Wild fisheries

Crab boat from theNorth Frisian Islands working in theNorth Sea

Global harvest of aquatic organisms in million tonnes, 1950–2010, as reported by the FAO^[1]

Awildfishery is anaturalbody of water with a sizeable free-rangingfish or otheraquatic animal (crustaceans andmolluscs) population that can beharvested for its commercial value. Wildfisheries can bemarine (saltwater) orlacustrine/riverine (freshwater), and rely heavily on thecarrying capacity of the localaquatic ecosystem.

Wild fisheries are sometimes calledcapture fisheries. The aquatic life they support is not artificially controlled in any meaningful way and needs to be "captured" or fished. Wild fisheries exist primarily in the oceans, and particularly aroundcoasts andcontinental shelves, but also exist inlakes andrivers. Issues with wild fisheries areoverfishing andpollution. Significant wild fisheries have collapsed or are in danger of collapsing, due to overfishing and pollution. Overall, production from the world's wild fisheries has levelled out, and may be starting to decline.

As a contrast to wild fisheries,farmed fisheries can operate in sheltered coastal waters, in rivers, lakes andponds, or in enclosed bodies of water such as pools orfish tanks. Farmed fisheries are technological in nature, and revolve around developments inaquaculture. Farmed fisheries are expanding, andChinese aquaculture in particular is making many advances. Nevertheless, the majority of fish consumed by humans continues to be sourced from wild fisheries. As of the early 21st century, fish is humanity's only significant wildfood source.

According to theFood and Agriculture Organization (FAO), theworld harvest bycommercial fisheries in 2010 consisted of 88.6 milliontonnes ofaquatic animals captured in wild fisheries, plus another 0.9 million tons ofaquatic plants (seaweed etc.). This can be contrasted with 59.9 million tonnes produced infish farms, plus another 19.0 million tons of aquatic plants harvested inaquaculture.^[1]

The productivity of marine fisheries is largely determined bymarine topography, including its interaction withocean currents and the diminishment of sunlight with depth. Marine topography is defined by variouscoastal and oceanic landforms, ranging from coastalestuaries andshorelines; tocontinental shelves andcoral reefs; to underwater anddeep sea features such as ocean rises andseamounts.

Anocean current is continuous, directed movement ofocean water. Ocean currents are rivers of relatively warm or cold water within the ocean. The currents are generated from the forces acting upon the water like the planet rotation, the wind, thetemperature andsalinity (henceisopycnal) differences and thegravitation of the moon. Thedepth contours, theshoreline and other currents influence the current's direction and strength.

More on currents

Ocean currents can flow for thousands of kilometers. Surface ocean currents are generally wind driven and develop their typical clockwise spirals in the northern hemisphere and counter-clockwise rotation in the southern hemisphere because of the imposed wind stresses. In wind driven currents, theEkman spiral effect results in the currents flowing at an angle to the driving winds. The areas of surface ocean currents move somewhat with theseasons; this is most notable in equatorial currents. Deep ocean currents are driven by density and temperature gradients.Thermohaline circulation, also known as the ocean's conveyor belt, refers to the deep ocean density-drivenocean basin currents. These currents, which flow under the surface of the ocean and are thus hidden from immediate detection, are calledsubmarine rivers.Upwelling anddownwelling areas in the oceans are areas where significant vertical movement of ocean water is observed. Surface currents make up about 10% of all the water in the ocean. Surface currents are generally restricted to the upper 400 meters of the ocean. The movement of deep water in the ocean basins is by density driven forces and gravity. The density difference is a function of different temperatures and salinity. Deep waters sink into the deep ocean basins at high latitudes where the temperatures are cold enough to cause the density to increase. The main causes of currents are: solar heating, winds and gravity. Ocean currents are also very important in the dispersal of many life forms. A dramatic example is thelife-cycle of the eel. Currents also determine the disposition ofmarine debris.

North Atlantic gyre North Atlantic gyre North Atlantic gyre Indian Ocean gyre North Pacific gyre South Pacific gyre South Atlantic         gyre World map of the five major ocean gyres Oceanic gyres are large-scaleocean currents caused by theCoriolis effect. Wind-driven surface currents interact with these gyres and the underwater topography, such as seamounts and the edge of continental shelves, to produce downwellings andupwellings.[3] These can transport nutrients and provide feeding grounds for plankton eatingforage fish. This in turn draws larger fish that prey on theforage fish, and can result in productive fishing grounds. Most upwellings are coastal, and many of them support some of the most productive fisheries in the world, such as small pelagics (sardines, anchovies, etc.). Regions of upwelling include coastalPeru,Chile,Arabian Sea, westernSouth Africa, easternNew Zealand and theCalifornia coast. External image Animation of the upwelling process.

