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Balfour Formation

Coordinates:32°18′S24°30′E / 32.3°S 24.5°E /-32.3; 24.5
From Wikipedia, the free encyclopedia
Geological formation in the Beaufort Group of South Africa
Balfour Formation
Stratigraphic range:Lopingian -Induan
256.3–251.2 Ma
Outcrops of the Balfour Formation are found in the hillsides surrounding Nieu-Bethesda, Eastern Cape, South Africa
TypeGeological formation
Unit ofAdelaide Subgroup,Beaufort Group
Sub-unitsOudeberg Member, Daggaboersnek Member, Ripplemead Member, Elandsberg Member, Palingkloof Member
UnderliesKatberg Formation
OverliesMiddleton Formation
Thicknessup to 2,150 m (7,050 ft)
Lithology
PrimaryMudstone,siltstone
OtherSandstone
Location
Coordinates32°18′S24°30′E / 32.3°S 24.5°E /-32.3; 24.5
Approximate paleocoordinates62°42′S29°18′W / 62.7°S 29.3°W /-62.7; -29.3
RegionNorthern Cape,Eastern Cape andFree State
Country South Africa

TheBalfour Formation is ageological formation that is found in theBeaufort Group, a majorgeological group that forms part of the greaterKaroo Supergroup inSouth Africa. The Balfour Formation is the uppermost formation of the Adelaide Subgroup which contains all the LatePermian - EarlyTriassic aged biozones of theBeaufort Group.Outcrops and exposures of the Balfour Formation are found from east of 24 degrees in the highest mountainousescarpments betweenBeaufort West andFraserburg, but most notably in the Winterberg and Sneeuberg mountain ranges nearCradock, theBaviaanskloof river valley,Graaff-Reniet andNieu Bethesda in theEastern Cape, and in the southernFree State province.[1][2]

Geology

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The Balfour Formation overlies the Middleton Formation of the Adelaide Subgroup and underlies the Katberg Formation of the lower Tarkastad Subgroup, all comprising the greaterBeaufort Group. The Balfour Formation is composed of five members which are listed below (from oldest to youngest):

  • Oudeberg Member
  • Daggaboersnek Member
  • Ripplemead Member
  • Elandsberg Member
  • Palingkloof member

The rocks of the Balfour Formation also incorporate the entireDaptocephalus Assemblage Zone, the lowermost portion of theLystrosaurus Assemblage Zone,[3] and the uppermost rocks of theCistecephalus Assemblage Zone.[4] Up until the middle section of the Ripplemead Member, the Balfour Formation correlates with the near contemporaneous Teekloof Formation west of the 24 degrees fromBeaufort West westwards, and to the Normadien Formation north of theOrange River. However, the Elandsberg and Palingkloof Members do not have any lateral correlates west of 24 degrees. This is either due to pasterosion of the upper, unknown members of the Teekloof Formation or there was a sudden cessation of sedimentary deposition in the western section of theKaroo Basin.[5][6][7][8]

Thesedimentary rocks of this formation are composed predominantly of alternating greenish-grey, bluish-grey, and grey-ish redmudstone that often containsiltstonelenses. Themudstones are very fine-grained, massive and exhibit blocky weathering.Claystone successions are also found which, along with themudstones, frequently contain desiccation cracks, raindrop impressions, andcalcareousnodules orconcretions are found throughout.Rhythmites are also found.Sandstones are less common, but some notable units have been studied in the Balfour Formation. In the lowermost section of the Balfour is asandstone-rich unit known as the Oudeberg Member. Thesandstones in this unit are very fine-grained and are rich infeldspar. Another sandstone unit in the middle of the Balfour Formation is the Daggaboersnek Member which contains thin, tabular sandstones, and ripple structures are common.[9][10]

