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Equus altidens

From Wikipedia, the free encyclopedia
Extinct species of equine from the Pleistocene

Equus altidens
Temporal range:Early Pleistocene–Middle Pleistocene
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Order:Perissodactyla
Family:Equidae
Genus:Equus
Species:
E. altidens
Binomial name
Equus altidens
Von Reichenau, 1915

Equus altidens is an extinct species of equine native to western Eurasia including Europe during theEarly Pleistocene to earlyMiddle Pleistocene.

Taxonomy

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Equus altidens was first described in 1915 from remains at the earlyMiddle Pleistocene Sussenborn locality in Germany.[1] Most recent authors consider the speciesEquus marxi also known from Sussenborn to be ajunior synonym ofE. altidens. The majority of authors have suggested the species is related to other "stenonine" equines known from Early Pleistocene of Europe such asEquus stenonis,[2] which are thought to be more closely related to livingzebras andasses than to true horses. Some authors have suggested a closer relationship to asses than to zebras forE. altidens.[3] Some authors have placed the species in the subgenusSussemionus, or place it with other "stenonines" in the separate genusAllohippus.[2]

"Orce Man", a supposed archaic human fossil from the Early Pleistocene of Spain, was later determined to actually be the skull remains of a juvenile of this species.[4] However, a later discovery of a tooth confirmed the presence of archaic humans at the same site.[5]

Description

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Equus altidens is a medium-sized equine species[2] with an estimated body mass of around 338–374 kilograms (745–825 lb).[6]ST2 The species had slender limbs, the most slender among the "stenonines", including themetapodial bones, and is distinguished from other equines by a number of dental morphological features, including narrow and small crowns, and deep molarectoflexid, though the dental morphology exhibits considerable variability in a number of characters.[2]

Distribution and chronology

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Equus altidens is known from remains found across Europe, Spanning from the Iberian Peninsula to Italy and Greece, northwards to Germany and Britain, and as far eastwards as Georgia in theCaucasus.[2] Other possible records are known from Romania, Turkey, Tajikistan, Hungary, and Russia.[7] Authors have differed about the timing of the earliest appearance of the species, ranging from as late as 1.2 million years ago to as early as 1.8 million years ago,[2] with recent research supporting the earlier end of the range, with the earliest records coming from theDmanisi site in Georgia.[3] The youngest remains of the species date to around 600,000 years ago,[2] when the species like other European "stenonine" equines was replaced by early caballine true horses belonging to the speciesEquus mosbachensis.[6]

Ecology

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Dental wear analysis thatEquus altidens had an abraisive, largely grazing based diet,[2][6] though it may have seasonally engaged in mixed feeding.[2] It often co-occurred alongside another larger "stenonine" equine,Equus suessenbornensis. Evidence suggests that this species had a diet that consisted of considerably more browse thanE. altidens, suggesting dietaryniche partitioning between the two species.[6]Equus altidens is thought to have primarily inhabited arid, open habitats.[3] Isotopic analysis from the Venta Micena locality in southeast Spain dating to the Early Pleistocene, around 1.6 million years ago, suggests that at this localityEquus altidens was regularly preyed upon by sabertooth cats, including the lion sizedHomotherium, and the smaller, jaguar sizedMegantereon.[8]

Relationship with humans

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At the Fuente Nueva-3 site in Spain, during the late Early Pleistocene around 1.2 million years ago, cut marks found on bones indicate butchery ofEquus altidens by archaic humans,[9] likelyHomo antecessor.[10]

