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Erketu

From Wikipedia, the free encyclopedia
Sauropod genus from the Late Cretaceous

Erketu
Temporal range:Late Cretaceous,
~96–89 Ma
Skeletal reconstruction of the holotype
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Dinosauria
Clade:Saurischia
Clade:Sauropodomorpha
Clade:Sauropoda
Clade:Macronaria
Clade:Titanosauriformes
Clade:Somphospondyli
Genus:Erketu
Ksepka & Norell, 2006
Type species
Erketu ellisoni
Ksepka & Norell, 2006

Erketu (meaning "Erketü tengri") is a genus ofsomphospondylandinosaur that lived inAsia during theLate Cretaceous, roughly between 96 million and 89 million years ago. It is known from theBayanshiree Formation ofMongolia, and its fossils were found between 2002 and 2003 during field expeditions.Erketu was first described in 2006 and later on in 2010 due to some cervicals that were left behind in the expedition. This genus represents the first sauropod described from the Bayanshiree Formation. The elongated cervical vertebrae indicate that it probably had the longest neck relative to its body size of any sauropod.

Discovery and naming

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The first remains ofErketu were found back in 2002 by the American Museum of Natural History–Mongolian Academy of Sciences expedition conducted inMongolia. The team discovered the outcrops of the new locality Bor Guvé, which overlies the Khara Kuthul locality and therefore is referable to theBayan Shireh Formation. The unearthed specimen,IGM 100/1803, was found in exposure at the sediments of Bor Guvé, mainly composed bysandstone andinterbedded greysiltstones, suggesting afluvial environment that is consistent with the Bayan Shireh Formation. Collected elements are mainly represented bycervical vertebrae and postcranial remains, such as the partial rightsternum,tibia andfibula withastragalus andcalcaneum.[1]

The remains became theholotype for the new genus and speciesErketu ellisoni, first described in2006 by Daniel Ksepka andMark Norell of theAmerican Museum of Natural History. The generic name of this particular sauropod species is named after the creator god (tengri) of the Mongolianshamanism,Erketü. The specific name,ellisoni, is in honor to the American Museum of Natural History's senior principalpaleoartist, and close friend of Norell: Mick Ellison.[1] In 2003 the team returned to the site and recovered three additional cervicals that were left behind during the first field expedition, and consequently described in2010.[2]

Description

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Four cervical vertebrae from the holotype

Erketu was a relatively large sauropod, with an estimated length of 15 m (49 ft) and a weight of 5 t (11,000 lb).[3] Its neck was estimated to be twice as long as its body, which may be a record for neck to body ratio. The exact ratio is unknown, because nodorsal vertebrae ofE. ellisoni have been reported, although some hindlimb material suggests the approximate size of the body. The long neck ofErketu is the result of the individual vertebrae being greatly elongated; it is unknown if the number of cervical vertebrae was increased.Erketu is also diagnosed by bifurcate anterior cervical neural spines, another unusual trait for a titanosauriform. The preserved cervicals include theatlas,axis and C3 to C9, however, the sixth is missing, giving a total of eight preserved cervical vertebrae.[1][2]

Size comparison with a human

The preserved sternum is rather thickened at some borders and shows a very deep lateral edge, it measures about 530 mm (53 cm) long. The right hindlimb elements were nicely found in articulation and they resemble those ofGobititan. The tibia is very straight, measuring 710 mm (71 cm) in total length, it has slightly expanded ends and in comparison to that ofOpisthocoelicaudia, the anterior and distal expansions are not very developed. The fibula is longer than the tibia with a length of 750 mm (75 cm), however, it appears to be broken on the distal end of one of thetrochanters.[1]

Classification

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The phylogenetic analysis of sauropods performed by its describers indicates thatErketu is a basal somphospondylian (theclade of allmacronarians closer totitanosaurs than tobrachiosaurids), and is most closely related toTitanosauria, being a close relative ofEuhelopus.[1]

During the description ofEuropatitan in 2017, a new cladistic analysis was performed. Here,Erketu was recovered in apolytomy alongChubutisaurus within the Somphospondyli:[4]

Somphospondyli

Paleoecology

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Erketu compared to the knownDinosauria of the Bayan Shireh Formation (Erketu in brown, third from left)

The remains ofErketu are attributed to theBayan Shireh Formation,[2] which is believed to have been dominated by fluvial and lacustrine environments, such as largemeanders andlakes but also a notorious semi-arid climate.[5][6] The age is estimated to be around 95.9 ± 6.0 million to 89.6 ± 4.0 million years ago,Cenomanian toConiacian stages.[7][8] Here,Erketu lived alongside diverse dinosaur fauna, such as the theropodsAchillobator andKhankhuuluu,[9][10] the abundant hadrosauroidGobihadros,[11] heavy armored ankylosaursTalarurus andTsagantegia,[12] and the well-known therizinosaursErlikosaurus andSegnosaurus.[13] The locality ofErketu has also yielded an indeterminate/unnamed titanosaur and very abundant fossilizedfruits.[1][2] Some ecological niches were filled by several dinosaurs, such as thebrowsersErlikosaurus,Segnosaurus[13] andTsagantegia; in contrast,Talarurus was agrazer.[12]Erketu in the other hand, due to its very long neck, was likely a high browser.[3]