In the ocean, thefood chain typically follows the course: Phytoplankton → zooplankton → predatory zooplankton →filter feeders → predatory organisms (usually constituting several trophic levels) Phytoplankton is usually theprimary producer (the first level in the food chain or the firsttrophic level). Phytoplankton converts inorganic carbon intoprotoplasm. Phytoplankton is consumed by microscopic animals calledzooplankton. These are the second level in the food chain, including krill, the larva of fish, squid, lobsters and crabs –as well as the smallcrustaceans calledcopepods, and many other types. Zooplankton is consumed by other, larger predatory zooplankton, fish, and baleen whales. Top ocean predators such as sharks, large seals, and dolphins then eat the fish or other organisms that eat zooplankton.[5] Trophic levels differ among food webs around the world, for example, whales may consume zooplankton directly - leading to an environment with one less trophic level compared to an environment where the predator does not eat the zooplankton directly. External image Animation of global primary production[6]

Primary biomass
Global primary production can be estimated fromsatellite observations. Satellites scan thenormalised difference vegetation index (NDVI) over terrestrial habitats, and scan sea-surfacechlorophyll levels over oceans. This results in 56.4 billiontonnesC/yr (53.8%), for terrestrial primary production, and 48.5 billion tonnes C/yr for oceanic primary production.[8] Thus, the totalphotoautotrophic primary production for the Earth is about 104.9 billion tonnes C/yr. This translates to about 426 gC/m2/yr for land production (excluding areas with permanent ice cover), and 140 gC/m2/yr for the oceans. However, there is a much more significant difference instanding stocks - while accounting for almost half of total annual production, oceanicautotrophs account for only about 0.2% of the total biomass. The most successfulanimal species, in terms of biomass, is probably theAntarctic krill,Euphausia superba, with a biomass of about 500 milliontonnes.[7][9] However, as a group, the small aquaticcrustaceans calledcopepods form the largest animal biomass on earth.[10]
BiomeEcosystem Type	Area	Mean Net Primary Production	World Primary Production	Mean biomass	World biomass	Minimum replacement rate
	(million km2)	(gram dryC / m2 / year)	(billion tonnes / year)	(kg dryC / m2)	(billion tonnes)	(years)
Open ocean	332.00	125.00	41.50	0.003	1.00	0.02
Upwelling zones	0.40	500.00	0.20	0.02	0.01	0.04
Continental shelf	26.60	360.00	9.58	0.01	0.27	0.03
Algal beds andreefs	0.60	2,500.00	1.50	2.00	1.20	0.80
Estuaries &mangroves	1.40	1,500.00	2.10	1.00	1.40	0.67
Total marine	361.00	152.01	54.88	0.01	3.87	0.07
Lakes andstreams	2.00	250.00	0.50	0.02	0.04	0.08
Terrestrial	147.00	554.51	114.90	12.55	1,873.38	16.15
Grand total	510.00	333.87	170.28	3.68	1,877.29	11.02
Source:Whittaker, R. H.;Likens, G. E. (1975)."The Biosphere and Man". In Leith, H; Whittaker, R H (eds.).Primary Productivity of the Biosphere. Springer-Verlag. pp. 305–328.ISBN 978-0-387-07083-4.; Ecological Studies Vol 14 (Berlin) Darci and Taylre are biomass specialists.