The presence of these rocks reveal much about the past environment that they were deposited in. The dominance of fine-grainedmudstone and less common, fine-grainedsandstones indicates that the rock sediments were deposited in a low-energy,fluvial environment, most likely one that hadmeandering rivers. At the time of sedimentary deposition, the Karooretroarcforeland system was in an overfilled phase, and purely terrestrial sediments occupied theKaroo Basin at this time. As this formation includes the rocks of both theCistecephalus,Daptocephalus,Lystrosaurus Assemblage Zones, the Balfour Formation preserves the geological record for theend Permian extinction event. This is important as the end Permian extinction event was the largest massextinction event in the Earth's history. This was followed by one of the worst biotic crises, which is reflected in the sudden and drasticsedimentaryfacies changes in the overlying Katberg Formation.[11][12][13][14][15]

Paleontology

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Where the Balfour Formation deposits correlate with theDaptocephalus Assemblage Zone, a great diversity ofvertebratefauna are found. This richness in species diversity observed in the Balfour Formation is especially true ofdicynodonts as numerous different species of this successful,herbivoroustherapsid have been recovered.[16][17] Various species ofburnetiamorphbiarmosuchians,rubidgeinegorgonopsians, andtherocephalianspecies such asMoschorhinus kitchingi and the earliestcynodont,Charassognathus gracilis, also appear.[18]Parareptile species,[19] namelycaptorhinids, theYounginiformeYoungina capensis,[20][21] and a variety oftemnospondylamphibians,[22] fishes, and plant fossils such asGlossopteris are likewise found.Glossopteris fossils or leaf impressions are particularly common in the Daggaboersnek Member. The uppermost unit of the Balfour Formation marks the Permian-aged side of thePermian-Triassic boundary. At this point in thebiostratigraphy there is a marked drop in species diversity as the Permian-Triassic extinction event began to take its course at the time of sediment deposition.

Paleofauna

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Reptiles

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Diapsids

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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
YounginaY. capensisKlipplaat, Doornplaas, Toverwater, Beeldhouersfortien, and Wellwood.Dicynodon Assemblage zone.LopingianAdiapsid reptile.

Parareptiles

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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
AnthodonA. serrariusChanghsingianA pareiasaur.
MilleropsisM. priceiDicynodon Assemblage zone.ChanghsingianAmillerettid parareptile.
MillerettaM. rubidgeiDicynodon Assemblage zone.ChanghsingianAmillerettid parareptile.
MillerosaurusM. nuffieldiChanghsingianAmillerettid parareptile.
PareiasaurusP. serridensWuchiapingian and Changhsingian.A pareiasaur.
OwenettaO. rubidgeiCistecephalus and Lystrosaurus Assemblage ZonesWuchiapingian and ChanghsingianA procolophonian parareptile.
SauropareionS. anoplusMiddleberg and Barendskraal Farm.Lystrosaurus assemblage zone.Induan.A basalprocolophonid parareptile.

Proterosuchids

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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
ProterosuchusP. fergusiBarendskraal farm and Middleburg.Lystrosaurus assemblage zone.Changhsingian.A complete skull.Aproterosuchidae reptile.

