Cockatoos are recognisable by their prominentcrests and curvedbills. Theirplumage is generally lesscolourful than that of other parrots, being mainly white, grey, or black and often with coloured features in the crest, cheeks, or tail. On average, they are larger than other parrots; however, thecockatiel, the smallest cockatoo species, is medium-sized.[3] The phylogenetic position of the cockatiel remains unresolved, except that it is one of the earliest offshoots of the cockatoo lineage. The remaining species are in two main clades. The five large black-coloured cockatoos of the genusCalyptorhynchus form one branch. The second and larger branch is formed by the genusCacatua, comprising 12 species of white-plumaged cockatoos and three monotypic genera that branched off earlier, namely the pink and greygalah, the mainly greygang-gang cockatoo and the large black-plumagedpalm cockatoo.
Cockatoos prefer to eat seeds,tubers,corms, fruit, flowers, and insects. They often feed in large flocks, particularly when ground-feeding. Cockatoos aremonogamous and nest intree hollows. Some cockatoo species have been adversely affected byhabitat loss, particularly from a shortage of suitable nesting hollows after large, mature trees are cleared; conversely, some species have adapted well to human changes and are considered agriculturalpests.
Cockatoos are popular birds inaviculture, but their needs are difficult to meet. The cockatiel is the easiest cockatoo species to maintain and is by far the most frequently kept in captivity. White cockatoos are more commonly found in captivity than black cockatoos. Illegal trade in wild-caught birds contributes to the decline of some cockatoo species in the wild.
Look upcockatoo in Wiktionary, the free dictionary.
The word cockatoo dates from the 17th century and is derived fromDutchkaketoe, which in turn is from theIndonesian/Malaykakatua. Seventeenth-century variants include cacato, cockatoon and crockadore, and cokato, cocatore and cocatoo were used in the 18th century.[4][5] The derivation has also been used for the family andgeneric names Cacatuidae andCacatua, respectively.[6]
InAustralian slang or vernacular speech, a person who is assigned to keep watch while others undertake clandestine or illegal activities, particularly gambling, may be referred to as a "cockatoo".[7] Proprietors of small agricultural undertakings are often jocularly or slightly disparagingly referred to as "cocky farmers".[8]
Cacatua – white cockatoos and corellas (13 species)
Genus levelcladogram of the cockatoos based on a 2023 molecular phylogenetic study by Brian Smith and collaborators.[9] The number of species in each genus is from the list maintained byFrank Gill,Pamela Rasmussen and David Donsker on behalf of the International Ornithological Committee (IOC), now theInternational Ornithologists' Union.[10]
The cockatoos were first defined as a subfamilyCacatuinae within the parrot familyPsittacidae by English naturalistGeorge Robert Gray in 1840, withCacatua the first listed and type genus.[11] This group has alternately been considered as either a full or subfamily by different authorities. American ornithologistJames Lee Peters in his 1937Check-list of Birds of the World andSibley andMonroe in 1990 maintained it as a subfamily, while parrot expertJoseph Forshaw classified it as a family in 1973.[12] Subsequent molecular studies indicate that the earliest offshoot from the original parrot ancestors were theNew Zealand parrots of the family Strigopidae, and following this the cockatoos, now a well-defined group orclade, split off from the remaining parrots, which then radiated across the Southern Hemisphere and diversified into the many species of parrots,parakeets,macaws,lories, lorikeets,lovebirds and othertrue parrots of the superfamilyPsittacoidea.[13][14][15][16][17][18][19][20][21]
The relationships among various cockatoo genera are largely resolved,[13][14][16][22][23][24] although the placement of thecockatiel (Nymphicus hollandicus) at the base of the cockatoos remains uncertain. The cockatiel is alternatively placed basal to all other cockatoo species,[13][23] as the sister taxon to the black cockatoo species of the genusCalyptorhynchus[16][22][24] or as the sister taxon to a clade consisting of the white and pink cockatoo genera as well as thepalm cockatoo.[14] The remaining species are within two main clades, one consisting of the black species of the genusCalyptorhynchus while the other contains the remaining species.[13][14][16][23][24] According to most authorities, the second clade includes the black palm cockatoo (Probosciger), the grey and reddishgalah (Eolophus), and thegang-gang cockatoo (Callocephalon),[13][14][16][23] althoughProbosciger is sometimes placed basal to all other species.[22] The remaining species are mainly white or slightly pinkish and all belong to the genusCacatua.[13][14][15][16][23] The generaEolophus andCacatua arehypomelanistic. The genusCacatua is further subdivided into thesubgeneraLicmetis, commonly known ascorellas, andCacatua, referred to as white cockatoos.[13][22][23][24][25] Confusingly, the term "white cockatoo" has also been applied to the whole genus.[26][27] The five cockatoo species of the genusCalyptorhynchus are commonly known as black cockatoos,[25] and are divided into two subgenera—Calyptorhynchus andZanda. The former group aresexually dichromatic, with the females having prominently barred plumage.[28] The two are also distinguished by differences in thefood-begging calls of juveniles.[29]
Thefossil record of cockatoos is even more limited than that of parrots in general, with only one truly ancient cockatoo fossil known: a species ofCacatua, most probably subgenusLicmetis, found in EarlyMiocene (16–23 millionyears ago) deposits ofRiversleigh, Australia.[30] Although fragmentary, the remains are similar to thewestern corella and the galah.[31] InMelanesia,subfossil bones ofCacatua species which apparently did not survive early human settlement have been found onNew Caledonia andNew Ireland.[32][33] The bearing of these fossils on cockatoo evolution and phylogeny is fairly limited, although the Riversleigh fossil does allow tentative dating of the divergence of subfamilies.
