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Atocetus

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Extinct genus of mammals

Atocetus Temporal range:Mid-Late Miocene ~13–7 Ma PreꞒ Ꞓ O S D C P T J K Pg N

Skull ofA. iquensis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Artiodactyla
Infraorder:	Cetacea
Family:	†Kentriodontidae
Subfamily:	†Pithanodelphininae
Genus:	†Atocetus De Muizon 1988
Species
A. iquensisDe Muizon 1988 (type) A. nasalis(Barnes 1985)

Atocetus is an extinctgenus ofpontoporiid dolphin found inMiocene-age marine deposits inPeru andCalifornia.^[1]

Description

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The type species,Atocetus iquensis, is known fromSerravallian-age strata of thePisco Formation of Peru,^[2] whileA. nasalis is known fromTortonian-age marine deposits of theModelo Formation in California.^[3] Barnes (1985) originally described the latter as a species ofPithanodelphis, but it was eventually transferred toAtocetus by Muizon (1988). The Miocene delphinidan"Champsodelphis" fuchsii from marine deposits inAustria was initially tentatively referred toAtocetus based on the discovery of partial skeletons from the Carpathian region with earbones similar to those ofAtocetus but was eventually reassigned toKentriodon following the discovery of additional earbones from Austria andRomania.^[4]^[5]

Phylogeny

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AlthoughAtocetus and other pithanodelphinines are usually assigned toKentriodontidae, the cladistic analysis of Lambert et al. (2017) showed thatAtocetus andPithanodelphis form a clade withTagicetus andLophocetus that is not only phylogenetically more derived than other kentriodontids but is also phylogenetically intermediate between thebaiji and members ofInioidea andPhocoenidae.^[6] However, a subsequent cladistic analysis by Post et al. (2017) recoversAtocetus as a member of Pontoporiidae, which includes thefranciscana.^[7]

References

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^Ichishima, H.; Barnes, L. G.; Fordyce, R. E.; Kimura, M.; Bohaska, D. J. (1994). "A review of kentriodontine dolphins (Cetacea; Deiphinoidea; Kentriodontidae): Systematics and biogeography".The Island Arc.3 (4): 486.Bibcode:1994IsArc...3..486I.doi:10.1111/j.1440-1738.1994.tb00127.x.
^De Muizon, 1988, p.131
^L. G. Barnes. 1985. The Late Miocene dolphin Pithanodelphis Abel, 1905 (Cetacea: Kentriodontidae) from California. Contributions in Science 367:1-27.
^E. Kazár, M. Vremir, and V. Codrea. 2004. Dolphin remains (Cetacea: Odontoceti) from the middle Miocene of Cluj-napoca, Romania. Acta Paleontologica Romanie 4:179-189.
^E. Kazár. 2006. Odontocete periotics (Mammalia: Cetacea) from the Carpathian Basin, Middle Miocene (Badenian and Sarmatian Stages), including the Vienna Basin, Austria. Beitrage zur Palaontologie 30:269-292.
^Olivier Lambert, Giovanni Bianucci, Mario Urbina, Jonathan H. Geisler; A new inioid (Cetacea, Odontoceti, Delphinida) from the Miocene of Peru and the origin of modern dolphin and porpoise families. Zool J Linn Soc 2017; 179 (4): 919-946. doi: 10.1111/zoj.12479
^Post K, Louwye S, Lambert O. (2017) Scaldiporia vandokkumi, a new pontoporiid (Mammalia, Cetacea, Odontoceti) from the Late Miocene to earliest Pliocene of the Westerschelde estuary (The Netherlands) PeerJ 5:e3991https://doi.org/10.7717/peerj.3991

Bibliography

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De Muizon, C (1988),"Les Vertébrés fossiles de la Formation Pisco (Pérou). Troisième partie: Les Odontocètes (Cetacea, Mammalia) du Miocène",Editions Recherche Sur les Civilisations,78:1–244, retrieved2019-03-13

Odontocete genera

Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Order:Artiodactyla
Infraorder:Cetacea

Odontoceti

†Agorophius †Ankylorhiza †Archaeodelphis †Argyrocetus †Ashleycetus †Caolodelphis †Chilcacetus †Enigmatocetus †Eoplatanista †Inticetus †Macrodelphinus †Microcetus †Mirocetus †Neosqualodon †Olympicetus †Papahu †Patriocetus †Phoberodon †Phococetus †Prosqualodon †Sachalinocetus †Simocetus †Sulakocetus †Tangaroasaurus
†Xenorophidae	Albertocetus Cotylocara Echovenator Inermorostrum Xenorophus
†Waipatiidae	Awamokoa Ediscetus Nihohae Nihoroa Otekaikea Urkudelphis Waipatia
†Squalodontidae	Eosqualodon Squalodon
†Squaloziphiidae	Crisocetus? Squaloziphius Yaquinacetus
†Eurhinodelphinidae	Ceterhinops Eurhinodelphis Iniopsis Mycteriacetus Phocaenopsis Schizodelphis Vanbreenia Xiphiacetus Ziphiodelphis
Crown-Odontoceti	see below↓