External image Conservation status map of the global 200 Aquatic habitats have been classified intomarine andfreshwaterecoregions by theWorldwide Fund for Nature (WWF). An ecoregion is defined as a "relatively large unit of land or water containing a characteristic set of natural communities that share a large majority of their species, dynamics, and environmental conditions (Dinerstein et al. 1995, TNC 1997).[11]

Estuaries are semi-enclosedcoastal bodies ofwater with one or morerivers orstreams flowing into them, and with a free connection to the opensea.[12] Estuaries are often associated with high rates of biological productivity. They are small, in demand, impacted by events far upstream or out at sea, and concentrate materials such as pollutants and sediments.[13][14] Lagoons are bodies of comparatively shallowsalt orbrackish water separated from the deepersea by a shallow or exposedsandbank,coral reef, or similar feature.Lagoon refers to both coastal lagoons formed by the build-up of sandbanks or reefs along shallow coastal waters, and the lagoons in atolls, formed by the growth of coral reefs on slowly sinking central islands. Lagoons that are fed by freshwater streams are estuaries. Theintertidal zone (foreshore) is the area that is exposed to theair atlow tide and submerged athigh tide, for example, the area between tide marks. This area can include many different types of habitats, including steep rocky cliffs, sandy beaches or vastmudflats. The area can be a narrow strip, as in Pacific islands that have only a narrow tidal range, or can include many meters of shoreline where shallow beach slope interacts with high tidal excursion. Thelittoral zone is the part of the ocean closest to the shore. The wordlittoral comes from theLatinlitoralis, which meansseashore.[15] The littoral zone extends from the high-water mark to near shore areas that are permanently submerged, and includes the intertidal zone. Definitions vary. Encyclopædia Britannica defines the littoral zone in a thoroughly vague way as the "marine ecological realm that experiences the effects of tidal and longshore currents and breaking waves to a depth of 5 to 10 metres (16 to 33 ft) below the low-tide level, depending on the intensity of storm waves".[16] The US Navy defines it as extending "from the shoreline to 600 feet (180 meters) out into the water"[17] Thesublittoral zone is the part of the ocean extending from the seaward edge of the littoral zone to the edge of thecontinental shelf.[18] It is sometimes called theneritic zone. Websters defines the neritic zone as the region of shallow water adjoining the seacoast. The wordneritic perhaps comes from theNeo-Latinnerita, which refers to a genus of marine snails, 1891.[19] The sublittoral zone is relatively shallow, extending to about 200 metres (110 fathoms), and generally has well-oxygenated water, low water pressure, and relatively stable temperature andsalinity levels. These, combined with presence of light and the resulting photosynthetic life, such asphytoplankton and floatingsargassum,[20] make the sublittoral zone the location of the majority of sea life. Voigt, Brian (1998)Glossary of Coastal TerminologyArchived 2008-09-16 at theWayback Machine Washington State Department of Ecology, publication 98–105 Pawson, M G; Pickett, G D and Walker, P (2002)The coastal fisheries of England and Wales, Part IV: A review of their status 1999–2001 Science Series, Technical Report 116.

Continental shelves are the extended perimeters of eachcontinent and associatedcoastal plain, which is covered duringinterglacial periods such as the current epoch by relatively shallowseas (known asshelf seas) and gulfs. The shelf usually ends at a point of decreasing slope (called theshelf break). The sea floor below the break is thecontinental slope. Below the slope is thecontinental rise, which finally merges into the deep ocean floor, theabyssal plain. The continental shelf and the slope are part of thecontinental margin. Continental shelves are shallow (averaging 140 metres or 460 feet), and the sunlight available means they can teem with life. The shallowest parts of the continental shelf are calledfishing banks.[21] There the sunlight penetrates to the seafloor and theplankton, on which fish feed, thrive.

Coral reefs arearagonite structures produced by living organisms, found in shallow, tropical marine waters with little to no nutrients in the water. High nutrient levels such as those found in runoff from agricultural areas can harm the reef by encouraging the growth ofalgae.[25] Although corals are found both in temperate and tropical waters, reefs are formed only in a zone extending at most from 30°N to 30°S of the equator.

