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Calocedrus

From Wikipedia, the free encyclopedia
Genus of conifer trees

Incense-cedar
Calocedrus decurrens
California incense cedar
Scientific classificationEdit this classification
Kingdom:Plantae
Clade:Tracheophytes
Clade:Gymnospermae
Division:Pinophyta
Class:Pinopsida
Order:Cupressales
Family:Cupressaceae
Subfamily:Cupressoideae
Genus:Calocedrus
Kurz
Type species
Calocedrus macrolepis
Synonyms[1]

HeyderiaK.Koch 1873 non Link 1833

California incense cedar, inLassen Volcanic National Park

Calocedrus, theincense cedar (alternatively spelledincense-cedar), is agenus ofconiferoustrees in the cypress familyCupressaceae first described as a genus in 1873.[2][3] Three species are native toeastern Asia and one to westernNorth America.[1]

Description

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The genus is related toThuja, and has similar overlapping scale-leaves.Calocedrus differs fromThuja in the scale leaves being in apparent whorls of four (actually opposite decussate pairs likeThuja, but not evenly spaced apart as inThuja, instead with the successive pairs closely then distantly spaced), and in thecones having just 2–3 pairs of moderately thin, erect scales, rather than 4–6 pairs of very thin scales inThuja.

Taxonomy

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The generic nameCalocedrus means "beautiful cedar".

Cladogram showing the evolutionary relationships:

Stull et al. 2021[4][5]
Calocedrus

C. decurrens(Torrey) Florin

C. macrolepisKurz

C. formosana(Florin) Florin

C. rupestrisAver., Nguyên & Lôc

C. decurrens foliage and male cones

Species

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Extant species

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ImageNameDescriptionDistribution
Calocedrus decurrens, California incense cedar (syn.Libocedrus decurrens)It is a large tree, typically reaching heights of 40–60 m and a trunk diameter of up to 3 m (maxima, 69 m tall and 3.9 m diameter), and with a broad conic crown of spreading branches. The leaves are bright green on both sides of the shoots, and the cones 2–2.5 cm long. It is by far the most widely known species in the genus, and is often simply called "incense cedar" without the regional qualifier.western North America
Calocedrus formosana, Taiwan incense cedarIt is very similar toC. macrolepis, and some botanists treat it as a variety of that,C. macrolepis var.formosana. It is a medium-sized tree, growing to 25–30 m tall, and is rare in the wild, occurring only as scattered trees in mixed forests. The leaves are glaucous green on the upper side of the shoots, and conspicuously marked with bright white stomatal patches on the underside. The cones are 1.5–2 cm long, carried on a 1–1.5 cm stem.[6]Taiwan
Calocedrus macrolepis, Chinese incense cedarIt is also a medium-size tree to 25–30 m tall, and likeC. formosana, is rare in the wild. The leaves and cones are similar toC. formosana, differing most obviously in the shorter cone stem, only 0.5 cm long.[7]southwestChina (fromGuangdong west toYunnan), and also in northernVietnam, northernLaos, extreme northernThailand and northeasternMyanmar
Calocedrus rupestrisThe most recently discovered living species ofCalocedrus, first described in 2004. It occurs exclusively on rocky limestone (karst) terrain, a habitat that has a very high level of endemism. The close proximity of these populations to theChinese andLaotian borders indicates that the species may occur in those countries as well. It is an evergreen, monoecious tree up to 25 m tall with a broadly rounded crown. The epithet "rupestris" means "rock-dwelling".[8][9]Vietnam

Extinct species

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NameDescriptionDistribution
Calocedrus huashanensisDescribed in 2012. It is known fromcompression fossils found in theOligocene ageNingming Formation of southern China.Calocedrus huashanensis is known from branches and leaves.southern China
Calocedrus suleticensisknown fromfossils found in theEarly Oligocene ofProbostov (Holy Kluk Hill) in the volcanic complex of the Ceske stredohori Mts.,Bohemia.Calocedrus suleticensis is known from a cone.Czech Republic

