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Fir

From Wikipedia, the free encyclopedia
(Redirected fromAbies)
Genus of conifers
For other uses, seeFIR (disambiguation) andFIRS (disambiguation).
"Fir tree" redirects here. For other uses, seeFir Tree (disambiguation).

Fir
Temporal range:49–0 MaEocene -Present[1] Possible Campanian record[2]
Korean fir (Abies koreana) cones and foliage
Scientific classificationEdit this classification
Kingdom:Plantae
Clade:Tracheophytes
Clade:Gymnospermae
Division:Pinophyta
Class:Pinopsida
Order:Pinales
Family:Pinaceae
Subfamily:Abietoideae
Genus:Abies
Mill.
Type species
Abies alba
Species

See text

Synonyms
  • PeuceRichard 1810

Firs areevergreenconiferous trees belonging to thegenusAbies (Latin:[ˈabieːs]) in the familyPinaceae. There are approximately 48–65[3][4] extant species, found on mountains throughout much of North andCentral America, Eurasia, andNorth Africa. The genus is most closely related toKeteleeria, a small genus confined to eastern Asia.[5]

The genus name is derived from the Latin "to rise" in reference to the height of its species.[6] The common English name originates with the Old Norsefyri or the Old Danishfyr.[7]

They are large trees, reaching heights of 10–80 metres (33–262 feet) tall with trunk diameters of 0.5–4 m (1 ft 8 in – 13 ft 1 in) when mature. Firs can be distinguished from other members of the pine family by the way in which their needle-likeleaves are attached singly to the branches with a base resembling asuction cup, and by theircones, which, like those ofcedars, stand upright on the branches like candles and disintegrate at maturity.

Identification of the different species is based on the size and arrangement of the leaves, the size and shape of the cones, and whether thebract scales of the cones are long and exserted, or short and hidden inside the cone.

Description

[edit]

Leaves

[edit]
  • A. alba – the underside of leaves have two whitish strips formed by wax-covered stomatal bands, while their bases are shaped like suction cups.
    A. alba – the underside of leaves have two whitish strips formed bywax-coveredstomatal bands, while their bases are shaped like suction cups.
  • A. grandis foliage – upper side of the leaves, showing the leaves lying flat either side of the shoot
    A. grandis foliage – upper side of the leaves, showing the leaves lying flat either side of the shoot
  • Foliage of A. pinsapo showing the radial leaf arrangement in this species
    Foliage ofA. pinsapo showing the radial leaf arrangement in this species
  • Most firs are inbetween the two extremes of flat and radial, with an intermediate arrangement, often with longer leaves at the sides, and shorter leaves above the shoot; here, A. mariesii in Japan
    Most firs are inbetween the two extremes of flat and radial, with an intermediate arrangement, often with longer leaves at the sides, and shorter leaves above the shoot; here,A. mariesii in Japan

Firs can be distinguished from other members of the pine family by the unique attachment of their needle-likeleaves to the twig by a base that resembles a smallsuction cup. The leaves are significantly flattened, sometimes even looking like they are pressed, as inA. sibirica.

The leaves have two whitish lines on the bottom, each of which is formed bywax-coveredstomatal bands. In most species, the upper surface of the leaves is uniformly green and shiny, withoutstomata or with a few on the tip, visible as whitish spots. Other species have the upper surface of leaves dull, greyish green or bluish to silvery (glaucous), coated by wax with variable number ofstomatal bands, and not always continuous. An example species with shiny green leaves isA. alba, and an example species with matt waxy leaves isA. concolor.

The tips of leaves are usually more or less notched (as inA. firma), but sometimes rounded or dull (as inA. concolor,A. magnifica) or sharp and prickly (as inA. bracteata,A. cephalonica,A. holophylla). The leaves of young plants are usually sharper.