Synapsids

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Therapsids

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Cynodonts
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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
CharassognathusC. gracilisAcynodont.
CynosaurusC. suppostusCistecephalus and Daptocephalus Assemblage Zones.Wuchiapingian - Changhsingian.A cynodont.
ProcynosuchusP. delaharpeaeNew Bathsheba.WuchiapingianA cynodont.
ProgalesaurusP. lootbergensisNew Lootsberg Pass.Lystrosaurus Assemblage Zone.LopingianA skull, dentition, and postcranial skeleton.A cynodont.
Vetusodon[23]V. elikhuluEnyezane.Daptocephalus Assemblage Zone.LopingianA partial skullA cynodont.
Dicynodonts
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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
DaptocephalusD. leonicepsDaptocephalus Assemblage Zone.ChanghsingianAdicynodont.
DolomitipesD. accordiiAdicynodont.
D. icelsi
DicynodontoidesD. recurvidensNooitgedacht 68.Uppermost part in Dicynodon Assemblage Zone.A skull and postcranial element.A dicynodont.
EmydopsE. arctatusAdicynodont.
LystrosaurusL. curvatusNooitgedacht 68 and Fairydale.lower part of Palingkloof Member, Dicynodon Assemblage Zone.Induan.A skull.Adicynodont.
Lystrosaurus
Lystrosaurus
L. declivisuppermost part of Lystrosaurus Assemblage Zone.Induan.A skull and a complete skeleton.
L. maccaigiPalingkloof Member, Dicynodon Assemblage ZoneChanghsingian.
L. murrayilower part of Palingkloof Member, Lystrosaurus Assemblage Zone.Induan.A skull and skeleton.
OudenodonO. bainiiA small dicynodont.
Gorgonopsids
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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
AloposaurusA. tenuisA gorgonopsid.
Alopsaurus
Alopsaurus
CyonosaurusC. kitchingiA gorgonopsid.
C. longiceps
DinogorgonD. rubidgeiCistecephalus and Daptocephalus assemblage.A gorgonopsid.
Inostrancevia[24]I. africanalower slope of the hill Loskop, Nooitgedacht 68.Daptocephalus Assemblage ZoneA partial skeleton including a complete skull and occluded mandible, entirety of the precaudal axial column, partial scapulae, partial right pelvis, the right femur, and a disarticulated left humerus, ulna, radius, and tibia.A predatoryinostranceviinaegorgonopsid.
LeontosaurusL. vanderhorstiDicynodon Assemblage Zone.A gorgonopsid.
LycaenopsL. ornatusA gorgonopsid.
ParagalerhinusP. rubidgeiWellwoodDicynodon Assemblage Zone.A partial skull.A gorgonopsid.
RubidgeaR. atroxCistecephalus and Daptocephalus assemblage zone.A gorgonopsid.
ScylacopsS. capensisA gorgonopsid.
Therocephalians
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GenusSpeciesLocalityStratigraphic positionTimeMaterialNotesImages
IctidochampsaI. platycepsDicynodon Assemblage Zone.Atherocephalian.
IctidosuchoidesI. intermediusAtherocephalian.
IctidosuchopsI. intermediusAtherocephalian.
MoschorhinusM. kitchingiSpitskop, Nooitgedacht 68.Dicynodon Assemblage Zone.A weathered skull along with a complete skull, partially articulated anterior skeleton, and disarticulated pelvis and hind limb.Atherocephalian.
Moschorhinus.
NotaelurodonN. kitchingiAtherocephalian.
OlivierosuchusO. parringtoniPalingkloof MemberLystrosaurus assemblege zone.A complete skull and anterior half of skeleton in articulation, including vertebrae, ribs, pectoral girdle, partial left humerus, complete right forelimb,

including terminal phalanges.

A therocephalian.
PromoschorhynchusP. platyrhinusAtherocephalian.
TheriognathusT. micropsAtherocephalian.
TetracynodonT. tenuisAtherocephalian.

Correlation

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The Balfour Formation is known to corresponds in age with theCis-Uralian fauna ofRussia[25] and theSanga do Cabral Formation,Paraná Basin ofBrazil. Correlations with other LatePermian - EarlyTriassic deposits abroad remain inconclusive.