References

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  1. ^Von Reichenau, W. (1915).Beiträge zur näheren Kenntnis fossiler Pferde aus Deutschem Pleistozän, inbesondere über die Entwicklung und die Abkaustadien des Gebisses vom Hochterrassenpferd. Equus mosbachensis v. R, Darmstadt, Germany.
  2. ^abcdefghiBoulbes, Nicolas; van Asperen, Eline N. (2019-09-10)."Biostratigraphy and Palaeoecology of European Equus".Frontiers in Ecology and Evolution.7.doi:10.3389/fevo.2019.00301.ISSN 2296-701X.
  3. ^abcBernor, Raymond L.; Cirilli, Omar; Bukhsianidze, Maia; Lordkipanidze, David; Rook, Lorenzo (September 2021)."The Dmanisi Equus: Systematics, biogeography, and paleoecology".Journal of Human Evolution.158: 103051.Bibcode:2021JHumE.15803051B.doi:10.1016/j.jhevol.2021.103051.PMID 34365132.
  4. ^Moyà-Solà, Salvador; Köhler, Meike (July 1997)."The Orce skull: anatomy of a mistake".Journal of Human Evolution.33 (1):91–97.Bibcode:1997JHumE..33...91M.doi:10.1006/jhev.1997.0121.PMID 9236079.
  5. ^Toro-Moyano, Isidro; Martínez-Navarro, Bienvenido; Agustí, Jordi; Souday, Caroline; Bermúdez de Castro, José María; Martinón-Torres, María; Fajardo, Beatriz; Duval, Mathieu; Falguères, Christophe; Oms, Oriol; Parés, Josep Maria; Anadón, Pere; Julià, Ramón; García-Aguilar, José Manuel; Moigne, Anne-Marie (2013)."The oldest human fossil in Europe, from Orce (Spain)".Journal of Human Evolution.65 (1):1–9.Bibcode:2013JHumE..65....1T.doi:10.1016/j.jhevol.2013.01.012.hdl:10072/338463.PMID 23481345.
  6. ^abcdSaarinen, Juha; Cirilli, Omar; Strani, Flavia; Meshida, Keiko; Bernor, Raymond L. (2021-02-26)."Testing Equid Body Mass Estimate Equations on Modern Zebras—With Implications to Understanding the Relationship of Body Size, Diet, and Habitats of Equus in the Pleistocene of Europe".Frontiers in Ecology and Evolution.9.doi:10.3389/fevo.2021.622412.hdl:11573/1505920.ISSN 2296-701X.
  7. ^Koufos, George D.; Vlachou, Theodora D.; Gkeme, Anastasia G. (2022), Vlachos, Evangelos (ed.),"The Fossil Record of Equids (Mammalia: Perissodactyla: Equidae) in Greece",Fossil Vertebrates of Greece Vol. 2, Cham: Springer International Publishing, pp. 351–401,doi:10.1007/978-3-030-68442-6_12,ISBN 978-3-030-68441-9, retrieved2024-12-15
  8. ^Palmqvist, P.; Perez-Claros, J. A.; Janis, C. M.; Figueirido, B.; Torregrosa, V.; Grocke, D. R. (November 2008)."Biogeochemical and Ecomorphological Inferences On Prey Selection and Resource Partitioning Among Mammalian Carnivores In An Early Pleistocene Community".PALAIOS.23 (11):724–737.Bibcode:2008Palai..23..724P.doi:10.2110/palo.2007.p07-073r.ISSN 0883-1351.
  9. ^Espigares, Ma Patrocinio; Martínez-Navarro, Bienvenido; Palmqvist, Paul; Ros-Montoya, Sergio; Toro, Isidro; Agustí, Jordi; Sala, Robert (May 2013)."Homo vs. Pachycrocuta: Earliest evidence of competition for an elephant carcass between scavengers at Fuente Nueva-3 (Orce, Spain)".Quaternary International.295:113–125.Bibcode:2013QuInt.295..113E.doi:10.1016/j.quaint.2012.09.032.
  10. ^Moyano, Isidro Toro; Barsky, Deborah; Cauche, Dominique; Celiberti, Vincenzo; Grégoire, Sophie; Lebegue, Frédéric; Moncel, Marie Hélène; de Lumley, Henry (October 2011)."The archaic stone tool industry from Barranco León and Fuente Nueva 3, (Orce, Spain): Evidence of the earliest hominin presence in southern Europe".Quaternary International.243 (1):80–91.Bibcode:2011QuInt.243...80M.doi:10.1016/j.quaint.2010.12.011.
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