The leaves are arranged spirally on the shoots, but by being twisted at their base, the way they spread from the shoot is diverse; in some species comb-like ('pectinate'), with the leaves flat on either side of the shoot (e.g.A. alba,A. grandis), in others, the leaves remain radial (e.g.A. pinsapo)[8]

Foliage in the upper crown on cone-bearing branches is different, with the leaves shorter, curved, and sometimes sharp.[9]

Cones

[edit]
  • Intact and disintegrated Bulgarian fir cones
    Intact and disintegratedBulgarian fir cones
  • Immature cones of some species are green; here Manchurian fir Abies holophylla
    Immature cones of some species are green; hereManchurian firAbies holophylla
  • Disintegrating cones of Manchurian fir
    Disintegrating cones of Manchurian fir
  • Pindrow fir Abies pindrow with dark purple cones
    Pindrow firAbies pindrow with dark purple cones
  • Caucasian Fir Abies nordmanniana young cones with reddish scales and yellow-green bracts
    Caucasian FirAbies nordmanniana young cones with reddish scales and yellow-green bracts
  • Noble fir Abies procera, with five heavy (20 cm, approx 0.5 kg each) cones, three erect, and two hanging where their weight has caused the branch to twist
    Noble firAbies procera, with five heavy (20 cm, approx 0.5 kg each) cones, three erect, and two hanging where their weight has caused the branch to twist

Firs differ from other conifers in having erect, cylindricalcones 5–25 cm (2–10 in) long that disintegrate at maturity to release the wingedseeds. In contrast tospruces, fir cones are erect; they do not hang, unless heavy enough to twist the branch with their weight.

The mature cones are usually brown. When young in summer, they can be green:

A. grandis,A. holophylla

or reddish:

A. alba,A. cephalonica,A. nordmanniana

or bloomed pale glaucous or pinkish:

A. numidica,A. pinsapo

or purple to blue, sometimes very dark blue, almost black:

A. forrestii,A. fraseri,A. homolepis,A. lasiocarpa,A. pindrow.

Many species arepolymorphic in cone colour, with different individuals of the same species producing either green or purple cones:

A. concolor,A. koreana (usually purple, rarely green, such as thecultivar 'Flava'),A. magnifica (usually green, occasionally purple),A. nephrolepis (f. chlorocarpa green),A. sibirica,A. veitchii (f. olivacea green)[8]

The cone scale bracts can be short and hidden in the mature cone, or long and exposed ('exserted'); this can vary even within a species, e.g. inAbies magnifica var.magnifica, the bracts are hidden, but in var.critchfieldii and var.shastensis, they are exserted. The bracts scales are often a different colour to the cone scales, which can make for a very attractive combination valued in ornamental trees.

Classification

[edit]

The oldest pollen assignable to the genus dates to the Late Cretaceous in Siberia, with records of leaves and reproductive organs across the Northern Hemisphere from theEocene onwards.[10]

Phylogeny ofAbies[11][12]
(Pseudotorreya)

