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Khoratpithecus

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Extinct genus of primates

Khoratpithecus Temporal range:Late Miocene, ~9–7 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓

Holotype mandible ofKhoratpithecus ayeyarwadyensis fromMyanmar
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	Primates
Suborder:	Haplorhini
Family:	Hominidae
Subfamily:	Ponginae
Tribe:	Pongini
Genus:	†Khoratpithecus Chaimaneeet al. 2004^[1]
Type species
†Khoratpithecus piriyai Chaimaneeet al. 2004
Species
K. piriyai^[1] K. ayeyarwadyensis^[2] K. chiangmuanensis^[3]

Khoratpithecus is anextinct genus ofpongin primates that lived during theLate Miocene (7–9 million years ago) inMyanmar andThailand.

Three species belong to this genus:

Khoratpithecus chiangmuanensis from Thailand(Chaimanee, Jolly, Benammi, Tafforeau, Duzer, Moussa & Jaeger, 2003) (formerlyLufengpithecus chiangmuanensis)
Khoratpithecus piriyai from ThailandChaimanee, Suteethorn, Jintasakul, Vidthayanon, Marandat & Jaeger, 2004
Khoratpithecus ayeyarwadyensis from MyanmarJaeger, Soe, Chavasseau, Coster, Emonet, Guy, Lebrun, Maung, Shwe, Tun, Oo, Rugbumrung, Bocherens, Benammi, Chaivanich, Tafforeau & Chaimanee, 2011

Palaeobiology

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Stable isotopic analysis consisting of pairedδ¹³C andδ¹⁸O measurements shows thatK. ayeyarwadyensis inhabited the treetops of highly forested habitats and fed exclusively onC₃ vegetation.^[4]Dental microwear and dental topography both indicate thatKhoratpithecus wasfrugivorous, preferring soft fruits over seeds and hard fruits.^[5]

References

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Wikispecies has information related toKhoratpithecus.

^^a ^bY. Chaimanee; V. Suteethorn; P. Jintasakul; C. Vidthayanon; B. Marandat; J.-J. Jaeger (2004). "A new orang-utan relative from the Late Miocene of Thailand".Nature.427 (6973):439–441.Bibcode:2004Natur.427..439C.doi:10.1038/nature02245.PMID 14749830.S2CID 4349664.
^J.-J. Jaeger; A. N. Soe; O. Chavasseau; P. Coster; E.-G. Emonet; F. Guy; R. Lebrun; A. A. Maung; H. Shwe; S. T. Tun; K. L. Oo; M. Rugbumrung; H. Bocherens; M. Benammi; K. Chaivanich; P. Tafforeau; Y. Chaimanee (2011)."First Hominoid from the Late Miocene of the Irrawaddy Formation (Myanmar)".PLOS ONE.6 (4):1–14.Bibcode:2011PLoSO...617065J.doi:10.1371/journal.pone.0017065.PMC 3080362.PMID 21533131.
^Y. Chaimanee; D. Jolly; Benammi, M.; Tafforeau, P.; Duzer, D.; Moussa, I. & J.-J. Jaeger (2003). "A Middle Miocene hominoid from Thailand and orangutan origins".Nature.422 (6927):61–65.Bibcode:2003Natur.422...61C.doi:10.1038/nature01449.PMID 12621432.S2CID 4422882.
^Habinger, S. G.; Chavasseau, O.; Jaeger, J.-J.; Chaimanee, Y.; Soe, A. N.; Sein, C.; Bocherens, H. (12 July 2022)."Evolutionary ecology of Miocene hominoid primates in Southeast Asia".Scientific Reports.12 (1).doi:10.1038/s41598-022-15574-z.ISSN 2045-2322.PMC 9276763.PMID 35821257. Retrieved19 August 2025.
^Merceron, Gildas; Taylor, Sarah; Scott, Robert; Chaimanee, Yaowalak; Jaeger, Jean-Jacques (8 April 2006)."Dietary characterization of the hominoid Khoratpithecus (Miocene of Thailand): evidence from dental topographic and microwear texture analyses".Naturwissenschaften.93 (7):329–333.doi:10.1007/s00114-006-0107-0.ISSN 0028-1042. Retrieved20 January 2026 – via Springer Nature Link.

