Star Fox is a series of spaceship shooter games published byNintendo. The main protagonist andplayer character of the series isFox McCloud, the leader of Star Fox, a team ofanthropomorphicanimals in the Lylatplanetary system. Gameplay involves control of futuristic aircraft called Arwings, as well as other vehicles and combat on foot.

According to programmerDylan Cuthbert, several names of animals were used in the names of the characters whenever the development team believed they, for those characters, "[sounded] good".^[1]

Star Fox,^[c] orTeam Star Fox, is a band of adventurers-for-hire often employed byGeneral Pepper. The team is headed by Fox McCloud.Star Fox 64 fills in the background of the team where at its founding, it was composed ofJames McCloud, Peppy Hare, andPigma Dengar (who later betrays the team and sides with Andross). Peppy later retires from active flight duty shortly beforeStar Fox: Assault and was more or less replaced byKrystal, Fox's love interest. The team's primary aircraft is theArwing, their land craft is theLandmaster, and the team's mothership is known as theGreat Fox. The team is made up of Fox McCloud, Falco Lombardi, Krystal, Slippy Toad, and Peppy Hare.

Fox McCloud^[d] is ared fox, themain character of the series, the leader of the team and James McCloud's son. Fox wears a green suit. When he hears about his father's disappearance, he drops out of the Corneria Defense Force. At the start of the series, Fox is young and still training under Peppy Hare, his father's friend and wingmate. He becomes an expert pilot, skilled in both air and ground-based attacking, as seen inStar Fox: Assault. Fox begins a relationship with Krystal inStar Fox Adventures which continues throughStar Fox: Assault and possibly ends inStar Fox: Command when Fox forces her off the team for fear of her safety, though the outcome can change depending on the route and ending.

Fox is also a playable character in all five games in theSuper Smash Bros. series.

The character is voiced in English by Steve Malpass inStar Fox Adventures,^[2] Jim Walker inStar Fox: Assault,^[3] and Mike West inStarlink: Battle for Atlas^[5] andSuper Smash Bros. Ultimate.^[7] In the Japanese versions, he is voiced byShinobu Satouchi inStar Fox 64,Super Smash Bros., andSuper Smash Bros. Melee,Kenji Nojima inStar Fox: Assault,^[8]Super Smash Bros. Brawl, andSuper Smash Bros. for Nintendo 3DS and Wii U and byTakashi Ohara fromStar Fox 64 3D onwards.^[9]

Falco Lombardi^[e] is a character who appeared in the firstStar Fox game for theSuper Nintendo Entertainment System. He subsequently appeared in the sequel,Star Fox 64 for theNintendo 64. In both games, he served as a member of the Star Fox Team, but a minor character nonetheless.^[10] Falco is a friend of Fox and the team's ace pilot, but he dislikes authority, and his brashness, overconfidence and tendency to act on his own tend to put him in danger. After the defeat of Andross at the end ofStar Fox 64, Falco disappears, and is not seen for a significant time.^[10] Falco does not appear during the majority ofStar Fox Adventures, as he could not be properly integrated into the plot in time for the game's release.^[11] He also appeared inStar Fox: Assault, andStar Fox Command.

Falco made his first appearance outside theStar Fox series inSuper Smash Bros. Melee. With a similar set of moves to that of Fox, Falco is considered to be one of the most viable characters in competitive play.^[12] Falco returned forSuper Smash Bros. Brawl,Super Smash Bros. for Nintendo 3DS andWii U andSuper Smash Bros. Ultimate as a playable character, trophies, and spirits.^[13] Falco is one of the game's fastest characters, and performs his moves very quickly.^[10] He jumps higher than Fox^[14] and uses a similar reflecting shield, blaster, and Landmaster tank. InMelee, Falco's appearance is based on his design inStar Fox 64, his design inBrawl andSuper Smash Bros. for Wii U and 3DS is based on his appearance inStar Fox: Assault andStar Fox Command, while his design inSuper Smash Bros. Ultimate combines elements from his appearance inStar Fox Zero and the previous twoSuper Smash Bros. games.^{[citation needed]}

Falco also appears in theNintendo Switch version ofStarlink: Battle for Atlas, along with the other members of the Star Fox team, joining forces with the Starlink team to save Atlas from the Forgotten Legion.^{[15][16][17][18]}

Farewell, Beloved Falco is amanga series that detailed the eight-year gap betweenStar Fox 64 andStar Fox Adventures. In the manga, Falco receives a request for help fromKatt Monroe. At the conclusion of the manga, he stays with Katt Monroe, while Fox returns to the team, explaining his departure from the Star Fox Team.^[19]