There are about 44 different birds in the cockatoo family Cacatuidae including recognized subspecies. The current subdivision of this family is as follows:[Note 1]
The cockatoos are generally medium to large parrots of stocky build, which range from 30–60 cm (12–24 in) in length and 300–1,200 g (0.66–2.65 lb) in weight; however, one species, the cockatiel, is considerably smaller and slimmer than the other species, being 32 cm (13 in) long (including its long pointed tail feathers) and 80–100 g (2.8–3.5 oz) in weight.[7][34][36][37] The movableheadcrest, which is present in all cockatoos, is spectacular in many species;[38] it is raised when the bird lands from flying or when it is aroused.[39] Cockatoos share many features with other parrots, including the characteristic curved beak shape and azygodactyl foot, with the two middle toes forward and the two outer toes backward.[40] They differ in the presence of an erectile crest and their lack of theDyck texture feather composition which causes the bright blues and greens seen in true parrots.[37]
Like other parrots, cockatoos have short legs, strong claws, a waddling gait[37] and often use their strong bill as a third limb when climbing through branches. They generally have long broad wings used in rapid flight, with speeds up to 70 km/h (43 mph) being recorded for galahs.[41] The members of the genusCalyptorhynchus and larger white cockatoos, such as thesulphur-crested cockatoo and thepink cockatoo, have shorter, rounder wings and a more leisurely flight.[41]
Cockatoos have a large bill, which is kept sharp by rasping the two jaws together when resting. The bill is complemented by a large muscular tongue which helps manipulate seeds inside the bill so that they can be de-husked before eating.[7] During the de-husking, the lower jaw applies the pressure, the tongue holds the seed in place and the upper jaw acts as an anvil. The eye region of the skull is reinforced to support muscles which move the jaws sideways.[37] The bills of male cockatoos are generally slightly larger than those of their female counterparts, but this size difference is quite marked in the palm cockatoo.[42]
Theplumage of the cockatoos is less brightly coloured than that of the other parrots, with species generally being either black, grey or white. Many species have smaller areas of colour on their plumage, often yellow, pink and red, usually on the crest or tail.[43] The galah and Major Mitchell's cockatoo are more broadly coloured in pink tones.[44] Several species have a brightly coloured bare area around the eye and face known as a periophthalmic ring; the large red patch of bare skin of the palm cockatoo is the most extensive and covers some of the face, while it is more restricted in some other species of white cockatoo, notably thecorellas andblue-eyed cockatoo.[44] The plumage of males and females is similar in most species. The plumage of the female cockatiel is duller than the male, but the most markedsexual dimorphism occurs in the gang-gang cockatoo and the two species of black cockatoos in the subgenusCalyptorhynchus, namely thered-tailed andglossy black cockatoos.[42] Theiris colour differs in a few species, being pink or red in the female galah and the pink cockatoo and red-brown in some other female white cockatoo species. The males all have dark brown irises.[42]
Awhite cockatoo's left foot clasping aviary bars showing claws, scaly skin andzygodactyly—the middle two toes forward and the outer two toes backward.
Cockatoos maintain their plumage with frequentpreening throughout the day. They remove dirt and oil and realign feather barbs by nibbling their feathers. They also preen other birds' feathers that are otherwise hard to get at. Cockatoos produce preen-oil from agland on their lower back and apply it by wiping their plumage with their heads or already oiled feathers.Powder-down is produced by specialised feathers in the lumbar region and distributed by the preening cockatoo all over the plumage.[45]
Moulting is very slow and complex. Black cockatoos appear to replace theirflight feathers one at a time, their moult taking two years to complete. This process is much shorter in other species, such as the galah andlong-billed corella, which each take around six months to replace all their flight feathers.[45]
Thevocalisations of cockatoos are loud and harsh.[7] They serve a number of functions, including allowing individuals to recognize one another, alerting others of predators, indicating individual moods, maintaining the cohesion of a flock and as warnings when defending nests. The use of calls and number of specific calls varies by species; the Carnaby's black cockatoo has as many as 15 types of call, whereas others, such as the pink cockatoo, have fewer. Some, like the gang-gang cockatoo, are comparatively quiet but do have softer growling calls when feeding. In addition to vocalisations, palm cockatoos communicate over large distances by drumming on a dead branch with a stick.[46] Cockatoo species also make a characteristic hissing sound when threatened.[39]
TheTanimbar corella is restricted to the islands of Tanimbar in Indonesia; a few feral escapees are found inSingapore.
