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- Article title
- Abstract
- Keywords
- Introduction
- Materials and methods
- Type revision, morphological terminology, and classification
- Geological settings
- Geometric morphometric analyses of forewing shape
- Shape discrimination at different taxonomic levels
- Assignment of the bee fossils
- Results
- Geometric morphometric analyses
- Systematics
- Family Apidae Latreille
- Late Eocene
- Eocene-Oligocene boundary
- Oligocene-Miocene boundary
- Lower Miocene
- Middle-Lower Miocene
- Upper Miocene
- Discussion
- Geometric morphometrics of forewing shape to discriminate taxa
- Taxonomic affinities of Calyptapis florissantensis and Oligobombus cuspidatus
- Origin and diversification of bumble bees
- Cenozoic extinctions of Corbiculata
- Acknowledgements
- References
- Supplementary materials
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