Slippy Toad^[f] is atoad who wears a blue suit. He's a childhood friend of Fox, and they studied together. He serves as theinventor andmechanic of the team. He attended Cornerian Academy, specializing in engineering. Slippy's father Beltino serves as the director of Engineering at Space Dynamics Co. Ltd.,^[20] and later serves the research director for the Cornerian Defense Forces. Slippy invents and constructs many inventions for the team, including the Blue-Marine, the Landmaster and the Roadmaster, creating the former two with his father. After the Aparoid race is destroyed inStar Fox: Assault, he temporarily withdraws from the Star Fox team after falling in love with a frog named Amanda, who returns his feelings, to spend more time with his girlfriend. However, when Fox needs backup inStar Fox Command, he comes to his assistance. In two endings inStar Fox Command, he and Amanda have children together. Due to his lower flying skills compared to his teammates, Slippy Toad tends to fall prey to enemy pilots, and as such calls upon the player for assistance. In bothStar Fox 64 andStar Fox: Assault, the player needs to save Slippy shortly after the start of the first mission.

In theSNES game andNintendo Power comic, Slippy has a croakingstutter, which was removed in later appearances.

Slippy makes a cameo appearance inSuper Smash Bros. Brawl as one ofSolid Snake's contacts on his Codec and sometimes a contact for Fox, Falco, or Wolf with a taunt on the Star Fox battlefield.

He is voiced in English byChris Seavor inSuper Smash Bros. Melee^[21] andStar Fox Adventures^[2] andMike McAuliffe inStar Fox: Assault.^[3] He is voiced byKyoko Tongu in the early Japanese versions of the series;^[8]Star Fox 64 3D andStar Fox Zero saw him being voiced by Kei Hayami.

Peppy Hare^[g] is arabbit who wears a red suit and is a member of the original Star Fox team, with leader James McCloud and wingmate Pigma Dengar. After Pigma's betrayal and James' capture, Peppy barely escapes to return to Papetoon to inform Fox McCloud of his father's fate. Peppy serves as Fox's mentor and gives Fox instructions and advice throughout the course of the games. Peppy's phrase fromStar Fox 64, "Do a barrel roll!", has become aninternet meme.^[22]Google created anEaster egg homage to it in November 2011. Typing "do a barrel roll" or "z or r twice" makes the user's web browser appear to spin.^[23][24] Cuthbert stated that he named Peppy.^[25]

According toStar Fox Command, he has a daughter namedLucy and is the widower of his wife Vivian. By the time ofStar Fox: Assault, he is retired from flight duty, taking up an advisory position in the Great Fox. Also according toCommand, he is eventually made general of the Corneria Army after Pepper grows ill.

In theNintendo Power comic, Peppy appears to have some form ofextrasensory perception, although it is not elaborated upon.

In English, Peppy was voiced byRick May inStar Fox 64,^[26]Chris Seavor inSuper Smash Bros. Melee^[21] andStar Fox Adventures,^[2] Henry Dardenne inAssault, and Jaz Adams inStar Fox 64 3D andStar Fox Zero.^[3] He was originally voiced by Tomohisa Asō in the Japanese versions of the series;^[8] Starting withStar Fox 64 3D, Kunpei Sakamoto took the rein of the character.

Krystal^[h] is a bluevixen who is the sole survivor of her doomed home planet, Cerinia. Krystal is calm and kind hearted. She first appears in the series in the early conceptDinosaur Planet and thenStar Fox Adventures, where she searches for answers on her planet's destruction and the death of her parents. She comes across a distress signal from Sauria. She joins Star Fox as the team'stelepath after the adventure. She fights in multiple battles inStar Fox: Assault. Before the start ofStar Fox Command, Krystal leaves the Star Fox team after a strained relationship with Fox. Several possible endings show her returning to both, including one where she marries Fox and their son Marcus leads a new Star Fox team decades in the future. InStar Fox: Assault, Krystal wears a bluecatsuit, blueknee-high boots, a silverchain belt, two silverrings on her tail, whitebeads in her hair, and a silverdiadem which has a turquoise jewel.

Krystal wore a loincloth and was originally designed to be a main character ofDinosaur Planet, before Nintendo turned the project into aStar Fox title. Her original design was that of a 16-year-old, who was raised by a wizard named Randorn. When Fox became the main character ofStar Fox Adventures, Krystal was redesigned into a 19-year-old and her role in the game became less central.

She makes a cameo appearance starting inSuper Smash Bros. Brawl as one of the contacts for Fox, Falco, or Wolf with a taunt on the Lylat Cruise stage. She appears as an Assist Trophy inSuper Smash Bros. Ultimate.

In English versions of the series, Krystal speaksEstuary English and is voiced by Estelle Ellis inStar Fox Adventures,^[2] and byAlésia Glidewell inStar Fox: Assault,^[3]Super Smash Bros. Brawl andSuper Smash Bros. Ultimate. In the Japanese version ofAssault, Krystal is voiced byAya Hara.^[8]

InStar Fox 2, completed in 1995 but officially released in 2017, the charactersMiyu,^[i] a femalelynx in a red suit, andFay,^[j] a femaledog in a blue suit, were added to the Star Fox team; neither of them were featured in games released after the making ofStar Fox 2.Dylan Cuthbert stated that he gave Miyu that name because he liked how it sounded, and that he named Fay after a crush he had in childhood prior to his teenage years.^[27] Cuthbert stated he did not remember whether he gave family names to Miyu and Fay.^[28]

TheNintendo Power'sStar Fox comic by Benimaru Itoh exclusively introduced the characterFara Phoenix, aFennec fox, as a love interest for Fox, starting off as a Cornerian test pilot and later becoming an unofficial member.

ROB 64 (known asNUS64^[k] in the original Japanese-language version) is the observation and analysis robot of Star Fox's mothership the Great Fox. ROB's name is a reference to theNES accessoryR.O.B.^{[citation needed]} The 64 is derived from the console of hisfirst appearance, theNintendo 64. ROB's Japanese name, "NUS", stands for "Nintendo Ultra Sixty-four", the original name of the Nintendo 64 system, and is part of the serial number of all N64 components: the controller, for example, is NUS-005.^[29]

ROB has the ability to completely control the Great Fox, piloting it, providing information to the team, and aiding them with items and vehicles. Prior toStar Fox Adventures, he has been rebuilt and upgraded several times by Slippy Toad, infusing him with more productivity and personality. After the first Great Fox is destroyed inStar Fox: Assault, he returns inStar Fox Command piloting the new version of the Great Fox. On some paths in the game, ROB joins Star Wolf when no "hero" character is available.

InStar Fox Zero, a tethered robot similar in appearance, Direct-i, also assists in operation of the Gyrowing and Roadmaster.

ROB is voiced in English by John Silke inStar Fox Adventures^[2] andDex Manley inStar Fox: Assault.^[3] He is voiced byDaisuke Sakaguchi in the Japanese version ofStar Fox 64,Yusuke Numata inStar Fox: Assault^[8] andAtsushi Abe inStar Fox 64 3D.

Tricky^[l] is the youngStyracosaurus^[30] prince of the EarthWalker Tribe from the planet Sauria. He serves as Fox's companion inStar Fox Adventures, in which Tricky performs various tasks for Fox, such as digging or breathing fire. He is made an honorary Star Fox member at the end ofAdventures. In his second appearance inStar Fox: Assault, Tricky has matured, and is now the king of the EarthWalker Tribe. After Fox and Krystal save his kingdom from the Aparoids, he promises to help repair Sauria.

He was voiced by Kevin Bayliss inStar Fox Adventures^[2] and Chet Morgan inStar Fox: Assault.^[3] In the Japanese version ofStar Fox: Assault, he is voiced byHirohiko Kakegawa,^[8] who also voiced Beltino and James McCloud in the same game. Tricky also cameos inSuper Smash Bros. Brawl as a collectible trophy.

Tricky was originally intended to be the same character as "Tricky the Triceratops" inDiddy Kong Racing. Nintendo now owns the rights to the Tricky character.^[31]

Star Wolf^[m] is a group of mercenaries originally created and hired by Andross to eliminate the Star Fox team. They have since been working on their own as Star Fox's rival team, occasionally helping Star Fox inStar Fox: Assault. Their primary aircraft is theWolfen. Though Star Wolf were planned to appear on the canceledStar Fox 2, they first appeared inStar Fox 64. They have appeared in every game except for the originalStar Fox andStar Fox Adventures. InStar Fox 64 andStar Fox Zero, the team is composed of Wolf O'Donnell, Leon Powalski, ex-Star Fox team member Pigma Dengar, and Andross' nephew Andrew Oikonny, while inStar Fox: Assault andStar Fox Command, the team is composed of Wolf O'Donnell, Leon Powalski, and Panther Caroso.