Coral reefs: Details

Coral reefs are estimated to cover 284,300 square kilometres, with theIndo-Pacific region (including theRed Sea,Indian Ocean,Southeast Asia and thePacific) accounting for 91.9% of the total.[citation needed] Southeast Asia accounts for 32.3% of that figure, while the Pacific includingAustralia accounts for 40.8%.Atlantic andCaribbean coral reefs only account for 7.6% of the world total.[26] Coral reefs are either restricted or absent from the west coast of theAmericas, as well as the west coast ofAfrica. This is due primarily toupwelling and strong cold coastal currents that reduce water temperatures in these areas.[27] Corals are also restricted from off the coastline ofSouth Asia fromPakistan toBangladesh.[26] They are also restricted along the coast around northeasternSouth America andBangladesh due to the release of vast quantities of freshwater from theAmazon andGanges Rivers respectively.[citation needed] Famous coral reefs and reef areas of the world include: TheGreat Barrier Reef - largest coral reef system in the world,Queensland, Australia; TheBelize Barrier Reef - second largest in the world, stretching from southern Quintana Roo, Mexico and all along the coast of Belize down to the Bay Islands of Honduras. TheRed Sea Coral Reef - located off the coast of Egypt and Saudi Arabia. Pulley Ridge - deepest photosynthetic coral reef,Florida Many of the numerous reefs found scattered over theMaldives TheNew Caledonia Barrier Reef - second longest double barrier reef in the world, with a length of about 1,500 km (932 mi). Coral reefs support an extraordinarybiodiversity; although they are located in nutrient-poor tropical waters. The process ofnutrient cycling between corals, zooxanthellae, and other reef organisms provides an explanation for why coral reefs flourish in these waters: recycling ensures that fewer nutrients are needed overall to support the community. Coral reefs are home to a variety of tropical or reeffish, such as the colorfulparrotfish,angelfish,damselfish, andbutterflyfish. Other fish groups found on coral reefs includegroupers,snappers,grunts andwrasses. Over 4,000 species of fish inhabit coral reefs.[26] It has been suggested that the high number of fish species that inhabit coral reefs are able to coexist in such high numbers because any free living space is rapidly inhabited by the first planktonic fish larvae that occupy it. These fish then inhabit the space for the rest of their life. The species that inhabit the free space is random and has therefore been termed 'a lottery for living space'.[28] Reefs are also home to a large variety of other organisms, includingsponges,Cnidarians (which includes some types of corals andjellyfish),worms,crustaceans (includingshrimp,spiny lobsters andcrabs),molluscs (includingcephalopods),echinoderms (includingstarfish, sea urchins andsea cucumbers),sea squirts,sea turtles andsea snakes.[26][29] Human activity may represent the greatest threat to coral reefs living in Earth'soceans. In particular,pollution and over-fishing are the most serious threats to these ecosystems. Physical destruction of reefs due to boat and shipping traffic is also a problem. Thelive food fish trade has been implicated as a driver of decline due to the use ofcyanide anddisaster for peoples living in the tropics. Hughes, et al., (2003), writes that "with increasedhuman population and improved storage and transport systems, the scale of human impacts on reefs has grown exponentially. For example, markets for fishes and othernatural resources have become global, supplying demand for reef resources far removed from their tropical sources."[31] Currently researchers are working to determine the degree various factors impact the reef systems. The list of factors is long but includes the oceans acting as acarbon dioxide sink, changes inEarth's atmosphere,ultraviolet light,ocean acidification, biologicalvirus, impacts ofdust storms carrying agents to far flung reef systems, various pollutants, impacts ofalgal blooms and others. Reefs are threatened well beyond coastal areas and so the problem is broader than factors from land development and pollution though those are too causing considerable damage. Southeast Asian coral reefs are at risk from damagingfishing practices (such ascyanide andblast fishing),overfishing, sedimentation, pollution and bleaching. A variety of activities, including education, regulation, and the establishment of marine protected areas are under way to protect these reefs.Indonesia, for example has nearly 33,000 square miles (85,000 km2) of coral reefs. Its waters are home to a third of the world's total corals and a quarter of its fish species. Indonesia's coral reefs are located in the heart of theCoral Triangle and have been victim to destructive fishing, unregulated tourism, and bleaching due to climatic changes. Data from 414 reef monitoring stations throughout Indonesia in 2000 found that only 6% of Indonesia's coral reefs are in excellent condition, while 24% are in good condition, and approximately 70% are in poor to fair condition (2003The Johns Hopkins University). General estimates show approximately 10% of the coral reefs around the world are already dead.[32][33] Problems range fromenvironmental effects of fishing techniques, described above, toocean acidification.[34]Coral bleaching is another manifestation of the problem and is showing up in reefs across the planet. Inhabitants of Ahus Island,Manus Province,Papua New Guinea, have followed a generations-old practice of restricting fishing in six areas of their reef lagoon. While line fishing is permitted, net and spear fishing are restricted based on cultural traditions. The result is that both thebiomass and individual fish sizes are significantly larger in these areas than in places where fishing is completely unrestricted.[35][36]It is estimated that about 60% of the world's reefs are at risk due to destructive, human-related activities. The threat to the health of reefs is particularly strong inSoutheast Asia, where an enormous 80% of reefs are consideredendangered. Organisations asCoral Cay,Counterpart[37] and theFoundation of the peoples of the South PacificArchived 2008-08-20 at theWayback Machine are currently undertaking coral reef/atoll restoration projects. They are doing so using simple methods ofplant propagation. Other organisations asPractical Action have released informational documents on how to set up coral reef restoration to the public.[38]