A. bracteata(Don) Poit.

(Religio)
section

A. mariesiiMasters

A. amabilis(Douglas ex Loudon) Forbes

A. proceraRehder

Amabilis
section

A. magnificaMurray

Nobiles
section

A. concolor(Gordon) Lindley ex Hildebr.

A. jaliscana(Martínez) Mantilla, Shalisko & Vázquez

A. guatemalensisRehder

A. hickeliiFlous & Gaussen

A. flinckiiRushforth

A. vejariiMartínez

A. durangensisMartínez

A. religiosa(Kunth) von Schlechtendal & von Chamisso

A. hidalgensisDebreczy, Rácz & Guízar

A. grandis(Douglas ex Don) Lindley

A. lowiana(Gordon) Murray

Grandis
(Abies)
section

A. albaMiller

A. pinsapoBoiss.

A. cephalonicaLoudon

A. nebrodensis(Lojac.) Mattei

A. nordmanniana(Steven) Spach

A. numidicade Lannoy ex Carrière

A. ×borisii-regisMattf.

A. cilicica(Antoine & Kotschy) Carrière

Abies
section

A. lasiocarpa(Hooker) Nuttall

A. ernestiiRehder

A. balsamea(von Linné) Miller

Balsamea
section

A. firmaSiebold & Zuccarini

A. sibiricaLedeb.

Sibiria
section

A. fanjingshanensisHuang, Tu & Fang

A. ziyuanensisFu & Mo

series

A. kawakamii(Hayata) Itô

A. chensiensisvan Tieghem

A. squamataMasters

A. beshanzuensisWu

A. pindrow(Royle ex Don) Royle

Squamatae
series

A. koreanaWilson

A. nephrolepis(Trautvetter ex Maxim.) Maxim.

A. holophyllaMaxim.

A. sachalinensis(Schmidt) Masters

A. fabri(Masters) Craib

A. veitchiiLindley

A. fraseri(Pursh) Poiret

A. delavayiFranchet

A. densaGriffith

A. spectabilis(Don) de Mirbel

A. nukiangensisCheng & Fu

A. ferreanaBordères & Gaussen

A. forrestiiColtm.-Rog.

A. georgeiOrr

A. homolepisSiebold & Zuccarini

A. yuanbaoshanensisLu & Fu

Spectabiles
Pseudopicea

SectionAbies

[edit]

SectionAbies is found in central, south, and eastern Europe and Asia Minor.

SectionBalsamea

[edit]

SectionBalsamea is found in northern Asia and North America, and high mountains further south.

SectionGrandis

[edit]

SectionGrandis is found in western North America to Mexico, Guatemala, Honduras and El Salvador, in lowlands in the north, moderate altitudes in south.

  • Abies grandis – grand fir or giant fir
    • Abies grandis var.grandis – Coast grand fir
    • Abies grandis var.idahoensis – interior grand fir
  • Abies concolor – white fir
    • Abies concolor subsp.concolor – Rocky Mountain white fir or Colorado white fir
    • Abies concolor subsp.lowiana – Low's white fir or Sierra Nevada white fir
  • Abies durangensis – Durango fir
    • Abies durangensis var.coahuilensis – Coahuila fir
  • Abies flinckii – Jalisco fir
  • Abies guatemalensis – Guatemalan fir
    • Abies guatemalensis var.guatemalensis
    • Abies guatemalensis var.jaliscana
  • Abies vejarii

SectionMomi

[edit]

SectionMomi is found in east and central Asia and the Himalaya, generally at low to moderate altitudes.

SectionAmabilis

[edit]

SectionAmabilis is found in the Pacific Coast mountains in North America and Japan, in high rainfall areas.

SectionPseudopicea

[edit]
A. fabri, Sichuan, China

SectionPseudopicea is found in theSino – Himalayan mountains at high altitudes.

SectionOiamel

[edit]

SectionOiamel is found in central Mexico at high altitudes.

SectionNobilis

[edit]
A. magnifica, California, USA

SectionNobilis (westernU.S., high altitudes)

SectionBracteata

[edit]

SectionBracteata (California coast)

SectionIncertae sedis

[edit]

SectionIncertae sedis

Ecology

[edit]

Firs are used as food plants by thecaterpillars of someLepidoptera species, includingChionodes abella (recorded onwhite fir),autumnal moth,conifer swift (a pest ofbalsam fir),the engrailed,grey pug,mottled umber,pine beauty and thetortrix mothsCydia illutana (whose caterpillars are recorded to feed onEuropean silver fir cone scales) andC. duplicana (on European silver fir bark around injuries orcanker).

Abies religiosa (sacred fir) trees giveroosting shelter to overwinteringmonarch butterflies.[15][16]

Phytochemistry

[edit]

Abies produce a variety ofterpenoids. The analyses of the Zavarin group – from Smedman et al. 1969 to Zavarin et al. 1977 – showed variation in terpenoid composition of the bark by genetics, geography, age and size of the tree.[17][18]

Uses

[edit]
Green fir twig pictured on top of heart in the coat of arms ofLaukaa

Wood of most firs is considered unsuitable for generaltimber use and is often used aspulp or for the manufacture ofplywood and rough timber. It is commonly used inCanadian Lumber Standard graded wood.[19] Because this genus has no insect or decay resistance qualities after logging, it is generally recommended in construction purposes for indoor use only (e.g. indoordrywall onframing). Firwood left outside cannot be expected to last more than 12 to 18 months, depending on the type of climate it is exposed to.[citation needed]