See also

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References

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  1. ^abcdefKsepka, D. T.; Norell, M. A. (2006)."Erketu ellisoni, a long-necked sauropod from Bor Guvé (Dornogov Aimag, Mongolia)"(PDF).American Museum Novitates (3508): 1−16.doi:10.1206/0003-0082(2006)3508[1:EEALSF]2.0.CO;2.hdl:2246/5783.
  2. ^abcdKsepka, D. T.; Norell, M. A. (2010)."The Illusory Evidence for Asian Brachiosauridae: New Material of Erketu ellisoni and a Phylogenetic Reappraisal of Basal Titanosauriformes"(PDF).American Museum Novitates (3700):1–27.doi:10.1206/3700.2.hdl:2246/6087.S2CID 86254470.
  3. ^abPaul, G. S. (2016).The Princeton Field Guide to Dinosaurs (2nd ed.). Princeton, New Jersey: Princeton University Press. pp. 223−224.ISBN 978-0-691-16766-4.
  4. ^Fernández-Baldor, F. T.; Canudo, J. I.; Huerta, P.; Moreno-Azanza, M.; Montero, D. (2017)."Europatitan eastwoodi, a new sauropod from the lower Cretaceous of Iberia in the initial radiation of somphospondylans in Laurasia".PeerJ.5 e3409.doi:10.7717/peerj.3409.PMC 5490465.PMID 28674644.
  5. ^Ishigaki, S.; Tsogtbaatar, K.; Saneyoshi, M.; Mainbayar, B.; Aoki, K.; Ulziitseren, S.; Imayama, T.; Takahashi, A.; Toyoda, S.; Bayardorj, C.; Buyantegsh, B.; Batsukh, J.; Purevsuren, B.; Asai, H.; Tsutanaga, S.; Fujii, K. (2016)."Report of the Okayama University of Science - Mongolian Institute of Paleontology and Geology Joint Expedition in 2016"(PDF).Bulletin of Research Institute of Natural Sciences (42): 33−46.
  6. ^Slowiak, J.; Szczygielski, T.; Ginter, M.; Fostowicz-Frelik, L. (2020). "Uninterrupted growth in a non‐polar hadrosaur explains the gigantism among duck‐billed dinosaurs".Palaeontology.63 (4):579–599.doi:10.1111/pala.12473.
  7. ^Danilov, I. G.; Hirayama, R.; Sukhanov, V. B.; Suzuki, S.; Watabe, M.; Vitek, N. S. (2014). "Cretaceous soft-shelled turtles (Trionychidae) of Mongolia: new diversity, records and a revision".Journal of Systematic Palaeontology.12 (7):799–832.doi:10.1080/14772019.2013.847870.S2CID 86304259.
  8. ^Kurumada, Y.; Aoki, S.; Aoki, K.; Kato, D.; Saneyoshi, M.; Tsogtbaatar, K.; Windley, B. F.; Ishigaki, S. (2020)."Calcite U–Pb age of the Cretaceous vertebrate‐bearing Bayn Shire Formation in the Eastern Gobi Desert of Mongolia: usefulness of caliche for age determination".Terra Nova.32 (4):246–252.doi:10.1111/ter.12456.
  9. ^Perle, A.; Norell, M. A.; Clark, J. (1999). "A new maniraptoran Theropod−Achillobator giganticus (Dromaeosauridae)−from the Upper Cretaceous of Burkhant, Mongolia".Contributions from the Geology and Mineralogy Chair, National Museum of Mongolia (101):1–105.OCLC 69865262.
  10. ^Voris, Jared T.; Zelenitsky, Darla K.; Kobayashi, Yoshitsugu; Modesto, Sean P.; Therrien, François; Tsutsumi, Hiroki; Chinzorig, Tsogtbaatar; Tsogtbaatar, Khishigjav (2025-06-11). "A new Mongolian tyrannosauroid and the evolution of Eutyrannosauria".Nature.doi:10.1038/s41586-025-08964-6.
  11. ^Tsogtbaatar, K.; Weishampel, D. B.; Evans, D. C.; Watabe, M. (2019)."A new hadrosauroid (Dinosauria: Ornithopoda) from the Late Cretaceous Baynshire Formation of the Gobi Desert (Mongolia)".PLOS ONE.14 (4) e0208480.Bibcode:2019PLoSO..1408480T.doi:10.1371/journal.pone.0208480.PMC 6469754.PMID 30995236.
  12. ^abPark, J. (2020). "Additional skulls of Talarurus plicatospineus (Dinosauria: Ankylosauridae) and implications for paleobiogeography and paleoecology of armored dinosaurs".Cretaceous Research.108 e104340.doi:10.1016/j.cretres.2019.104340.
  13. ^abZanno, L. E.; Tsogtbaatar, K.; Chinzorig, T.; Gates, T. A. (2016)."Specializations of the mandibular anatomy and dentition of Segnosaurus galbinensis (Theropoda: Therizinosauria)".PeerJ.4 e1885.doi:10.7717/peerj.1885.PMC 4824891.PMID 27069815.
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Erketu
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