Aseamount is an underwatermountain, rising from theseafloor that does not reach to the water's surface (sea level), and thus is not anisland. They are defined byoceanographers as independent features that rise to at least 1,000 meters above the seafloor. Seamounts are common in the Pacific Ocean. Recent studies suggest there may be 30,000 seamounts in the Pacific, about 1,000 in the Atlantic Ocean and an unknown number in the Indian Ocean.[39]

Worldwide, freshwater lakes have an area of 1.5 million square kilometres.^[44] Saline inland seas add another 1.0 million square kilometres.^[45] There are 28 freshwater lakes with an area greater than 5,000 square kilometres, totalling 1.18 million square kilometres or 79 percent of the total.^[46]

Freshwater fisheries are essential to supporting human life around the globe whether they are used for recreation or commercial use. Climate change presents several challenges in sustaining these fisheries as waters become warmer resulting in decreased dissolved oxygen, as the toxicity of pollutants increases, and as the physiological changes in fishes and changes in their habitat systems alter what we are used to. Deoxygenation and eutrophication are two major effects that are detrimental to fish and ecosystem health and the problem is more prevalent as the size of the body of water decreases.^[47] Details on the changes occurring in fish physiology and their habitats can be found at the respective citation.

Increased management and surveillance on freshwater fisheries will be vital to the longevity, sustainability, and productivity of the fisheries and essential to maintaining our food production from that source.

Pollution is the introduction of contaminants into an environment. Wild fisheries flourish in oceans, lakes, and rivers, and the introduction of contaminants is an issue of concern, especially as regards plastics, pesticides, heavy metals, and other industrial and agricultural pollutants which do not disintegrate rapidly in the environment. Land run-off and industrial, agricultural, and domestic waste enter rivers and are discharged into the sea.Pollution from ships is also a problem.

Marine debris is human-created waste that ends up floating in the sea. Oceanic debris tends to accumulate at the centre of gyres and coastlines, frequently washing aground where it is known as beach litter. Eighty percent of all known marine debris is plastic - a component that has been rapidly accumulating since the end of World War II.^[48] Plastics accumulate because they don'tbiodegrade as many other substances do; while they willphotodegrade on exposure to the sun, they do so only under dry conditions, aswater inhibits this process.^[49]

Discardedplastic bags,six-pack rings and other forms ofplastic waste which finish up in the ocean present dangers to wildlife and fisheries.^[50] Aquatic life can be threatened through entanglement, suffocation, and ingestion.^[51][52][53]

Nurdles, also known as mermaids' tears, are plastic pellets typically under five millimetres in diameter, and are a major contributor to marine debris. They are used as a raw material in plastics manufacturing, and are thought to enter thenatural environment after accidental spillages. Nurdles are also created through thephysical weathering of larger plastic debris. They strongly resemblefish eggs, only instead of finding a nutritious meal, any marine wildlife that ingests them will likely starve, be poisoned and die.^[54]

Many animals that live on or in the seaconsume flotsam by mistake, as it often looks similar to their natural prey.^[55] Plastic debris, when bulky or tangled, is difficult to pass, and may become permanently lodged in the digestive tracts of these animals, blocking the passage of food and causing death through starvation or infection.^[56] Tiny floating particles also resemblezooplankton, which can leadfilter feeders to consume them and cause them to enter the oceanfood chain. In samples taken from theNorth Pacific Gyre in 1999 by the Algalita Marine Research Foundation, the mass of plastic exceeded that of zooplankton by a factor of six.^[48][57] More recently, reports have surfaced that there may now be 30 times more plastic than plankton, the most abundant form of life in the ocean.^[58]

