilla in the familyMotacillidae and are not close relatives of the fantails.
The genus Rhipidura was introduced in 1827 by the naturalistsNicholas Vigors andThomas Horsfield.[2][3] Thetype species was subsequently designated asMuscicapa flabelliferaGmelin, JF, 1788 by the English zoologistGeorge Gray in 1840.[4] This is ajunior synonym ofMuscicapa fuliginosaSparrman, 1787, theNew Zealand fantail.[5] The genus name combines theAncient Greekrhipis,rhipidos meaning "fan" with "oura" meaning "tail".[6]
The fantails are small bodied (11.5–21 cm long) birds with long tails; in some species the tail is longer than the body and in most the tail is longer than the wing.[7] When the tail is folded it is rounded at the end, but when spread in display or aerial foraging it has a characteristic fan shape that gives the family its name.
Fantails adopt a hunched horizontal posture most of the time, with the wings drooped and held away from the body and the tail half cocked. There are some exceptions to this, particularly thenorthern fantail of New Guinea and theCockerell's fantail of theSolomon Islands, which have a more upright posture reminiscent of themonarch flycatchers.
The wings of fantails are tapered and have sacrificed speed for agility, making fantails highly efficient at catching insect prey. Overall the fantails are strong fliers, and some species can undertake long migrations, but the thicket fantails (sooty thicket fantail,white-bellied thicket fantail andblack thicket fantail) are very weak fliers, and need to alight regularly.
The bills of fantails are typical for aerial insect eating birds, being flat and triangular. Thegape is surrounded by two rows ofrictal bristles which are long, often as long as the bill. The bills of most species are fairly weak, limiting fantails to softer insects, although the more terrestrial willie wagtail has a stronger bill.
Theplumage of most fantails shows some variation, most species are relatively uniform with some markings.[7] A few species, such as theRennell fantail, have uniform plumage, while others have striking if sombre patterns. The colours of most species are greys, blacks, whites and browns, although a few species have yellow or even striking blue feathers. In most species there is nosexual dimorphism in plumage; the notable exception being theblack fantail ofNew Guinea where the male has all-over black plumage and the female is almost entirelyrufous. In a few species, such as theNew Zealand fantail, there exist two colourmorphs, the common pied morph and the rarer black morph (which is most common in theSouth Island).[8]
Fantails are an Australasianfamily that has spread from as far asSamoa to northern India. In the south theNew Zealand fantail ranges as far asThe Snares off New Zealand, in the eastern extent of the family has several endemic forms in westernPolynesia. There are numerous species inIndonesia, thePhilippines and in South East Asia, and the family ranges into southern China, India and theHimalayas. Some species have a widespread distribution, particularly thewillie wagtail,grey fantail,white-throated fantail andnorthern fantail; others have a highly restricted range and in the case of some insular species may be restricted to a single island. TheMussau fantail is restricted to a single island in theBismarck Archipelago, and theKadavu fantail has a similarly restricted distribution in theKadavu Group ofFiji.
Most fantails, particularly the tropical or insular forms, are sedentary and undertake no migration. Some northern and southern species undertake a variety of movements; theyellow-bellied fantail of theHimalayas is an altitudinal migrant, breeding between 1500 and 4000 metres, but moving to lower altitudes (as low as 180 m) in the winter. Some Australian fantails undertake seasonal migrations, although these show considerable variation even within individual species. Most populations of therufous fantail exhibit little migratory behaviour, but the south-eastern population movesen masse to northernQueensland andNew Guinea.
Fantails exhibit wide tastes in habitat; while the majority of species are found inrainforests fantails exist in most available habitats from deserts andmangrove forests to highly modified agricultural and urban environments. Most species are able to survive in a variety of habitats. Of all the species themangrove fantail has the most restricted habitat requirements, being entirely restricted to mangrove forests over some of its range, although it can exist 3 km away in the absence of other fantails.[9] Some of the more primitive species are generally more restricted to primary rainforest, but most other species can survive in more disturbed forest. The most adaptable species is the willie wagtail, which is abundant in every habitat type in Australia except for dense rainforest.
The behaviour of many species of fantail has not been studied, but overall the family is highly uniform in its habits. Anecdotal observations of less studied species suggest a high degree of similarity with the better studied species. Fantails are highly active birds, with several of the smaller species continuously on the move; even when perched they continue to rock back and forth, spin 180° on the spot, wag their tail from side to side or fan it. In flight they are highly agile and undertake highly aerobatic and intricate looping flights while using their fanned tail to catch insects in flight.
The majority of the diet of fantails composes of small insects and invertebrates. The larger willie wagtail is capable of tackling smallskinks, but this is exceptional. Insect prey is generally small and easily handled, but larger items sometimes need to be subdued by being banged on branches, an action that also removes the wings of larger prey items like moths.
There are two general techniques used by the family in order to obtain prey.[10] The first is known as "static searching", where the fantail will remain at a perch and watch for aerial prey which it will then sally towards and snatch from the air before returning to the perch in order to consume and resume searching. The second method used is known as "progressive searching", where the fantail moves through vegetation searching for insect prey which it gleans; the movement of the searching fantail also flushes out hidden prey which is also pursued and consumed. The willie wagtail performs a terrestrial version of this technique, pumping its tail from side to side and undertaking quick darting movements across open ground in order to flush out prey.
Fantails frequently form associations with other species in order obtain prey. Some species perch on the backs of cattle, which they use both as a vantage point and because the cattle flush up insects. This behaviour has given the willie wagtail the nickname "shepherd's companion". Fantails are often very bold around people and will approach them closely in order to capture insects flushed by them. Different species are also frequently found inmixed-species feeding flocks, travelling with other small insectivorous birds on the periphery of the flocks taking advantage of flushed prey.
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Fantails areterritorial and aggressively defend their territories from conspecifics (other members of the same species) as well as other fantail species and other flycatchers.[7] Within the territory the female selects the nesting site, these sites are often close to the previous year's nest. Breeding responsibilities, nest building, incubation and chick feeding, are shared between both sexes.
The nest, a smallcup of grass stems neatly bound together inspider silk, takes around 10 days to construct. Many species incorporate a trailing tail into the base of the nest; this possibly breaks up the shape of the nest, although little other effort is made to conceal the nest. To compensate for the high visibility of the nest fantails will aggressively defend their chicks from potential predators.
Female fantails will also distract a potential predator byappearing to be injured and luring the predator away from the nest. While the female is pretending to be injured the male may continue to attack the predator. In spite of this fantails have a generally low nesting success.
Based on theIOC's listing as of December 2023, the genus contains 61 species:[11]
Formerly, some authorities also considered the following species (or subspecies) as species within the genusRhipidura:
