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Dr.Charles M. Clarke (born inMelbourne,Australia) is anecologist andbotanist specialising in thecarnivorous plantgenusNepenthes, for which he is regarded as a world authority.^[1] Clarke has anhonours degree in Botany fromMonash University inMelbourne, and aPh.D. inEcosystem management at theUniversity of New England, inArmidale,New South Wales.

Clarke first travelled to Borneo in search ofpitcher plants in 1987. In 1989 and 1990 he lived inBrunei, studying theecology ofNepenthes. In between travels, Clarke has taughtEcology andBiometrics atJames Cook University inQueensland, and worked as ahorticultural consultant inHong Kong.

He now works at theCairns Botanic Garden.^[1] Clarke has writtenfive books and guides onNepenthes, which present a synthesis of the research performed on his travels around theMalay Archipelago. The most significant of these works are the monographsNepenthes of Borneo (1997) andNepenthes of Sumatra and Peninsular Malaysia (2001).^[2]

Clarke has described six species ofNepenthes:N. baramensis (now known asN. hemsleyana^[3]),N. benstonei,N. chaniana,N. izumiae,N. jacquelineae, andN. tenax.

• Clarke, C., Schlauer, J., Moran, J. & Robinson, A.S., 2018. Systematics and evolution ofNepenthes.In: Ellison, A.M. & Adamec, L., eds.:Carnivorous plants: physiology, ecology, and evolution. Oxford University Press, 58-69.doi:10.1093/oso/9780198779841.003.0005
• Clarke, C.M. 1992. The ecology of metazoan communities inNepenthes pitcher plants in Borneo, with special reference to the community ofNepenthes bicalcarata Hook.f. Ph.D. thesis,University of New England,Armidale,N.S.W. 269 pp.
• Clarke, C.[M.] 1993."The Possible Functions of the Thorns ofNepenthes bicalcarata (Hook.f.) Pitchers"(PDF). (194 KiB)Carnivorous Plant Newsletter22(1–2): 27–28.
• Clarke, C.M. & R.L. Kitching 1993. The Metazoan Food Webs from Six BorneanNepenthes Species.Ecological Entomology18: 7–16.
• Clarke, C.M. & J.A. Moran 1994. A further record of aerial pitchers inNepenthes ampullaria Jack.Malayan Nature Journal47: 321–323.
• Clarke, C.M. & R.L. Kitching 1995. Swimming Ants and Pitcher Plants: a Unique Ant-Plant Interaction from Borneo.Journal of Tropical Ecology11(4): 589–602.
• Clarke, C.M. 1997.Nepenthes of Borneo.Natural History Publications (Borneo),Kota Kinabalu. xi + 207 pp.
• Clarke, C.M. 1997.Another nice trip to Sumatra.Carnivorous Plant Newsletter26(1): 4–10.
• Clarke, C.M. 1997. The effects of pitcher dimorphism on the metazoan community of the carnivorous pitcher plantNepenthes bicalcarata Hook.f.Malayan Nature Journal50: 149–157.
• Clarke, C.M. 1998. Initial colonisation and prey capture inNepenthes bicalcarata (Nepenthaceae) pitchers in Brunei.Sandakania12: 27–36.
• Clarke, C.M. 1998. The aquatic arthropod community of the pitcher plant,Nepenthes bicalcarata (Nepenthaceae) in Brunei.Sandakania11: 55–60.
• Clarke, C.M. 1998. A re-examination of geographical variation inNepenthes food webs.Ecography21(4): 430–436.doi:10.1111/j.1600-0587.1998.tb00408.x
• Clarke, C.M. 1999.Nepenthes benstonei (Nepenthaceae), a new pitcher plant from Peninsular Malaysia.Sandakania13: 79–87.