Toxicadditives used in the manufacture of plastic materials canleach out into their surroundings when exposed to water. Waterbornehydrophobic pollutantscollect and magnify on the surface of plastic debris,^[59] thus making plastic far more deadly in the ocean than it would be on land.^[48] Hydrophobic contaminants are also known tobioaccumulate in fatty tissues,biomagnifying up the food chain and putting great pressure onapex predators. Some plastic additives are known to disrupt theendocrine system when consumed, others can suppress the immune system or decrease reproductive rates.^[57]

Apart from plastics, there are particular problems with other toxins which do not disintegrate rapidly in the marine environment. Heavy metals are metallic chemical elements that have a relatively high density and are toxic or poisonous at low concentrations. Examples aremercury,lead,nickel,arsenic andcadmium. Other persistent toxins arePCBs,DDT,pesticides,furans,dioxins andphenols.

Such toxins can accumulate in the tissues of many species of aquatic life in a process calledbioaccumulation. They are also known to accumulate inbenthic environments, such asestuaries andbay muds: a geological record of human activities of the last century.

Some specific examples are

• Chinese and Russian industrial pollution such asphenols and heavy metals in theAmur River have devastated fish stocks and damaged itsestuary soil.^[60]
• Wabamun Lake inAlberta,Canada, once the bestwhitefish lake in the area, now has unacceptable levels of heavy metals in its sediment and fish.
• Acute and chronicpollution events have been shown to impact southern Californiakelp forests, though the intensity of the impact seems to depend on both the nature of the contaminants and duration of exposure.^{[61][62][63][64][65]}
• Due to their high position in thefood chain and the subsequentaccumulation ofheavy metals from their diet,mercury levels can be high in larger species such as bluefin andalbacore. As a result, in March 2004 theUnited StatesFDA issued guidelines recommending that pregnant women, nursing mothers and children limit their intake of tuna and other types of predatory fish.^[66]
• Some shellfish and crabs can survive polluted environments, accumulating heavy metals or toxins in their tissues. For example,mitten crabs have a remarkable ability to survive in highly modifiedaquatic habitats, including polluted waters.^[67] The farming and harvesting of such species needs careful management if they are to be used as a food.^[68][69]

• Mining has a poor environmental track record. For example, according to theUnited States Environmental Protection Agency, mining has contaminated portions of the headwaters of over 40% of watersheds in the western continental US.^[70] Much of this pollution finishes up in the sea.

• Heavy metals enter the environment throughoil spills - such as thePrestige oil spill on theGalician coast - or from other natural oranthropogenic sources.^[71]

Eutrophication is an increase in chemicalnutrients, typically compounds containingnitrogen orphosphorus, in anecosystem. It can result in an increase in the ecosystem'sprimary productivity (excessive plant growth and decay), and further effects including lack of oxygen and severe reductions in water quality, fish, and other animal populations.

The biggest culprit are rivers that empty into the ocean, and with it the many chemicals used asfertilizers in agriculture as well as waste fromlivestock andhumans. An excess of oxygen depleting chemicals in the water can lead tohypoxia and the creation of adead zone.^[72]

Surveys have shown that 54% of lakes inAsia areeutrophic; inEurope, 53%; inNorth America, 48%; inSouth America, 41%; and inAfrica, 28%.^[73]Estuaries also tend to be naturally eutrophic because land-derived nutrients are concentrated where run-off enters the marine environment in a confined channel. TheWorld Resources Institute has identified 375 hypoxic coastal zones around the world, concentrated in coastal areas in Western Europe, the Eastern and Southern coasts of the US, and East Asia, particularly in Japan.^[74] In the ocean, there are frequentred tide algae blooms^[75] that kill fish and marine mammals and cause respiratory problems in humans and some domestic animals when the blooms reach close to shore.

