Aprostrate shrub is awoody plant, most of the branches of which lie upon or just above the ground, rather than being held erect as are the branches of most trees and shrubs.
Prostration may occur because the supporting tissues in stems are not strong enough to support the weight of the plant, causing the plant to bend until it reaches the ground. Alternatively, it may occur because of a genetic disposition for branches to grow horizontally on or just under the ground; for example, as a strategy to avoid overly strong sunlight. Finally, environmental factors such as strong winds laden with sand or salt may tend to prune away erect branches, thereby creating a prostrate habit in plants that may not be predisposed to prostration.
Prostrate shrubs are used inhorticulture asgroundcovers and inhanging baskets, and to bind soils and preventerosion in remediallandscaping. They are also important components ofrock gardens. The shrinking size ofurban gardens has meant an increase in demand for and desirability of dwarf and prostrate forms of many garden plants. More recently, prostrate shrubs have received attention for their usefulness in plantinggreen roofs andgreen walls, where they can contribute toenvironmental conservation.
Prostrate plant forms may arise from deliberate breeding and hybridisation, such as the groundcovergrevilleas, includingGrevillea "Poorinda Royal Mantle",[1] andG. "Bronze Rambler",[2] or by selection of forms of plants growing in exposed areas. The windswept coastal heathlands and cliffs of far southeastern New South Wales have yieldedBanksia 'Roller Coaster' andBanksia 'Pygmy Possum'.[3] These are flattened varieties of less than 0.5 m in height of the speciesBanksia integrifolia andBanksia serrata, respectively, both of which grow to sizeable trees of 10 m (30 ft) or more in height.[3] The origin of others is unclear; Cootamundra Wattle (Acacia baileyana) is a popular and widely used garden plant in Australia, a prostrate form of which is also in cultivation. Its origin is unknown, possibly a chance seedling in cultivation. It itself is a popular garden plant, its cascading horizontal branches good for rockeries.[4]
Manyspecies of roses grow as long canes that can spread prostrate on the ground, and some of these have been hybridized to formclimbing and rambling roses that can be allowed to grow on the ground without support, as well as varieties known as "carpet roses" that are intended to be grown as ground covers.[5] Prostrate juniper,Juniperus communis subsp.alpina is a naturally occurring form of a popularevergreen.[6] Many species ofCotoneaster, such asC. apiculatus, are prostrate ornamentals,[7] as are the closely relatedPyracanthas, and thehumilis variant of sweetbox,Sarcococca hookeriana var.humilis. Heaths (Erica carnea and related species) and heathers (Calluna vulgaris) are prostrate shrubs often featured prominently in rock gardens.[8] Wintergreens such asGaultheria humifusa andGaultheria procumbens are also prostrate growers.[9]
Although, strictly speaking, herbaceous, the North American bunchberry (Cornus canadensis), a member of thedogwood genus, can appear as a prostrate shrub when growing in the garden.[10] Manydaphnes, particularly those derived fromDaphne odora, are prostrate growers, including forms that are highly fragrant and have variegated leaves, although they are short-lived.[11] Some forms of the culinaryherbrosemary (Rosmarinus officinalis) form prostrate shrubs when grown inMediterranean climates, although they die back to the ground when grown where winters are colder.[12]
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