Cockatoos have a much more restricted range than the true parrots, occurring naturally only inAustralia,Indonesia, thePhilippines, and some Pacific regions.[7] Eleven of the 21 species exist in the wild only in Australia, while seven species occur only in the islands of thePhilippines, Indonesia,Papua New Guinea and theSolomon Islands. No cockatoo species are found inBorneo, despite their presence on nearbyPalawan andSulawesi or many Pacific islands,[47] although fossil remains have been recorded fromNew Caledonia.[32]
Three species occur in both New Guinea and Australia.[48] Some species have widespread distributions, with the galah, for example, occurring over most of Australia, whereas other species have tiny distributions, confined to a small part of the continent, such as theBaudin's black cockatoo of Western Australia or to a small island group, such as theTanimbar corella, which is restricted to theTanimbar Islands of Indonesia. Some cockatoos have been introduced accidentally to areas outside their natural range such as New Zealand, Singapore, andPalau,[49] while two Australian corella species have been introduced to parts of the continent where they are not native.
Cockatoos occupy a wide range of habitats from forests in subalpine regions to mangroves. However, no species is found in all types of habitat.[50] The most widespread species,[7] such as the galah and cockatiel,[51] are open-country specialists that feed on grass seeds.[7] They are often highly mobile fast flyers and are nomadic. Flocks of birds move across large areas of the inland, locating and feeding on seed and other food sources. Drought may force flocks from more arid areas to move further into farming areas.[51] Other cockatoo species, such as the glossy black cockatoo, inhabit woodlands, rainforests, shrublands and even alpine forests. Thered-vented cockatoo inhabits mangroves and its absence from northernLuzon may be related to the lack of mangrove forests there.[52] Forest-dwelling cockatoos are generally sedentary, as the food supply is more stable and predictable.[53] Several species have adapted well to human modified habitats and are found in agricultural areas and even busy cities.[54]
Cockatoos arediurnal and require daylight to find their food.[7] They are not early risers, instead waiting until the sun has warmed their roosting sites before feeding. All species are generally highly social and roost, forage and travel in colourful and noisyflocks. These vary in size depending on availability of food; in times of plenty, flocks are small and number a hundred birds or less, while in droughts or other times of adversity, they may swell up to contain thousands or even tens of thousands of birds; one record from theKimberley noted a flock of 32,000little corellas. Species that inhabit open country form larger flocks than those of forested areas.[55]
Some species require roosting sites that are located near drinking sites; other species travel great distances between the roosting and feeding sites.[56] Cockatoos have several characteristic methods of bathing; they may hang upside down or fly about in the rain or flutter in wet leaves in the canopy.[39] Cockatoos have a preferred "footedness" analogous to human handedness. Most species are left-footed with 87–100% of individuals using their left feet to eat, but a few species favor their right foot.[57]
Cockatoos aremonogamous breeders, withpair bonds that can last many years. Many birds pair up in flocks before they reach sexual maturity and delay breeding for a year at least. Females breed for the first time anywhere from three to seven years of age and males are often older. Sexual maturity is delayed so birds can develop the skills for raising and parenting young, which is prolonged compared with other birds; the young of some species remain with their parents for up to a year.[58] Cockatoos may also displaysite fidelity, returning to the same nesting sites in consecutive years.[46] Courtship is generally simple, particularly for established pairs, with the black cockatoos alone engaging incourtship feeding. Established pairs do engage in preeningeach other, but all forms of courtship drop off afterincubation begins, possibly due to the strength of the pair-bond.[59]
Like most parrots, the cockatoos are cavitynesters, nesting in holes in trees,[60] which they are unable to excavate themselves.[61] These hollows are formed from decay or destruction of wood by branches breaking off, fungi or insects such as termites or evenwoodpeckers where their ranges overlap.[62] In many places these holes are scarce and the source of competition, both with other members of the same species and with other species and types of animal.[63] In general, cockatoos choose hollows only a little larger than themselves, hence different-sized species nest in holes of corresponding (and different) sizes. If given the opportunity, cockatoos prefer nesting over 7 or 8 metres (23 or 26 ft) above the ground[62] and close to water and food.[64]
The nesting hollows are lined with sticks, wood chips and branches with leaves. The eggs of cockatoos are oval and initially white, as their location makes camouflage unnecessary.[65] However, they do become discoloured over the course of incubation. They range in size from 55 mm × 37 mm (2.2 in × 1.5 in) in the palm and red-tailed black cockatoos, to 26 mm × 19 mm (1.02 in × 0.75 in) in the cockatiel.[65]Clutch size varies within the family, with the palm cockatoo and some other larger cockatoos laying only a single egg and the smaller species laying anywhere between two and eight eggs. Food supply also plays a role inclutch size.[66] Some species can lay a second clutch if the first fails.[67] Around 20% of eggs laid are infertile.[68] The cockatoos' incubation and brooding responsibilities may either be undertaken by the female alone in the case of the black cockatoos or shared amongst the sexes as happens in the other species. In the case of the black cockatoos, the female is provisioned by the male several times a day. The young of all species are born covered in yellowish down, bar the palm cockatoo, whose young are born naked.[69] Cockatoo incubation times are dependent on species size, with the smaller cockatiels having a period of around 20 days and the larger Carnaby's black cockatoo incubating its eggs for up to 29 days.[7]
Thenestling period also varies by species size, with larger species having longer nestling periods. It is also affected by season and environmental factors and by competition with siblings in species with clutch sizes greater than one. Much of what is known about the nestling period of some species is dependent on aviary studies – aviary cockatiels canfledge after 5 weeks and the large palm cockatoos after 11 weeks.[7] During this period, the young become covered in juvenile plumage while remaining in the hollow. Wings and tail feathers are slow to grow initially but more rapid as the primary feathers appear. Nestlings quickly reach about 80–90% of adult weight about two-thirds of the time through this period, plateauing before they leave the hollow; they fledge at this weight with wing and tail feathers still to grow a little before reaching adult dimensions.[70] Growth rate of the young, as well as numbers fledged, are adversely impacted by reduced food supply and poor weather conditions.[71]
Wildlong-billed corellas inPerth. The bird on the right is using its long beak to dig for food in short grass.