Wolf O'Donnell^[n] (sometimes called Lord O'Donnell) is awolf, the leader of Star Wolf team, and Fox's rival, fueled by his prior rivalry with his father, James.^[32] He was said to be skillfully goaded by Pigma Dengar into forming Star Wolf and work for Andross to take down the Star Fox team.^[32] After the Star Wolf team fails their mission and the Venom army is destroyed, Wolf and his team go their separate way.^[33]

In the English games, Wolf is voiced byGrant Goodeve inStar Fox: Assault^[3] and by Jay Ward inSuper Smash Bros. Brawl,Starlink: Battle for Atlas,^[5] andSuper Smash Bros. Ultimate.^[7] In the Japanese versions, Wolf is voiced byHisao Egawa inStar Fox 64,Mahito Oba inStar Fox: Assault^[8] andSuper Smash Bros. Brawl, and Kosuke Takaguchi inStar Fox 64 3D,Star Fox Zero,Starlink: Battle for Atlas andSuper Smash Bros. Ultimate.^[9]

Leon Powalski^[o] is achameleon who is a member of the Star Wolf team, and is the only member other than Wolf that has been on the team since its creation. Leon's past is unknown, with no records of him belonging to any prior organization, and he is also believed to be an alien that migrated from another star system,^[32] but he most likely had some past rivalry with Falco Lombardi whom he mainly concentrates on during dogfights inStar Fox 64 and taunts mercilessly withinStar Fox: Assault.

Leon's English personality has changed noticeably sinceStar Fox 64. He has gone from a classy, competent pilot to a crazed, ruthless assassin who usually gives a deranged laugh after defeating a foe. His English voice has also changed to suit this; originally, his tone was calm and sinister, but it has become high-pitched and maniacal.

Despite his ruthless swagger and love of the hunt, Leon loves to adorn his maniacal disposition to maintain his spot as the assassin on the Star Wolf team. He is somewhat of anarcissist, and enjoys his feared reputation.

His personality inStar Fox Command seems to contradict his cold-blooded image. It is implied that, beneath his facade, he has a soft spot for peace. This is hinted at in his profile in the Pilot Gallery and his rejoicing after defeating the Anglar Emperor because he claims that he will look forward to parades with flowers and other such niceties. Leon has a customized ship called the Rainbow Delta, which may allude to his supposed soft spot or reference the fact that chameleons can change color.

Leon has been voiced byShinobu Satouchi in most of the Japanese versions and byTakashi Ohara inStar Fox 64 3D andStar Fox Zero. InStar Fox: Assault, he and Panther Caroso were voiced in English by David Scully.

InStar Fox: Assault, Leon and Panther Caroso were voiced in English by David Scully.^[3] The former has been voiced byShinobu Satouchi in most of the Japanese versions^[8] and byTakashi Ohara inStar Fox 64 3D andStar Fox Zero.

Pigma Dengar^[p] is apig who was one of the original members of the Star Fox team. He is a greedypirate and mechanic with a twisted personality, who is willing to deceive others and does not care who he hurts so long as he is paid.^[32] Pigma is also sadistic as evidenced by his introductory quote from the Bolse level.^[34] Said to be a researcher under Andross' direct supervision during his time at the Corneria Defense Force Scientific Research Institute, he eventually schemes with Andross to betray the Star Fox team, causing the presumed death of James McCloud. Afterwards, he persuades Wolf to create and lead the Star Wolf team, secretly manipulating him and his team in accordance to Andross' orders.^[32] When fighting Star Fox inStar Fox 64, Pigma targets Peppy due to their previously having been teammates. Pigma is also said to have created the Star Fox team emblem,^[35] as well as (in collaboration with Andross) custom engineering the Wolfen for the Star Wolf team to utilize.^[32]

Pigma is later forcibly fired from Star Wolf by Wolf O'Donnell for his greed and treachery. He attempts to profit off a Memory Cube inStar Fox: Assault. As a result of his greed, he undergoes Aparoid assimilation which fuses him with a spacecraft. Fox McCloud defeats him, causing him to explode. He nonetheless returns and serves as a boss inStar Fox Command, revealing that he survived the previous encounter as a cube-shaped cyborg. Whether the player encounters him depends on decisions made during gameplay.

According toStar Fox Zero, Pigma Dengar managed to volunteer as a member of Star Fox shortly before betraying the team. InStar Fox Guard, he is shown to have some degree of mechanical expertise.

In the English version ofStar Fox: Assault, Pigma is voiced by Lev Liberman.^[3] He was voiced byDaisuke Gōri in the early Japanese versions,^[8] andTsuguo Mogami in the Japanese version ofStar Fox 64 3D,Star Fox Zero andStar Fox Guard.

In the Japanese version ofStar Fox 64, Pigma speaks in theKansai dialect. His last name refers to the way people who speak in Kansai often end their sentences with "dengar".^[36] The name, "Pigma", is a spin on the worddogma, pointing to his antidogmatic nature.

Andrew Oikonny^[q] is amonkey who is the nephew ofAndross. InStar Fox 64, Andross is said to have effectively forced Pigma to include him on the team, as he is otherwise an inexperienced pilot with an elitist attitude, using his uncle's authority as a shield to be arrogant, causing him to be unpopular with his crew mates; essentially, the only reason Wolf tolerates him is because he is using his connections to Andross to provide the team with immeasurable technological and financial support.^[32] In battle, he primarily targets Slippy usually saying, "Stick to the pond, froggie".