In addition toland runoff, atmosphericanthropogenicfixed nitrogen can enter the open ocean. A study in 2008 found that this could account for around one third of the ocean's external (non-recycled) nitrogen supply and up to three per cent of the annual new marine biological production.^[76] It has been suggested that accumulating reactive nitrogen in the environment may have consequences as serious as putting carbon dioxide in the atmosphere.^[77]

The oceans are normally a naturalcarbon sink, absorbing carbon dioxide from the atmosphere. Because the levels of atmospheric carbon dioxide are increasing, the oceans are becomingmore acidic.^[78][79]The potential consequences of ocean acidification are not fully understood, but there are concerns that structures made of calcium carbonate may become vulnerable to dissolution, affecting corals and the ability of shellfish to form shells.^[80]

A report fromNOAA scientists published in the journal Science in May 2008 found that large amounts of relatively acidified water are upwelling to within four miles of the Pacificcontinental shelf area of North America. This area is a critical zone where most local marine life lives or is born. While the paper dealt only with areas fromVancouver to northern California, other continental shelf areas may be experiencing similar effects.^[81]

Fishing nets that have been left or lost in the ocean by fishermen are calledghost nets, and can entanglefish,dolphins,sea turtles,sharks,dugongs,crocodiles,seabirds,crabs, and other creatures. Acting as designed, these nets restrict movement, causing starvation, laceration and infection, and—in those that need to return to the surface to breathe—suffocation.^[82]

Fishing operations often use trawl netting dragging and dredging them across the ocean bottom. Numerous habitats and ecosystems are disturbed and destroyed by trawling including coral reefs, sediments, and grasses that provide feeding and breeding grounds for a plethora of marine organisms. Coastal habitats such as mangroves are often sites of aquaculture farming practices in which the mangroves are either destroyed for easier use of the land or experience harmful conditions due to the farm being abandoned once the area becomes too polluted with excess nutrients.^[83]

Some specific examples of overfishing.

• On the east coast of theUnited States, the availability of bay scallops has been greatly diminished by the overfishing of sharks in the area. A variety of sharks have, until recently, fed onrays, which are a main predator of bay scallops. With the shark population reduced, in some places almost totally, the rays have been free to dine on scallops to the point of greatly decreasing their numbers.^{[citation needed]}
• Chesapeake Bay's once-flourishingoyster populations historically filtered theestuary's entire water volume of excess nutrients every three or four days. Today that process takes almost a year,^[84] and sediment, nutrients, and algae can cause problems in local waters. Oysters filter these pollutants, and either eat them or shape them into small packets that are deposited on the bottom where they are harmless.
• The Australian government alleged in 2006 thatJapan illegally overfishedsouthern bluefin tuna by taking 12,000 to 20,000 tonnes per year instead of their agreed 6,000 tonnes; the value of such overfishing would be as much as US$2 billion. Such overfishing has resulted in severe damage to stocks. "Japan's huge appetite for tuna will take the most sought-after stocks to the brink of commercial extinction unless fisheries agree on more rigid quotas" stated theWWF.^[85][86] Japan disputes this figure, but acknowledges that some overfishing has occurred in the past.^[87]
• Jackson, Jeremy B C et al. (2001)Historical overfishing and the recent collapse of coastal ecosystems Science 293:629-638.

Our World in Data provides a figure showing the trend in global fishing exploitation over a few decades to reveal the intensifying circumstances at hand:

Overfishing presents many threats to fish population densities, obviously. However, as these populations plummet below the maximum sustainable yield (MSY) value for the specific population, you are now risking the loss of biodiversity and possibility for extinction due to less diversity. This loss in diversity is especially concerning as we deal with environmental changes from climate change since less diversity decreases a populations ability to adapt and survive the alterations of the habitat.

• 
  This graph demonstrates the importance of following a specific quota of exploitation to sustain a resource such as a fish population.

Eachspecies in anecosystem is affected by the other species in that ecosystem. There are very few single prey-single predator relationships. Most prey are consumed by more than one predator, and most predators have more than one prey. Their relationships are also influenced by other environmental factors. In most cases, if one species is removed from an ecosystem, other species will most likely be affected, up to the point of extinction.

Speciesbiodiversity is a major contributor to the stability of ecosystems. When an organism exploits a wide range of resources, a decrease in biodiversity is less likely to have an impact. However, for an organism which exploit only limited resources, a decrease in biodiversity is more likely to have a strong effect.

Reduction of habitat, hunting and fishing of some species toextinction or near extinction, and pollution tend to tip the balance ofbiodiversity. For a systematic treatment of biodiversity within atrophic level, seeunified neutral theory of biodiversity.

The global standard for recordingthreatened marine species is theIUCN Red List of Threatened Species.^[88] This list is the foundation for marine conservation priorities worldwide. A species is listed in the threatened category if it is considered to becritically endangered,endangered, orvulnerable. Other categories arenear threatened anddata deficient.