Cockatoos are versatile feeders and consume a range of mainly vegetable food items. Seeds form a large part of the diet of all species; these are opened with their large and powerful bills. The galahs, corellas and some of the black cockatoos feed primarily on the ground; others feed mostly in trees.[7] The ground-feeding species tend to forage in flocks, which form tight, squabbling groups where seeds are concentrated and dispersed lines where food is more sparsely distributed;[72] they also prefer open areas where visibility is good. Thewestern and long-billed corellas have elongated bills to excavate tubers and roots and the pink cockatoo walks in a circle around the doublegee (Emex australis) to twist out and remove the underground parts.[73]
Many species forage for food in the canopy of trees, taking advantage ofserotiny (the storage of a large supply of seed in cones or gumnuts by plant genera such asEucalyptus,Banksia andHakea), a natural feature of the Australian landscape in dryer regions. These woody fruiting bodies are inaccessible to many species and harvested in the main by parrots, cockatoos and rodents in more tropical regions. The larger cones can be opened by the large bills of cockatoos but are too strong for smaller animals.[74] Many nuts and fruits lie on the end of small branches which are unable to support the weight of the foraging cockatoo, which instead bends the branch towards itself and holds it with its foot.[75]
While some cockatoos are generalists taking a wide range of foods, others are specialists. The glossy black cockatoo specialises in the cones of trees of the genusAllocasuarina, preferring a single species,A. verticillata. It holds the cones in its foot and shreds them with its powerful bill before removing the seeds with its tongue.[76] Some species take large numbers of insects, particularly when breeding; in fact the bulk of theyellow-tailed black cockatoo's diet is made up of insects. The large bill is used in order to extract grubs and larvae from rotting wood. The amount of time cockatoos have to spend foraging varies with the season.[75] During times of plenty they may need to feed for only a few hours in the day, in the morning and evening, then spend the rest of the day roosting or preening in trees, but during the winter most of the day may be spent foraging. The birds have increased nutritional requirements during the breeding season, so they spend more time foraging for food during this time. Cockatoos have largecrops, which allow them to store and digest food for some time after retiring to a tree.[77]
Theperegrine falcon andlittle eagle have been reported taking galahs and thewedge-tailed eagle has been observed killing a sulphur-crested cockatoo.[78] Eggs and nestlings are vulnerable to many hazards. Various species of monitor lizard (Varanus) are able to climb trees and enter hollows. Other predators recorded include thespotted wood owl on Rasa Island in the Philippines; theamethystine python,black butcherbird and rodents including thegiant white-tailed rat[79] in Cape York; andbrushtail possum on Kangaroo Island. Furthermore, galahs and little corellas competing for nesting space with the glossy black cockatoo on Kangaroo Island have been recorded killing nestlings of the latter species there. Severe storms may also flood hollows drowning the young and termite or borer activity may lead to the internal collapse of nests.[80]
Like other parrots, cockatoos can be afflicted bypsittacine beak and feather disease (PBFD). The viral infection causes feather loss and beak malformation and reduces the bird's overall immunity. Particularly prevalent in sulphur-crested cockatoos, little corellas and galahs, it has been recorded in 14 species of cockatoo to date. Although unlikely to significantly impact on large, healthy populations of birds in the wild, PBFD may pose a high risk to smaller stressed populations.[81]
Awhite cockatoo and a sulphur-crested cockatoo were found to be infected with the protozoonHaemoproteus and another sulphur-crested cockatoo had themalaria parasitePlasmodium on analysis of faecal samples at Almuñecar ornithological garden inGranada in Spain.[82] Like amazon parrots and macaws, cockatoos frequently develop cloacalpapillomas. The relationship with malignancy is unknown, as is the cause, although a parrotpapilloma virus has been isolated from agrey parrot with the condition.[83]
Cockatoos have been shown to learn new skills through social interaction. InNew South Wales, researchers andcitizen scientists were able to track the spread of lid-flipping skills as cockatoos learned from each other to open garbage bins. Bin-opening spread more quickly to neighbouring suburbs than suburbs further away. In addition, birds in different areas developed their own variants for accomplishing the complex task.[84][85]
Human activities have had positive effects on some species of cockatoo and negative effects on others. Many species of open country have benefited greatly fromanthropogenic changes to the landscape, with the great increase in reliable seed food sources, and available water contributing to their survival, as well as their adaption to a diet including foreign foodstuffs. This benefit appears to be restricted to Australian species, as cockatoos favouring open country outside Australia have not become more abundant. Predominantly forest-dwelling species have suffered greatly fromhabitat destruction; in the main, they appear to have a more specialised diet and have not been able to incorporate exotic food into their diet. A notable exception is the yellow-tailed black cockatoo in eastern Australia.[86]