According to the OfficialStar Fox 64 Player's Guide his favorite book is"The Apes of Wrath".

Sometime after Andross is destroyed, Oikonny, no longer having the support of his uncle, is kicked out of Star Wolf,^[33] and from there he plans to avenge his uncle's death by gathering the remnants of the Venom army to stage a rebellion against Corneria, as seen in the beginning ofStar Fox: Assault, where Oikonny pilots his ship resembling Andross on Fortuna, but a large Aparoid destroys it.

He returns inStar Fox Command where he joins the Anglars and pilots the Death Crab. He again attempts to get revenge on Fox and confronts him and his team in Fichina.

InStar Fox 2, a similar character named Algy appears as a member of the Star Wolf wing, who seems to be a lemur and is unrelated to Andross.

In English, he is voiced by John Hugil inStar Fox: Assault.^[3] In the Japanese versions of the series, he is voiced byDaisuke Sakaguchi inStar Fox 64,Yūsuke Numata inStar Fox: Assault,^[8] andAtsushi Abe inStar Fox 64 3D andStar Fox Zero.

Panther Caroso (known asPanther Caluroso^[r][37] in Japanese-language versions) is ablack panther who makes his first appearance as the newest member of Star Wolf inStar Fox: Assault. His signature symbol, which is on his Wolfen, is a redrose. Panther is conceited and a flirt, considering himself aladies' man. In the game, he unsuccessfully tries to flirt with Krystal several times.

Panther reappears inStar Fox Command, where he attempts to begin a relationship with Krystal after she joins the Star Wolf team following her breakup with Fox. In the English version ofCommand, Panther speaks in the third person, despite his normal speaking habits inStar Fox: Assault.

He is voiced in English by David Scully inStar Fox: Assault.^[3] His Japanese voice is provided byTetsu Inada in bothStar Fox: Assault^[8] andSuper Smash Bros. Brawl.

Andross's army^[s] (also known as theImperial Guard^[t]) is the main army of the planet Venom, which operates under Andross. Although most of the army is decimated by the Star Fox team and the Corneria Army, a small group of them are present for the Aparoid invasion.

Andross (known asAndorf^[u][38] in the Japanese versions) is the mainantagonist of the series, directly appearing in every game except forStar Fox: Assault, and has beenFox McCloud'sarchenemy ever since his defeat against him at the end of the Lylat Wars.

Before the series, Andross was ascientist working for Corneria. His original intentions are supposedly pure and beneficial; he devotes most of his life to finding ways to protect Corneria. However, a growing obsession with his engine and bio-technology drives him to the brink of madness, despite multiple warnings from the planet's ruling council. The increasing perversion of his experiments results in unleashing a deadly weapon within a Cornerian city, which may or may not have been accidental.

Subsequently captured by General Pepper and banished for treason to the wasteland planet Venom, he builds an army and plots to dominate the Lylat System and rebuild it in his name. The original Star Fox team is sent to investigate. The investigation ends when Pigma Dengar defects to Andross's side and sends his friends to Andross, resulting in the capture (and possible death) of James McCloud. Andross then proclaims himselfemperor of the Lylat System, and begins to invade the neighboring worlds, initiating the Lylat Wars.

Five years later after a new Star Fox team is formed under Fox McCloud, they begin their mission to liberate the Lylat System from Andross. Upon arriving at Venom, Fox engages Andross in a duel. Andross is shown to have apparently experimented with himself and become a giant disembodied head, but Fox is able to defeat him and Andross self-destructs.

Andross' spirit survives. After a botched attempt at a revival on Titania, Andross travels to Sauria (also known as Dinosaur Planet). Weakened, Andross takes refuge in Krazoa Palace, and uses the SharpClaw Tribe to keep him alive. He manipulates their leaderGeneral Scales into collecting various artifacts infused with the spirits of the Krazoa and imprisons Krystal in the palace. Through her, he is able to generate the energy from the Krazoa Spirits to restore himself back to full power, intending to obliterate the entire Lylat System. Fox again defeats him, after which Andross again self-destructs.

While he does not appear inStar Fox: Assault, his reign continues in his nephew Andrew Oikonny, who constructs a new fleet with Andross' remaining Venomian units. They attack Fortuna at the beginning of the game, with Andrew using a robotic copy of his uncle as the game's first boss.

He reappears inStar Fox Command as a spirit in control of a bio-weapon, but is no longer a threat.

Dialogue between Zazan and the Anglar Emperor in "Fox and Krystal" implies that the Anglar race was created by Andross.

Andross appears inSuper Smash Bros. Brawl as an Assist Trophy character, resembling his metallic-looking form from the firstStar Fox game.