References

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  1. ^Rubidge, B. S. (ed.) 1995b.Biostratigraphy of the Beaufort Group (Karoo Supergroup). South African Committee of Stratigraphy.Biostratigraphic Series 1. Pretoria, Council for Geoscience.
  2. ^Catuneanu, O., Elango, H. N. (2001-04-15). "Tectonic control on fluvial styles: the Balfour Formation of the Karoo Basin, South Africa".Sedimentary Geology.140 (3–4):291–313.Bibcode:2001SedG..140..291C.doi:10.1016/S0037-0738(00)00190-1.ISSN 0037-0738.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  3. ^Botha, Jennifer; Smith, Roger M. H. (2007-06-01). "Lystrosaurus species composition across the Permo–Triassic boundary in the Karoo Basin of South Africa".Lethaia.40 (2):125–137.Bibcode:2007Letha..40..125B.doi:10.1111/j.1502-3931.2007.00011.x.ISSN 1502-3931.
  4. ^Kammerer, Christian F.; Bandyopadhyay, Saswati; Ray, Sanghamitra (2016-11-01). "A new taxon of cistecephalid dicynodont from the upper Permian Kundaram Formation of India".Papers in Palaeontology.2 (4):569–584.Bibcode:2016PPal....2..569K.doi:10.1002/spp2.1055.ISSN 2056-2802.S2CID 88833541.
  5. ^Visser, J.N.J. and Dukas, B.A., 1979. Upward-fining fluviatile megacycles in the Beaufort Group north of Graaff-Reinet, Cape Province.Trans. Geol. Soc. S. Afr,82(149), p.e154.
  6. ^Johnson, M.R.; Vuuren, C.J. Van; Visser, J.N.J.; Cole, D.I.; Wickens, H. De V.; Christie, A.D.M.; Roberts, D.L. (1997-01-01).Chapter 12 The foreland karoo basin, south africa. Vol. 3. pp. 269–317.doi:10.1016/S1874-5997(97)80015-9.ISBN 9780444825711.ISSN 1874-5997.{{cite book}}:|journal= ignored (help)
  7. ^Rubidge, B. S.; Hancox, P. J.; Catuneanu, O. (1998-12-01)."Reciprocal flexural behaviour and contrasting stratigraphies: a new basin development model for the Karoo retroarc foreland system, South Africa".Basin Research.10 (4):417–439.Bibcode:1998BasR...10..417C.doi:10.1046/j.1365-2117.1998.00078.x.ISSN 1365-2117.S2CID 56420970.
  8. ^Katemaunzanga, David; Gunter, Cornelis Janse (2009-10-01). "Lithostratigraphy, Sedimentology, and Provenance of the Balfour Formation (Beaufort Group) in the Fort Beaufort–Alice Area, Eastern Cape Province, South Africa".Acta Geologica Sinica - English Edition.83 (5):902–916.Bibcode:2009AcGlS..83..902K.doi:10.1111/j.1755-6724.2009.00110.x.ISSN 1755-6724.S2CID 130345218.
  9. ^Viglietti, Pia; Rubidge, Bruce; Malcom Harris Smith, Roger (2017-03-01)."Revised lithostratigraphy of the upper Permian Balfour and Teekloof formations of the main Karoo Basin, South Africa".South African Journal of Geology.120 (1):45–60.Bibcode:2017SAJG..120...45V.doi:10.25131/gssajg.120.1.45.
  10. ^Hiller, N., Stavrakis, N. (1984-02-01). "Permo-Triassic fluvial systems in the southeastern Karoo Basin, South Africa".Palaeogeography, Palaeoclimatology, Palaeoecology.45 (1):1–21.Bibcode:1984PPP....45....1H.doi:10.1016/0031-0182(84)90106-8.ISSN 0031-0182.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  11. ^Ward, Peter D.; Koch, Paul L.; Smith, Roger M. H.; MacLeod, Kenneth G. (2000-03-01). "Timing of mammal-like reptile extinctions across the Permian-Triassic boundary in South Africa".Geology.28 (3):227–230.Bibcode:2000Geo....28..227M.doi:10.1130/0091-7613(2000)28<227:TOMREA>2.0.CO;2.ISSN 0091-7613.
  12. ^Ward, Peter D.; Smith, Roger M. H. (2001-12-01). "Pattern of vertebrate extinctions across an event bed at the Permian-Triassic boundary in the Karoo Basin of South Africa".Geology.29 (12):1147–1150.Bibcode:2001Geo....29.1147S.doi:10.1130/0091-7613(2001)029<1147:POVEAA>2.0.CO;2.ISSN 0091-7613.