Several species of cockatoo can be serious agriculturalpests.[87] They are sometimescontrolled by shooting,poisoning or capture followed bygassing. Non-lethal damage mitigation methods used include scaring, habitat manipulation and the provision of decoy food dumps or sacrifice crops to distract them from the main crop. They can be a nuisance in urban areas due to destruction of property. They maintain their bills in the wild by chewing on wood, but in suburbia, they may chew outdoor furniture, door and window frames;[54] soft decorative timbers such aswestern redcedar are readily demolished.[88] Birds may also target external wiring and fixtures such as solar water heaters,[54] television antennae and satellite dishes.[88] A business in central Melbourne suffered as sulphur-crested cockatoos repeatedly stripped the silicone sealant from the plate glass windows.[89] Galahs and red-tailed black cockatoos have stripped electrical cabling in rural areas and tarpaulin is targeted elsewhere.[89] Outside Australia, the Tanimbar corella is a pest onYamdena Island where it raids maize crops.[90]
Sulphur-crested cockatoos damaging the Sturt Mall shopping centre facade, made of polystyrene
In 1995 theGovernment of thestate ofVictoria published a report on problems caused by long-billed corellas, sulphur-crested cockatoos and galahs, three species which, along with the little corella, have large and growing populations, having benefited from anthropogenic changes to the landscape. Subsequent to the findings and publication of the report, these three species were declared unprotected by aGovernor in Council Order under certain conditions and are allowed to be killed where serious damage is being caused by them to trees, vineyards, orchards, recreational reserves and commercial crops.[91] Damage covered by the report included not only that to cereal crops, fruit and nut orchards and some kinds of vegetable crops but also to houses and communications equipment.[92] The little corella is a declared pest of agriculture in Western Australia, where it is an aviculturally introduced species. The birds damagesorghum, maize,sunflower,chickpeas and other crops. They alsodefoliateamenity trees in parks and gardens, dig for edible roots andcorms onsports grounds andrace tracks, as well as chew wiring and household fittings.[93] In South Australia, where flocks can number several thousand birds and the species is listed as unprotected, they are accused of defoliatingred gums and other native or ornamental trees used for roosting, damagingtarpaulins on grain bunkers, wiring and flashing on buildings, taking grain from newly seededpaddocks and creating anoise nuisance.[94]
Several rare species and subspecies, too, have been recorded as causing problems. TheCarnaby's black cockatoo, athreatenedWestern Australianendemic, has been considered a pest in pineplantations where the birds chew off the leading shoots of growing pine trees, resulting in bent trunks and reduced timber value.[95] They are also known to damage nut and fruit crops,[96] and have learnt to exploitcanola crops.[97] The Baudin's black cockatoo, also endemic to the south-west of Western Australia, can be a pest in apple and pearorchards where it destroys the fruit to extract the seeds.[95]Muir's corella, the nominate subspecies of the western corella, is also a declared pest of agriculture in Western Australia, as well as being nationallyvulnerable and listed under state legislation as being "rare or likely to become extinct".[98]
The principal threats to cockatoos arehabitat loss and thewildlife trade. All cockatoos are dependent on trees for nesting and are vulnerable to their loss; in addition many species have specialised habitat requirements or live on small islands and have naturally small ranges, making them vulnerable to the loss of these habitats.[104] Cockatoos are popular as pets and the capture and trade has threatened some species; between 1983 and 1990, 66,654 recordedsalmon-crested cockatoos were exported fromIndonesia, a figure that does not include the number of birds caught for the domestic trade or that were exported illegally.[105] The capture of many species has subsequently been banned but the trade continues illegally. Birds are put in crates or bamboo tubing and conveyed on boats out of Indonesia and the Philippines.[106] Not only are the rare species smuggled out of Indonesia but also common and rare cockatoos alike are smuggled out of Australia; birds are sedated, covered in nylon stockings and packed intoPVC tubing which is then placed in unaccompanied luggage on international flights.[106] Mortality is significant (30%) and eggs, more easily hidden on the bodies of smugglers on flights, are increasingly smuggled instead. Trafficking is thought to be run by organised gangs, who also trade Australian species for overseas species such as macaws coming the other way.[107]
All species of cockatoo except the cockatiel are protected by the Convention on International Trade in Endangered Species of Wild Fauna and Flora (CITES), which restricts import and export of wild-caught parrots to special licensed purposes. Five cockatoo species (including all subspecies)—the Tanimbar corella (Cacatua goffiniana), red-vented cockatoo (Cacatua haematuropygia), Moluccan cockatoo (Cacatua moluccensis), yellow-crested cockatoo (Cacatua sulphurea) and palm cockatoo (Probosciger aterrimus)—are protected on the CITES Appendix I list. With the exception of the cockatiel, all remaining cockatoo species are protected on the CITES Appendix II list.[108]
A wing-clipped pet.Salmon-crested cockatoos, also known as Moluccan cockatoos,[109] are the largest white-coloured cockatoo species at about 52 cm (20 in) long and weighing 775–935 grams.[110] Cockatoos can be noisy and demanding pets.