• Clarke, C.M. 2001. Ecology & Conservation of MontaneNepenthes (Nepenthaceae) in Sumatra. Fourth International Flora Malesiana Symposium. 20–24 July 1998, Kuala Lumpur.
• Clarke, C.M. 2001.Nepenthes of Sumatra and Peninsular Malaysia.Natural History Publications (Borneo),Kota Kinabalu. ix + 325 pp.
• Clarke, C.M. 2001.A Guide to the Pitcher Plants of Sabah.Natural History Publications (Borneo),Kota Kinabalu. iv + 40 pp.
• Moran, J.A., M.A. Merbach, N.J. Livingston, C.M. Clarke & W.E. Booth 2001."Termite Prey Specialization in the Pitcher PlantNepenthes albomarginata - Evidence from Stable Isotope Analysis". Archived fromthe original(PDF) on 4 January 2020.Annals of Botany88: 307–311.
• Clarke, C.M. 2002.A Guide to the Pitcher Plants of Peninsular Malaysia.Natural History Publications (Borneo),Kota Kinabalu. iv + 32 pp.
• Clarke, C.M., T. Davis & R. Tamin 2003.Nepenthes izumiae (Nepenthaceae): a new species from Sumatra.Blumea48: 179–182.
• Moran, J.A., C.M. Clarke & B.J. Hawkins 2003. From Carnivore to Detritivore? Isotopic Evidence for Leaf Litter Utilization by the Tropical Pitcher PlantNepenthes ampullaria.International Journal of Plant Sciences164: 635–639.
• Dong, T.T.X., Q.M. Xiao, C.M. Clarke, H.S. Zhong, N.J. Zhao, K.L. Chun & K.W.K. Tsim 2003. Phylogeny ofAstragalus in China: Molecular evidence from the DNA sequences of 5S rRNA spacer, ITS, and 18S rRNA.Journal of Agricultural and Food Chemistry51(23): 6709–6714.
• Clarke, C.M. & C.C. Lee 2004.A Pocket Guide: Pitcher Plants of Sarawak.Natural History Publications (Borneo),Kota Kinabalu. vi + 81 pp.
• Cantley, R., C.M. Clarke, J. Cokendolpher, B. Rice & A. Wistuba 2004.Nepenthes clipeata Survival Project. International Carnivorous Plant Society.
• Clarke, C.M., C.C. Lee & S. McPherson 2006.Nepenthes chaniana (Nepenthaceae), a new species from north-western Borneo.Sabah Parks Journal7: 53–66.
• Clarke, C.M & R. Kruger 2006.Nepenthes tenax C.Clarke and R.Kruger (Nepenthaceae), a new species from Cape York Peninsula, Queensland.Austrobaileya7(2): 319–324.
• Clarke, C.M. 2006. Introduction. In: Danser, B.H.The Nepenthaceae of the Netherlands Indies. Natural History Publications (Borneo), Kota Kinabalu. pp. 1–15
• Clarke, C.M., U. Bauer, C.C. Lee, A.A. Tuen, K. Rembold & J.A. Moran 2009."Tree shrew lavatories: a novel nitrogen sequestration strategy in a tropical pitcher plant"(PDF). Archived fromthe original(PDF) on 19 July 2011.Biology Letters5(5): 632–635.doi:10.1098/rsbl.2009.0311
• Chin, L., J.A. Moran & C. Clarke 2010. Trap geometry in three giant montane pitcher plant species from Borneo is a function of tree shrew body size.New Phytologist186 (2): 461–470.doi:10.1111/j.1469-8137.2009.03166.x
• Clarke, C.M., J.A. Moran & L. Chin 2010. Mutualism between tree shrews and pitcher plants: perspectives and avenues for future research.Plant Signaling & Behavior5(10): 1187–1189.doi:10.4161/psb.5.10.12807
• Clarke, C., J.A. Moran & C.C. Lee 2011.Nepenthes baramensis (Nepenthaceae) – a new species from north-western Borneo .Blumea56(3): 229–233.doi:10.3767/000651911X607121

• Clarke, C.M. 1997.Nepenthes of Borneo.Natural History Publications (Borneo),Kota Kinabalu. xi + 207 pp. ISBN 983-812-057-X

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