  13. ^Smith, R. M. H., Botha, J. (2005-09-01). "The recovery of terrestrial vertebrate diversity in the South African Karoo Basin after the end-Permian extinction".Comptes Rendus Palevol.4 (6–7):623–636.Bibcode:2005CRPal...4..623S.doi:10.1016/j.crpv.2005.07.005.ISSN 1631-0683.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  14. ^Smith, R. M. H., Botha, J. (2006-08-01). "Rapid vertebrate recuperation in the Karoo Basin of South Africa following the End-Permian extinction".Journal of African Earth Sciences.45 (4–5):502–514.Bibcode:2006JAfES..45..502B.doi:10.1016/j.jafrearsci.2006.04.006.ISSN 1464-343X.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  15. ^Smith, Roger; Kirschvink, Joseph L.; Garrison, Geoffrey H.; Erwin, Douglas H.; Kock, Michiel O. De; Buick, Roger; Botha, Jennifer; Ward, Peter D. (2005-02-04). "Abrupt and Gradual Extinction Among Late Permian Land Vertebrates in the Karoo Basin, South Africa".Science.307 (5710):709–714.Bibcode:2005Sci...307..709W.CiteSeerX 10.1.1.503.2065.doi:10.1126/science.1107068.ISSN 1095-9203.PMID 15661973.S2CID 46198018.
  16. ^Fröbisch, Jörg (2008-11-17)."Global Taxonomic Diversity of Anomodonts (Tetrapoda, Therapsida) and the Terrestrial Rock Record Across the Permian-Triassic Boundary".PLOS ONE.3 (11): e3733.Bibcode:2008PLoSO...3.3733F.doi:10.1371/journal.pone.0003733.ISSN 1932-6203.PMC 2581439.PMID 19011684.
  17. ^Viglietti, P. A., Smith, R. M. H., Angielczyk, K. D., Kammerer, C. F., Fröbisch, J., Rubidge, B. S. (2016-01-01). "The Daptocephalus Assemblage Zone (Lopingian), South Africa: A proposed biostratigraphy based on a new compilation of stratigraphic ranges".Journal of African Earth Sciences.113:153–164.Bibcode:2016JAfES.113..153V.doi:10.1016/j.jafrearsci.2015.10.011.ISSN 1464-343X.{{cite journal}}: CS1 maint: multiple names: authors list (link)
  18. ^Kammerer, Christian F. (2016-01-26)."Systematics of the Rubidgeinae (Therapsida: Gorgonopsia)".PeerJ.4: e1608.doi:10.7717/peerj.1608.ISSN 2167-8359.PMC 4730894.PMID 26823998.
  19. ^Dias-Da-Silva, Sérgio; Modesto, Sean Patrick; Schultz, Cesar Leandro (2006). "Canadian Science Publishing".Canadian Journal of Earth Sciences.43 (11):1685–1693.Bibcode:2006CaJES..43.1685D.doi:10.1139/e06-043.S2CID 140641975.
  20. ^Gow, Chris E. (1975)."The morphology and relationships of Youngina capensis Broom and Prolacerta broomi Parrington".Palaeontologia Africana.ISSN 0078-8554.
  21. ^Gardner, Nicholas; Holliday, Casey; O’Keefe, F. (2010-11-01)."The Braincase of Youngina Capensis (Reptilia, Diapsida): New Insights from High-Resolution CT Scanning of the Holotype".Biological Sciences Faculty Research.
  22. ^Damiani, R. J. (2004-01-01). "Temnospondyls from the Beaufort Group (Karoo Basin) of South Africa and Their Biostratigraphy".Gondwana Research.7 (1):165–173.Bibcode:2004GondR...7..165D.doi:10.1016/S1342-937X(05)70315-4.ISSN 1342-937X.
  23. ^Abdala, Fernando & Gaetano, Leandro. (2019). A new, large cynodont from the Late Permian of the Karoo Basin, South Africa and its bearings in epicynodont phylogeny.. Zoological Journal of the Linnean Society. 10.1093/zoolinnean/zlz004.
  24. ^Kammerer, Christian F.; Viglietti, Pia A.; Butler, Elize; Botha, Jennifer (2023). "Rapid turnover of top predators in African terrestrial faunas around the Permian-Triassic mass extinction".Current Biology.33 (11): 2283–2290.e3.Bibcode:2023CBio...33E2283K.doi:10.1016/j.cub.2023.04.007.PMID 37220743.S2CID 258835757.
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