Kept for their appearance, intelligence, and engaging personalities,[7] cockatoos can nonetheless be problematic pets orcompanion parrots.[111] Generally, they are not good atmimicking human speech,[112] although thelittle corella is a renowned talker.[113] As social animals, wild cockatoos have been known to learn human speech from ex-captive birds that have integrated into a flock.[114] Their care is best provided by those experienced in keeping parrots.[111] Cockatoos are social animals and their social needs are difficult to cater for,[111] and they can suffer if kept in a cage on their own for long periods of time.[115]
The cockatiel is by far the cockatoo species most frequently kept in captivity. Among U.S. bird keepers that participated in a survey by APPMA in 2003/04, 39% had cockatiels, as opposed to only 3% that had (other) cockatoo species.[116] The white cockatoos are more often encountered in aviculture than the black cockatoos.[117] Black cockatoos are rarely seen in European zoos due to export restrictions on Australian wildlife but birds seized by governments have been loaned.[118]
Cockatoos are often very affectionate with their owner and at times other people but can demand a great deal of attention. It has been suggested that cockatoos' need for physical attention from humans may stem from suboptimal rearing techniques – young birds being removed from parental care for hand-rearing too early in the belief that this will produce a more suitable pet, leading the bird to seek out physical contact from humans as a parent substitute.[119] Furthermore, their intense curiosity means they must be given a steady supply of objects to tinker with, chew, dismantle and destroy. Parrots in captivity may suffer from boredom, which can lead tostereotypic behaviour patterns, such asfeather-plucking. Feather plucking is likely to stem from psychological rather than physical causes.[120] Other major drawbacks include their painful bites,[121] and their piercing screeches.[122] The salmon-crested[123] and white cockatoo species are particular offenders.[124] All cockatoos have a finepowder on their feathers, which may induceallergies in certain people.[121] In general, the smaller cockatoo species such as Goffin's and quieter Galah's cockatoos are much easier to keep as pets.[125] The cockatiel is one of the most popular and easiest parrots to keep as a pet,[126][127] and manycolour mutations are available in aviculture.[36]
A petcockatiel. This parrot is about 32 cm (13 in) long and is by far the smallest and lightest cockatoo.