Uses

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Archery

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Incense cedar was one of the favored varieties of wood used to make bows by Native Americans in California. Likejuniper, andPacific yew, the other two coveted bow woods among Pacific Natives, this wood has excellent flexibility and compression strength-weight ratio. When backed with sinew, it produces extremely flexible, fast, hard-hitting bows, which are rivaled only by horn-sinew composite bows for their ability to store and release elastic energy. The archer Saxton Pope observed thatIshi used this wood to produce short bows.[10]

Lumber

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Thewood ofCalocedrus is soft, moderately decay-resistant, and with a strong spicy-resinous fragrance. That ofC. decurrens is the primary material for woodenpencils, because it is soft and tends to sharpen easily without forming splinters. The two Asian species were (at least in the past) in very high demand forcoffin manufacture in China, due to the scent of the wood and its decay resistance. It is likely that pastover-exploitation is responsible for their current rarity.

Incense cedar was the preferredhearth board of the Native Peoples ofNorthern California forlighting fires by friction.

Cultivation

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Calocedrus decurrens, the California incense cedar, is a popularornamental tree, grown particularly in locations with cool summer climates likeBritain,Washington andBritish Columbia. Its very narrow columnar crown in landscape settings, an unexplained consequence of the climatic conditions in these areas, is not shown by trees in their native 'wild'habitat. The California incense cedar is also valued for itsdrought tolerance. The Asian species are rarely cultivated.[11]

References

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  1. ^ab"Calocedrus".World Checklist of Selected Plant Families (WCSP).Royal Botanic Gardens, Kew. Archived fromthe original on October 10, 2018.
  2. ^Kurz, Wilhelm Sulpiz (1873). "On a few new plants from Yunan".Journal of Botany, British and Foreign.11:196.OCLC 1642195.
  3. ^"Calocedrus Kurz".Tropicos.Missouri Botanical Garden.
  4. ^Stull, Gregory W.; Qu, Xiao-Jian; Parins-Fukuchi, Caroline; Yang, Ying-Ying; Yang, Jun-Bo; Yang, Zhi-Yun; Hu, Yi; Ma, Hong; Soltis, Pamela S.; Soltis, Douglas E.; Li, De-Zhu; Smith, Stephen A.; Yi, Ting-Shuang; et al. (2021)."Gene duplications and phylogenomic conflict underlie major pulses of phenotypic evolution in gymnosperms".Nature Plants.7 (8):1015–1025.bioRxiv 10.1101/2021.03.13.435279.doi:10.1038/s41477-021-00964-4.PMID 34282286.S2CID 232282918.
  5. ^Stull, Gregory W.; et al. (2021)."main.dated.supermatrix.tree.T9.tre". Figshare.doi:10.6084/m9.figshare.14547354.v1.{{cite journal}}:Cite journal requires|journal= (help)
  6. ^Fu, Liguo; Yu, Yong-fu; Adams, Robert P.; Farjon, Aljos."Calocedrus macrolepis var.formosana".Flora of China. Vol. 4 – via eFloras.org,Missouri Botanical Garden, St. Louis, MO &Harvard University Herbaria, Cambridge, MA.
  7. ^Fu, Liguo; Yu, Yong-fu; Adams, Robert P.; Farjon, Aljos."Calocedrus macrolepis".Flora of China. Vol. 4 – via eFloras.org,Missouri Botanical Garden, St. Louis, MO &Harvard University Herbaria, Cambridge, MA.
  8. ^Averyanov, H.T. Nguyen & L.K. Phan. Issues of Basic Research in Life Sciences with direction in upland agriculture and forestry. Proceedings, the 2004th [sic] National Conference on Life Sciences Thai Nguyen University, September 23, 2004 41–43, 1.
  9. ^"Calocedrus".World Checklist of Selected Plant Families (WCSP).Royal Botanic Gardens, Kew. Archived fromthe original on May 13, 2019.
  10. ^"How Ishi made his bow and his method of shooting, from: Hunting with the Bow and Arrow by Saxton Pope, 1923". 11 February 2019.
  11. ^Munz, P. A. 1974. Flora of Southern California 1–1086. University of California Press, Berkeley.

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