Crown-Odontoceti

Physeteroidea

†Acrophyseter †Albicetus †Brygmophyseter †Eudelphis †Hoplocetus †Livyatan †Miophyseter †Rhaphicetus †Scaldicetus †Zygophyseter
Kogiidae	†Aprixokogia Kogia †Koristocetus †Nanokogia †Platyscaphokogia †Pliokogia †Praekogia †Scaphokogia †Thalassocetus?
Physeteridae	†Aulophyseter †Cozzuoliphyseter †Diaphorocetus †Idiophyseter †Idiorophus †Orycterocetus Physeter †Physeterula †Placoziphius †Thalassocetus?

Platanistoidea

†Aondelphis †Dolgopolis †Ensidelphis †Perditicetus †Eurhinodelphinidae? †Squalodontidae? †Waipatiidae?
†Allodelphinidae?	Allodelphis Arktocara Goedertius Ninjadelphis Zarhinocetus
†Squalodelphinidae?	Furcacetus Huaridelphis Macrosqualodelphis Medocinia Notocetus Phocageneus Squalodelphis
Platanistidae	†Araeodelphis †Dilophodelphis †Grimadelphis †Pachyacanthus? †Pebanista Platanista †Pomatodelphis †Prepomatodelphis †Zarhachis?

Ziphiidae

†Aporotus †Beneziphius †Caviziphius †Chavinziphius †Chimuziphius †Choneziphius †Dagonodum †Globicetus †Imocetus †Messapicetus †Nazcacetus †Nenga †Ninoziphius †Notoziphius †Pterocetus Tasmacetus †Tusciziphius †Xhosacetus †Ziphirostrum
Berardiinae	†Archaeoziphius Berardius †Microberardius
Ziphiinae	†Izikoziphius Ziphius
Hyperoodontinae	†Africanacetus Hyperoodon †Ihlengesi Indopacetus Mesoplodon

Delphinida	see below↓

Delphinida

†Albireo †Atocetus †Brevirostrodelphis †Cammackacetus †Delphinodon †Hadrodelphis †Herbeinodelphis †Liolithax †Lophocetus †Macrokentriodon †Miminiacetus †Odobenocetops †Pictodelphis †Pithanodelphis †Tagicetus †Westmorelandelphis
†Kentriodontidae	Belonodelphis? Kampholophos Kentriodon Platysvercus Sophianacetus Rudicetus Wimahl
Lipotidae	Lipotes †Parapontoporia †Prolipotes

Inioidea

†Awadelphis
Iniidae	†Brujadelphis †Goniodelphis Inia †Ischyrorhynchus †Isoninia? †Isthminia †Kwanzacetus †Meherrinia †Plicodontinia †Saurocetes
Pontoporiidae	†Atocetus? †Auroracetus †Brachydelphis †Piscorhynchus †Pliopontos †Pontistes †Protophocaena †Samaydelphis †Scaldiporia †Stenasodelphis Pontoporia

Delphinoidea

Monodontidae	†Bohaskaia †Casatia Delphinapterus †Denebola †Haborodelphis Monodon
Phocoenidae	†Archaeophocaena †Australithax †Haborophocoena †Lomacetus †Miophocaena Neophocaena †Numataphocoena Phocoena Phocoenoides †Piscolithax †Pterophocaena? †Semirostrum †Septemtriocetus

Delphinidae

†Arimidelphis †Australodelphis †Eodelphinus †Etruridelphis †Hemisyntrachelus Lagenorhynchus †Norisdelphis Orcinus †Platalearostrum †Pliodelphis †Pterophocaena?
Lissodelphininae	Cephalorhynchus Lissodelphis
Delphininae	Delphinus Lagenodelphis Sotalia Sousa Stenella Tursiops
Globicephalinae	Feresa Globicephala Grampus Orcaella Peponocephala Pseudorca Steno

Taxon identifiers
Atocetus	Wikidata:Q19720723 GBIF:4575950 IRMNG:1284637 Open Tree of Life:3612350 Paleobiology Database:36669

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