Larger cockatoos can live 30 to 70 years depending on the species, or occasionally longer, and cockatiels can live for about 20 years. As pets they require a long-term commitment from their owners. Their longevity is considered a positive trait as it reduces instances of the loss of a pet.[7] The oldest cockatoo in captivity was a pink cockatoo namedCookie, residing atBrookfield Zoo in Chicago, which lived to be 83 years old (1933–2016).[128][129][130] A salmon-crested cockatoo named King Tut who resided at theSan Diego Zoo was nearly 69 when he died in 1990 and a palm cockatoo reached 56 inLondon Zoo in 2000.[131] However, anecdotal reports describe birds of much greater ages.[131] Cocky Bennett ofTom Ugly's Point in Sydney was a celebrated sulphur-crested cockatoo who was reported to have reached an age of 100 years or more. He had lost his feathers and was naked for much of his life.[132] A palm cockatoo was reported to have reached 80 or 90 years of age in an Australian zoo,[46] and a little corella that was removed from a nest in central Australia in 1904 was reported still alive in the late 1970s.[78] In February 2010, a white cockatoo named Arthur was claimed to be 90 years old; he had lived with a family for generations inDalaguete,Cebu, before being taken toCebu City Zoo.[133]
Trained cockatoos are sometimes seen in bird shows in zoos. They are generally less motivated by food than other birds; some may respond more to petting or praise than food. Cockatoos can often be taught to wear aparrot harness, enabling their owners to take them outdoors. Cockatoos have been used inanimal-assisted therapy, generally in nursing homes.[134]
Cockatoos often have pronounced responses to musical sounds and numerous videos exist showing the birds dancing to popular music. Research conducted in 2008 with anEleonora cockatoo namedSnowball had indicated that this particular individual is indeed capable ofbeat induction—perceiving human-created music and synchronizing his body movements to the beat.[135]
The earliest European depiction of a cockatoo is in the falconry bookDe arte venandi cum avibus, written byFrederick II, Holy Roman Emperor.[136] The next European depiction of a cockatoo, previously thought to be the earliest, is present in the 1496 painting byAndrea Mantegna titledMadonna della Vittoria. Later examples were painted by Hungarian artistJakob Bogdani (1660–1724), who resided in Amsterdam from 1683 and then England,[137] and appeared with numerous other birds in the bird pieces of the Dutch painterMelchior d'Hondecoeter (1636–1695).[138] A cockatoo is the unlucky subject inAn Experiment on a Bird in the Air Pump by English artistJoseph Wright of Derby, its fate unclear in the painting.[139] Cockatoos were among the many Australian plants and animals which featured in decorative motifs inFederation architecture of the early 20th century.[140] A visit to a Camden Town pet shop in 1958 inspired English painter William Roberts to paintThe Cockatoos, in the collection of the Tate Gallery.[141][142] American artist and sculptorJoseph Cornell was known for placing cutout paper cockatoos in his works.[143]
Two 1970s police dramas featured protagonists with pet cockatoos. In the 1973 filmSerpico,Al Pacino's character had a pet white cockatoo and the television showBaretta sawRobert Blake's character with Fred the Triton cockatoo.[147] The popularity of the latter show saw a corresponding rise in popularity of cockatoos as pets in the late 1970s.[148] Cockatoos have been used frequently in advertising; a cockatoo appeared in a 'cheeky' (and later toned-down) 2008 advertising campaign for Cockatoo Ridge Wineries.[149]
^J. Simpson; E. Weiner, eds. (1989). "cockatoo".Oxford English Dictionary (2nd ed.). Oxford: Clarendon Press.ISBN978-0-19-861186-8.
^Mynott, Jeremy (2009).Birdscapes: Birds in Our Imagination and Experience. Princeton, New Jersey: Princeton University Press. p. 319.ISBN978-0-691-13539-7.
^abde Kloet, RS; de Kloet SR (2005). "The evolution of the spindlin gene in birds: Sequence analysis of an intron of the spindlin W and Z gene reveals four major divisions of the Psittaciformes".Molecular Phylogenetics and Evolution.36 (3):706–21.Bibcode:2005MolPE..36..706D.doi:10.1016/j.ympev.2005.03.013.PMID16099384.
^Schweizer M, Seehausen O, Güntert M, Hertwig ST (2010). "The evolutionary diversification of parrots supports a taxon pulse model with multiple trans-oceanic dispersal events and local radiations".Molecular Phylogenetics and Evolution.54 (3):984–994.doi:10.1016/j.ympev.2009.08.021.PMID19699808.S2CID1831016.
^Leo Joseph; Alicia Toon; Erin E. Schirtzinger; Timothy F. Wright (2011). "Molecular systematics of two enigmatic generaPsittacella andPezoporus illuminate the ecological radiation of Australo-Papuan parrots (Aves: Psittaciformes)".Molecular Phylogenetics and Evolution.59 (3):675–684.Bibcode:2011MolPE..59..675J.doi:10.1016/j.ympev.2011.03.017.PMID21453777.
^abcdAdams M, Baverstock PR, Saunders DA, Schodde R, Smith GT (1984). "Biochemical systematics of the Australian cockatoos (Psittaciformes: Cacatuinae)".Australian Journal of Zoology.32 (3):363–77.doi:10.1071/ZO9840363.
^Courtney, J (1996). "The juvenile food-begging calls, food-swallowing vocalisation and begging postures in Australian Cockatoos".Australian Bird Watcher.16:236–49.ISSN0045-0316.
^Boles, Walter E (1993). "A new cockatoo (Psittaciformes: Cacatuidae) from the Tertiary of Riversleigh, northwestern Queensland, and an evaluation of rostral characters in the systematics of parrots".Ibis.135 (1):8–18.doi:10.1111/j.1474-919X.1993.tb02804.x.
^abcMurphy S, Legge S, Heinsohn R (2003). "The breeding biology of palm cockatoos (Probosciger aterrimus): a case of a slow life history".Journal of Zoology.261 (4):327–39.CiteSeerX10.1.1.475.7031.doi:10.1017/S0952836903004175.
^abcTemby, Ian (1999)."Urban wildlife issues in Australia"(PDF). In Shaw Williams; Lisa Harris; Larry Vandruff (eds.).Proceedings of the 4th International Symposium on Urban Wildlife Conservation. Tucson, Arizona: University of Arizona.Archived(PDF) from the original on 27 September 2011. Retrieved11 December 2009.