According toStar Fox Zero, Andross secretly worked on a Cornerian teleportation device for peacekeeping purposes, but his lust for power made him claim it as his own for his schemes. After discovering the truth, General Pepper and James McCloud used the teleporter to exile Andross to another dimension.

InStar Fox 2, a direct sequel to the original game, Andross is revealed to have survived his encounter with Fox McCloud and his army advances towards Corneria. The Star Fox team hastily scrambles to destroy Andross once again.

Andross appears in several different forms throughout theStar Fox series:

• In the first game, he takes the form of a large metallic head (or possibly a hologram) that contains a cube covered in images of his face.
• InStar Fox 64 andStar Fox Adventures, the shell is a large version of his head, which contains an enlarged brain that can act and move independently. The latter Andross is more demonic-looking than the other, resembling a largegorilla head. InStar Fox: Assault, his nephew Andrew uses a robot with a similar design.
  • InStar Fox Zero, Andross is first shown in a form that outwardly resembles his appearance inStar Fox 64, but he is shown to have become robotic.
  • InStar Fox 2, his image appears as a disheveled version of his previous self with a cybernetic implant over the left side of his head, and his boss form initially appears as a mask and later a form somewhat resembling hisStar Fox 64 self.

Andross was voiced byRick May in the English version ofStar Fox 64.^[26] He was also voiced byDaisuke Gōri in the Japanese version ofStar Fox 64 and byTsuguo Mogami inStar Fox 64 3D andStar Fox Zero.

Captain Shears^[v] is adog who appears in the mangaStar Fox: Farewell, Beloved Falco, which was released on the officialStar Fox Adventures Japanese website. Shears conducts his operations in a base on Titania. Working as a double agent, he tells the Star Fox team he and his group of scientists are planning to reconstruct Andross's research, claiming there might still be Venom soldiers on the planet. Katt Monroe does not trust him and tells Falco of Shears' plan, leading to a dogfight between Fox and Falco. While Fox prevails, Falco is unharmed.

The Star Fox team and Katt's gang team up and learn Shears is not planning to reconstruct Andross' research, but to resurrect Andross. Fox storms into Shears' base and fights him. After losing, Shears attempts to escape, only to be crushed to death under the weight of the newly constructed Andross.

General Scales^[w] appears inStar Fox Adventures. Scales leads the SharpClaw Tribe, which strives to be the ruling tribe of Sauria. After being rejected by the other tribes, General Scales amasses an army of Sharpclaws and breaks four pieces of the planet apart by removing the four SpellStones where his army occupies each chunk. Andross gives him power in return for capturing Krystal and the Krazoa Spirits, while keeping himself relatively unknown to Scales at the same time. At the end of the game, Andross stops the fight between Scales and Fox and has him relinquish the last Krazoa spirit to Fox and collapses. Before he relinquishes the Krazoa Spirit, General Scales tells Fox that he will regret this.

General Scales is voiced by John Silke.^[2]

TheAparoid Queen^[x] is the leader of the Aparoid race and the primary antagonist ofStar Fox: Assault. From within the Homeworld, the Queen rules over the Aparoids like a queen bee over her hive. Beltino Toad determines that if the Queen is infected with a self-destruct program, then the rest of the Aparoids will die along with her. She can emulate personalities of people who are infected, or people close to her enemies. Star Fox make their way into the Queen's chamber, and infect her with the self-destruct program. The rest of the Aparoids are wiped out. She seems to take a peaceful approach when directly confronted, as she attempts to convince the Star Fox team to give in. She describes the assimilation as "not sacrifice, but evolution". She also believes that everything in the universe exists for the Aparoids (or herself).

TheAnglar army^[y] are a race ofanglerfish-like creatures that reside in the Toxic Seas of Venom. They first appeared inStar Fox Command. It is implied in the Venom Level of the storyline "Fox and Krystal" that the Anglar were created by Andross through mutations caused by his experiments on Venom's oceans.

TheAnglarEmperor^[z] is the leader of the Anglar race, and the primary antagonist ofStar Fox Command. Originating on Venom, he appears as a final boss for several missions. He pilots a large serpent-like creature known as the Arrowhead. In some of the final missions, he becomes a very largeangler fish, capable of taking the player's ship into his mouth, which is reminiscent of an attack used byAndross in other games.

Octoman^[aa], anoctopus-like creature, is one of the Anglar Emperor's henchmen.

He is based on Octoman from theF-Zero franchise.

Zako^[ab] is ahammerhead shark-like bounty hunter. He is defeated in the Venom Sea.

Zazan^[ac] is an Anglar who led the attack on Corneria. He is defeated during the fight on Venom.

Zoldge^[ad] is asquid-like mercenary that pilots the Solar Satellite. He is defeated when the Solar Satellite is shot down.