Haplorhini

Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Order:Primates

Haplorhini

†"Omomyidae"

Altanius? Altiatlasius? Bownomomys Kohatius
Microchoerinae	Indusomys Melaneremia Microchoerus Nannopithex Necrolemur Paraloris Pseudoloris Quercyloris Vectipithex
"Anaptomorphinae"	Absarokius Anaptomorphus Anemorhysis Arapahovius Aycrossia Chlororhysis Gazinius Pseudotetonius Sphacorhysis Strigorhysis Tatmanius Teilhardina Tetonius Tetonoides Trogolemur Walshina
"Omomyinae"	Ageitodendron Asiomomys Brontomomys Chipetaia Chumashius Diablomomys Dyseolemur Ekwiiyemakius Gunnelltarsius Hemiacodon Jemezius Loveina Macrotarsius Nesomomys Omomys Ourayia Rooneyia Shoshonius Steinius Stockia Uintanius Utahia Washakius Wyomomys Yaquius
Tarkadectinae	Tarka Tarkadectes Tarkops

Tarsiiformes

†Archicebus †Afrotarsiidae? †"Omomyidae"?
Tarsiidae	Carlito Cephalopachus †Hesperotarsius †Oligotarsius Tarsius †Xanthorhysis?

Simiiformes

- see below↓

Simiiformes

†Afrotarsiidae?

Afrasia
Afrotarsius

†Eosimiidae

†Amphipithecidae

†Parapithecoidea

Arsinoea Biretia
Proteopithecidae	Proteopithecus Serapia
Parapithecidae	Abuqatrania Apidium Parapithecus Qatrania Ucayalipithecus

Platyrrhini

†Parvimico †Perupithecus †Szalatavus †Canaanimico †Branisella †Lagonimico
Aotidae	Aotus †Tremacebus
Pitheciidae	†Antillothrix Cacajao Callicebus †Carlocebus †Cebupithecia Cheracebus Chiropotes †Homunculus †Insulacebus †Mazzonicebus †Miocallicebus †Nuciruptor Pithecia Plecturocebus †Proteropithecia †Soriacebus †Xenothrix
Atelidae	Alouatta Ateles Brachyteles †Caipora †Cartelles †Chilecebus Lagothrix †Mohanamico Oreonax †Paralouatta †Protopithecus †Solimoea †Stirtonia
Cebidae	†Acrecebus Cebus †Dolichocebus †Killikaike †Panamacebus †Patasola Saimiri Sapajus
Callitrichidae	Callimico Callithrix Cebuella Leontocebus Leontopithecus Mico †Micodon Saguinus

Catarrhini	see below↓

Catarrhini

†Micropithecus
†Saadanius

†Oligopithecidae

†Propliopithecidae

†Pliopithecoidea

Krishnapithecus Paidopithex †Kapi
Pliopithecidae	Epipliopithecus Pliopithecus
Dionysopithecidae	Dionysopithecus Platodontopithecus
Crouzeliidae	Anapithecus Barberapithecus Crouzelia Egarapithecus Fanchangia Laccopithecus Plesiopliopithecus Pliobates

Cercopithecoidea

†Alophe

†Victoriapithecidae

Colobinae

Cercopithecinae

Cercopithecini	Allenopithecus Allochrocebus Cercopithecus Chlorocebus Erythrocebus Miopithecus †Nanopithecus
Papionini	Cercocebus †Dinopithecus †Gorgopithecus Lophocebus Macaca Mandrillus Papio †Paradolichopithecus †Parapapio †Pliopapio †Procercocebus †Procynocephalus Rungwecebus †Soromandrillus Theropithecus

Hominoidea	see below↓

Hominoidea

†Afropithecus †Ekembo †Equatorius †Griphopithecus †Kamoyapithecus? †Kenyapithecus †Morotopithecus †Nacholapithecus †Otavipithecus †Proconsul †Samburupithecus
†Dendropithecidae	Dendropithecus Micropithecus? Nyanzapithecus Oreopithecus? Rangwapithecus Rukwapithecus Simiolus Turkanapithecus
Hylobatidae	†Bunopithecus Hoolock Hylobates †Junzi Nomascus Symphalangus †Yuanmoupithecus

Hominidae

†Buronius †Graecopithecus †Nakalipithecus †Ouranopithecus
Ponginae	†Ankarapithecus †Gigantopithecus †Indopithecus †Khoratpithecus †Lufengpithecus †Meganthropus Pongo †Sivapithecus

Pan
Hominina	†Ardipithecus? †Australopithecus †Graecopithecus? Homo †Kenyanthropus †Orrorin? †Sahelanthropus? †Paranthropus

Taxon identifiers
Khoratpithecus	Wikidata:Q143034 Wikispecies:Khoratpithecus EoL:23402862 GBIF:4827700 IRMNG:1255655 Open Tree of Life:4117749 Paleobiology Database:89267

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