^Lindenmayer, DB; Pope MP; Cunningham RB; Donnelly CF; Nix HA (1996). "Roosting of the Sulphur-Crested CockatooCacatua galerita".Emu.96 (3):209–12.Bibcode:1996EmuAO..96..209L.doi:10.1071/MU9960209.
^Heinsohn, R; Murphy S; Legge S (2003). "Overlap and competition for nest holes among eclectus parrots, palm cockatoos and sulphur-crested cockatoos".Australian Journal of Zoology.51 (1):81–94.doi:10.1071/ZO02003.
^Crowley, GM; Garnett S (2001). "Food value and tree selection by Glossy Black-CockatoosCalyptorhynchus lathami".Austral Ecology.26 (1):116–26.doi:10.1046/j.1442-9993.2001.01093.x (inactive 3 December 2024).{{cite journal}}: CS1 maint: DOI inactive as of December 2024 (link)
^Cordon, GP; Hitos Prados A; Romero D; Sánchez Moreno M; Pontes A; Osuna A; Rosales MJ (2009). "Intestinal and haematic parasitism in the birds of the Almunecar (Granada, Spain) ornithological garden".Veterinary Parasitology.165 (3–4):361–66.doi:10.1016/j.vetpar.2009.07.027.PMID19682800.
^Stedman NL, Latimer KS, Rakich PM (1998)."Cloacal papillomas in psittacine birds: A retrospective histopathologic review".Proceedings of International Virtual Conferences in Veterinary Medicine: Diseases of Psittacine Birds.International Virtual Conferences in Veterinary Medicine. Athens, GA: College of Veterinary Medicine, University of Georgia. Archived fromthe original on 20 July 2011. Retrieved1 March 2011.
^Environment and Natural Resources Committee (Parliament of Victoria) (1995).Problems in Victoria caused by Long-billed Corellas, Sulphur-crested Cockatoos and Galahs. Victorian Government Printer.
^"Fauna Note No.20: Little Corella"(PDF). Western Australian Department of Environment and Conservation. 24 July 2007. Archived fromthe original(PDF) on 1 September 2008. Retrieved10 December 2009.
^"Calyptorhynchus latirostris".Species Profile and Threats Database. Department of the Environment, Water, Heritage and the Arts, Canberra. 2009.Archived from the original on 17 May 2018. Retrieved10 December 2009.
^Maron, M (2005). "Agricultural change and paddock tree loss: Implications for an endangered subspecies of Red-tailed Black-Cockatoo".Ecological Management & Restoration.6 (3):206–11.Bibcode:2005EcoMR...6..206M.doi:10.1111/j.1442-8903.2005.00238.x.
^Kinnaird, M; O'Brien TG; Lambert FR; Purmias D (2003). "Density and distribution of the endemic Seram cockatooCacatua moluccensis in relation to land use patterns".Biological Conservation.109 (2):227–35.Bibcode:2003BCons.109..227K.doi:10.1016/S0006-3207(02)00150-7.
^King, C. E.; Heinhuis, H.; Brouwer, K. (January 2000). "Management and husbandry of black cockatoosCalyptorhynchus spp in captivity".International Zoo Yearbook.37 (1):87–116.doi:10.1111/j.1748-1090.2000.tb00710.x.
^Garner, MM; Clubb SL; Mitchell MA; Brown L (2008). "Feather-picking psittacines: histopathology and species trends".Veterinary Pathology.45 (3):401–08.doi:10.1354/vp.45-3-401.PMID18487502.S2CID1023194.
^abBrouwer, K; Jones M; King C; Schifter H (2000). "Longevity records for Psittaciformes in captivity".International Zoo Yearbook.37:299–316.doi:10.1111/j.1748-1090.2000.tb00735.x.
^Swift, W. Bradford (1997). "The healing touch – animal-assisted therapy".Animals.16 (4):130–32.
^Patel AD, Iversen JR, Bregman MR, Schulz I, Schulz C (August 2008).Investigating the human-specificity of synchronization to music(PDF).Proceedings of the 10th Intl. Conf. On Music Perception and Cognition. Adelaide: Causal Productions. Archived fromthe original(PDF) on 6 January 2009. Retrieved14 November 2008.
^Fraser, Hugh; Joyce, Ray (1989).The Federation House – Australia's Own Style. Willoughby, NSW: Weldon Publishing. p. 103.ISBN978-1-86302-033-6.
^"The Cockatoos 1958".Tate Collection: William Roberts 1895–1980. Tate Gallery.Archived from the original on 11 January 2012. Retrieved12 December 2009.
^d'Harnoncourt, A (June 1978). "The Cubist Cockatoo: A Preliminary Exploration of Joseph Cornell's Homages to Juan Gris".Philadelphia Museum of Art Bulletin.74 (321):3–17.doi:10.2307/3795312.JSTOR3795312.
^"ACT Flags and Emblems". Chief Minister's Department, ACT Government.Archived from the original on 28 October 2018. Retrieved10 December 2009.
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