General Pepper^[ae] is thebloodhoundCommander-in-chief of the Corneria Defense Force.

He originally exiles Andross to Venom and hires the Star Fox team to investigate it five years later. Two more years later, he becomes the primary source for the new Star Fox's missions. Pepper assigned Star Fox to investigate Dinosaur Planet (Sauria) inStar Fox Adventures.

During the Aparoid invasion inStar Fox: Assault, Pepper comes close to death when Aparoids fuse to him and his flagship. At the General's command, Fox destroys the ship with Pepper still on board. He is rescued by Peppy Hare.

Afterwards, Pepper grows ill and retires his position as general of the Corneria Army. He passes this role on to his friend Peppy Hare prior to the events ofStar Fox Command.

In English, he is voiced by John Silke inStar Fox Adventures^[2] and Gray Eubank inAssault.^[3] In the Japanese version ofStar Fox 64, he was voiced byDaisuke Gōri,Michihiro Ikemizu inStar Fox: Assault,^[8] and currently byTsuguo Mogami inStar Fox 64 3D andStar Fox Zero.

James McCloud^[af] is Fox's father and the principal founder of the Star Fox team. On a scouting mission to Venom, he was betrayed by a former member of Star Fox named Pigma Dengar and the team is captured by the evil Andross. In the ensuing conflict, James McCloud courageously creates a diversion so that Peppy Hare could escape, and is presumed dead. He appears toFox McCloud inStar Fox 64 to lead Fox out of Andross's exploding base and inStar Fox Command during a final boss fight.

James makes his first playable appearance inCommand piloting an Arwing like Peppy which resembles theAssault design. None of the other characters acknowledge him and Fox questions if he is seeing things.

He also appears inStar Fox Zero, where Andross acknowledges him by loudly asking why he won't stay dead.

In theStar Fox for SNES, it is stated that Fox's father is missing and was last seen at a black hole. However, the Black Hole is actually a warp point that was created by Andross' experiments. In theNintendo Power comic, he is given the name Fox McCloud Sr.

InStar Fox: Assault, the Aparoid Queen uses James' voice, hinting that James may have been infected at one point. It is also possible that the Aparoid Queen learned of James through the memories of those the Aparoids had infected.

James is also the name of a character in theF-Zero series. The instruction booklet forF-Zero X describes him as working to raise money for the debts of his mercenary organization, "Galaxy Dog". He also uses a racing car similar to an Arwing.

James was voiced by Tomohisa Asō inStar Fox 64, and byHirohiko Kakegawa inStar Fox: Assault. InStar Fox 64 3D, James is voiced by Kunpei Sakamoto.

Katt Monroe^[ag] is a female pilot and an old acquaintance of Falco from their days as a part of a space gang known as "FREE-AS-A-BIRD".^[32] She who comes to the aid of the Corneria Defense Force and the Star Fox team. Katt resembles acat, and appears with pink fur inStar Fox 64 andStar Fox Zero, but appears inStar Fox Command with black fur.

She appears inStar Fox 64 on the planet Zoness, and will either appear to defend the Great Fox in Sector Z, or shoot switches on Macbeth. She has a crush on Falco Lombardi, wishing to join Star Fox to be close to him. InStar Fox Command, she appears in missions with Falco in the Asteroid Belt and Sector X. She and Falco have an argument in which Falco insults her and Katt vows to never help him again, though later in the storyline, she does say she "can't just sit and watch him from the sidelines".

In the manga, "Farewell, Beloved Falco", Katt appears to have a romantic relationship with Kool, another cat with blue fur. Though she implies she likes him, she still shows a liking for Falco.

Katt is voiced byKyōko Tongū in the Japanese version ofStar Fox 64, and Kei Hayami in the Japanese version ofStar Fox 64 3D andStar Fox Zero.

There are variousDinosaurTribes that live on Sauria as seen inStar Fox Adventures.

When the Aparoids invaded Sauria duringStar Fox: Assault, some of the Dinosaur Tribes lost some of their members to the battle.

TheCloudRunner Tribe^[ah] are one of the two ruling tribes on Sauria. They resemblePteranodons.

TheEarthWalker Tribe^[ai] are one of the two ruling tribes on Sauria. They resemble variousCeratopsid genera.

TheHighTop Tribe^[aj] are the largest dinosaur tribe on Sauria. They are based on theApatosaurus and otherSauropods.

TheLightFoot Tribe^[ak] are a secretive and reclusive tribe on Sauria. They are loosely based on theOrnitholestes and the Chief LightFoot has a frill similar to that of afrilled lizard and theJurassic Park version of theDilophosaurus. They live in the LightFoot Village between ThornTail Hollow and Cape Claw. The LightFoots do not interact with other Tribes.