Many marine species are under increasing risk of extinction and marinebiodiversity is undergoing potentially irreversible loss due to threats such asoverfishing,bycatch,climate change,invasive species and coastal development.

By 2008, theIUCN had assessed about 3,000 marine species. This includes assessments of known species of shark, ray, chimaera, reef-building coral, grouper, marine turtle, seabird, and marine mammal. Almost one-quarter (22%) of these groups have been listed as threatened.^[89]

• Sharks, rays, and chimaeras: are deep waterpelagic species, which makes them difficult to study in the wild. Not a lot is known about their ecology and population status. Much of what is currently known is from their capture innets from both targeted and accidental catch. Many of these slow growing species are not recovering from overfishing by shark fisheries around the world.
• Groupers: Major threats are overfishing, particularly the uncontrolled fishing of small juveniles and spawning adults.
• Coral reefs: The primary threats tocorals are bleaching and disease which has been linked to an increase in sea temperatures. Other threats include coastal development, coral extraction, sedimentation and pollution. Thecoral triangle (Indo-Malay-Philippine archipelago) region has the highest number of reef-building coral species in threatened category as well as the highest coral species diversity. The loss of coral reef ecosystems will have devastating effects on many marine species, as well as on people that depend on reef resources for their livelihoods.
• Marine mammals: includewhales,dolphins,porpoises,seals,sea lions,walruses,sea otter,marine otter,manatees,dugong and thepolar bear. Major threats include entanglement inghost nets, targeted harvesting, noise pollution from military and seismic sonar, and boat strikes. Other threats are water pollution, habitat loss from coastal development, loss of food sources due to the collapse of fisheries, and climate change.
• Seabirds: Major threats includelongline fisheries andgillnets,oil spills, and predation by rodents and cats in their breeding grounds. Other threats are habitat loss and degradation from coastal development, logging and pollution.
• Marine turtles: Marine turtles lay their eggs on beaches, and are subject to threats such as coastal development, sand mining, and predators, including humans who collect their eggs for food in many parts of the world. At sea, marine turtles can be targeted by small scalesubsistence fisheries, or becomebycatch duringlongline andtrawling activities, or become entangled inghost nets or struck by boats.

An ambitious project, called the Global Marine Species Assessment, is under way to make IUCN Red List assessments for another 17,000 marine species by 2012. Groups targeted include the approximately 15,000 known marine fishes, and important habitat-formingprimary producers suchmangroves,seagrasses, certainseaweeds and the remainingcorals; and important invertebrate groups includingmolluscs andechinoderms.^[89]

Freshwater fisheries have a disproportionately high diversity of species compared to other ecosystems. Although freshwater habitats cover less than 1% of the world's surface, they provide a home for over 25% of known vertebrates, more than 126,000 known animal species, about 24,800 species offreshwater fish,molluscs,crabs anddragonflies, and about 2,600macrophytes.^[89]Continuing industrial and agricultural developments place huge strain on these freshwater systems. Waters are polluted or extracted at high levels, wetlands are drained, rivers channelled, forests deforestated leading to sedimentation, invasive species are introduced, and over-harvesting occurs.

In the 2008IUCN Red List, about 6,000 or 22% of the known freshwater species have been assessed at a global scale, leaving about 21,000 species still to be assessed. This makes clear that, worldwide, freshwater species are highly threatened, possibly more so than species in marine fisheries.^[90] However, a significant proportion of freshwater species are listed asdata deficient, and more field surveys are needed.^[89]

A recent paper published by theNational Academy of Sciences of the USA warns that: "Synergistic effects ofhabitat destruction, overfishing, introduced species, warming, acidification, toxins, and massive runoff of nutrients are transforming once complex ecosystems like coral reefs and kelp forests into monotonous level bottoms, transforming clear and productive coastal seas into anoxic dead zones, and transforming complex food webs topped by big animals into simplified, microbially dominated ecosystems with boom and bust cycles of toxic dinoflagellate blooms, jellyfish, and disease".^[91]

• Fishing by country
• List of countries by seafood consumption
• List of harvested aquatic animals by weight
• Ocean fisheries
• Population dynamics of fisheries
• World fish production

• International Nitrogen Initiative:Web site
• Population Distribution within 100 km of Coastlines (2000)World Resources Institute.
• NOAA:Carbon cycle science

• Fishing industry

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