Aromatic rice produces more 2-acetyl-1-pyrroline than usual due to a loss-of-function mutation in the BADH2 gene. The BADH2 mutation can be transferred by conventional breeding; it is arecessive trait.[4]Gene editing can be used to induce a similar mutation in ordinary rice, turning them aromatic without affecting other traits.[5]
Thearomaticcultivar group originated in the Indian subcontinent about 2400–4000 years ago, from an admixture of the newly-introducedjaponica rice (carrying the BADH2 mutation) and the localaus rice. Not all members of the cultivar group are fragrant.[6] Thearomatic group is considered part of thejaponica subspecies.[7]
^Ghareyazie, Behzad; Alinia, Faramarz; Menguito, Corazon A.; Rubia, Leila G.; De Palma, Justina M.; Liwanag, Evelyn A.; Cohen, Michael B.; Khush, Gurdev S.; Bennett, John (1997). "Enhanced resistance to two stem borers in an aromatic rice containing a synthetic cryIA(b) gene".Molecular Breeding.3 (5):401–414.doi:10.1023/A:1009695324100.S2CID34294922.