TheRedEye Tribe^[al] are a tribe of carnivorous dinosaurs that are locked up in the Walled City on Sauria by the EarthWalkers. They are based on theTyrannosaurus. The RedEyes are released from the Walled City by General Scales.

TheShadowHunters are a tribe of dinosaurs that are mostly seen in the Test of Fear. They evoke the appearances of theVelociraptor andCoelophysis.

TheSharpClaw Tribe^[am] are a race of humanoidAllosaurus that are more advanced and violent than the other tribes. The SharpClaw originated from a fragment of the second moon of Sauria which crashed into the planet and became known as the Moon Mountain Pass. Eventually, they left their home and spread.

TheSnowHorn Tribe^[an] are a race ofWoolly Mammoths that reside in the SnowHorn Wastes.

TheThornTail Tribe^[ao] are a peaceful tribe of dinosaurs that live in ThornTail Hollow on Sauria. They are based on theNodosaurus.

Beltino Toad^[ap] is Slippy Toad's father. He works as the research director for the Corneria Defense Force. Beltino originally worked for Space Dynamics Co. Ltd. as an engineer. He and Slippy construct many inventions for the Star Fox team, including the Blue-Marine and the Landmaster. Beltino is first mentioned in theStar Fox 64 Player's Guide, but does not make an in-game appearance untilStar Fox: Assault. Beltino discovers the way to defeat the Aparoids. Star Fox destroys them using a program he developed. In the English version ofAssault, Beltino Toad is voiced by Scott Burns,^[3] and his Japanese voice actor isHirohiko Kakegawa.^[8]

Amanda Toad^[aq] is Slippy's fiancée, who first appears inStar Fox Command. They meet two years prior to the events ofStar Fox Command, and instantly fall in love. Amanda occasionally helps the team out. She pilots theTadpole,^[ar] a ship armed with a multi-lock. She considers herself more of the leader in her relationship with Slippy, and always wants to be on his side during a fight. In two endings, she and Slippy settle down with children, one of whom joins Fox and Krystal's son, Marcus, as well as Lucy Hare's daughter and Falco Lombardi himself in forming a new Star Fox team. In another ending, she herself joins the Star Fox team to be close to Slippy.

TheCorneria Army,^[as] also known as theCorneria Defense Force,^[at] is one of the main military forces on Corneria. Their purpose is to protect Corneria and all of the Lylat System from all types of threats. Besides General Pepper, Captain Shears, and the generic soldiers, among the members of the Corneria Army are:

Bill Grey^[au] is adog who is Fox's friend and appears to help him inStar Fox 64,Star Fox Command andStar Fox Zero. The two were originally in training together in the Cornerian Academy, and after Fox leaves to join Star Fox, Bill rises through the ranks and is elected commander of the Husky and Bulldog squadrons of Corneria. In his first two games, he assists Fox when Star Fox reaches Katina. The name of the character is a reference to General William Grey, from the 1996 blockbuster filmIndependence Day. The battle on Katina inStar Fox 64 also greatly resemblesIndependence Day. Bill is voiced byDaisuke Sakaguchi in the Japanese version ofStar Fox 64, andAtsushi Abe in the Japanese version ofStar Fox 64 3D andStar Fox Zero.

Dash Bowman, known asAsh Bowman^[av] in the Japanese version, is amonkey who is a young pilot for the Corneria Army, as well as the grandson of Andross and cousin once removed to Andrew Oikonny. He thinks very highly of Fox and Falco, and wishes to join the Star Fox team. He makes his first appearance inStar Fox Command. Despite his grandfather's dark ways, he still appreciates the man's forgotten aspirations, and wishes to carry on his legacy of making Lylat a better place. Dash's future remains uncertain. One ending shows him joining Falco and Katt to form the Star Falco team, while another two show him becoming ruler of Venom. One of the latter shows him converting Venom into a peaceful planet and the other has him becoming corrupt with power like Andross.

Lucy Hare^[aw] is arabbit who debuts inStar Fox Command, and is Peppy Hare's only daughter. Her motherVivian Hare^[ax] died shortly after the Lylat Wars due to an illness. She is a bit of atomboy, but is still very polite to her elders. She is an astrophysics teacher on the planet Fichina, but wants to become a pilot like her father. She and Krystal are good friends. In some missions, she fights the Anglar attackers in her aircraft, theSky Bunny,^[ay] armed with plasma lasers and a single lock. In one of the endings, her daughter becomes a member of the new Star Fox team, led by Marcus, Fox's son.

Grippy Toad^[az] is Slippy's uncle who appeared inStar Fox Guard.

When reviewingStar Fox Adventures,GameSpot reviewer Greg Kasavin commented that the game has a likable cast of characters who manage to be cute and pretty cool at the same time.^[39]