Caucasian fir,noble fir,Fraser's fir andbalsam fir are popularChristmas trees, generally considered to be the best for this purpose, with aromatic foliage that does not shed many needles on drying out. Many are also decorative garden trees, notablyKorean fir and Fraser's fir, which produce brightly coloured cones even when very young, still only 1–2 m (3 ft 3 in – 6 ft 7 in) tall. Many fir species are grown in botanic gardens and other specialist tree collections in Europe and North America.[20]

Abies spectabilis or Talispatra is used inAyurveda as an antitussive (cough suppressant) drug.[21][22]

See also

[edit]

References

[edit]
  1. ^abSchorn, Howard; Wehr, Wesley (1986). "Abies milleri, sp. nov., from the Middle Eocene Klondike Mountain Formation, Republic, Ferry County, Washington".Burke Museum Contributions in Anthropology and Natural History.1:1–7.
  2. ^"Drumheller region (Cretaceous to of Canada)".PBDB.org.
  3. ^Torres, Leon Nahuel; Shi, Xiao; Na, Yuling; Wang, Bing; Tian, Chi; Chen, Jun (2024-03-01)."First study on fossil wood from the Middle Pleistocene of the Songliao Plain, Northeast China".Review of Palaeobotany and Palynology.322: 105063.Bibcode:2024RPaPa.32205063T.doi:10.1016/j.revpalbo.2024.105063.ISSN 0034-6667.
  4. ^Debreczy Zsolt Rácz István and Kathy Musial. 2011.Conifers Around the World : Conifers of the Temperate Zones and Adjacent Regions. Budapest: DendroPress.
  5. ^Leslie, Andrew B.; et al. (2018)."ajb21143-sup-0004-AppendixS4"(PDF).American Journal of Botany.105 (9):1531–1544.doi:10.1002/ajb2.1143.PMID 30157290.
  6. ^Coombes, Allen J. (2012).The A to Z of plant names : a quick reference guide to 4000 garden plants (1st ed.). Portland, Or.: Timber Press. pp. 17, 23.ISBN 978-1-60469-196-2.OCLC 741564356.
  7. ^"fir | Origin and meaning of fir by Online Etymology Dictionary".www.etymonline.com.Archived from the original on 2021-08-08. Retrieved2020-10-01.
  8. ^abSeneta, Włodzimierz (1981).Drzewa i krzewy iglaste (Coniferous trees and shrubs) (in Polish) (1st ed.). Warsaw: Państwowe Wydawnictwo Naukowe (PWN).ISBN 978-83-01-01663-0.
  9. ^Arno, Stephen F.; Hammerly, Ramona P. (2020) [1977].Northwest Trees: Identifying & Understanding the Region's Native Trees (field guide ed.). Seattle:Mountaineers Books. p. 125.ISBN 978-1-68051-329-5.OCLC 1141235469.Archived from the original on 2023-09-19. Retrieved2022-02-26.
  10. ^Xiang, Xiaoguo; Cao, Ming; Zhou, Zhekun (October 2007)."Fossil history and modern distribution of the genus Abies (Pinaceae)".Frontiers of Forestry in China.2 (4):355–365.doi:10.1007/s11461-007-0058-4.ISSN 1673-3517.Archived from the original on 2024-01-01. Retrieved2023-02-15.
  11. ^Stull, Gregory W.; Qu, Xiao-Jian; Parins-Fukuchi, Caroline; Yang, Ying-Ying; Yang, Jun-Bo; Yang, Zhi-Yun; Hu, Yi; Ma, Hong; Soltis, Pamela S.; Soltis, Douglas E.; Li, De-Zhu; Smith, Stephen A.; Yi, Ting-Shuang; et al. (2021)."Gene duplications and phylogenomic conflict underlie major pulses of phenotypic evolution in gymnosperms".Nature Plants.7 (8):1015–1025.Bibcode:2021NatPl...7.1015S.bioRxiv 10.1101/2021.03.13.435279.doi:10.1038/s41477-021-00964-4.PMID 34282286.S2CID 232282918.Archived from the original on 2022-01-10. Retrieved2023-03-30.
  12. ^Stull, Gregory W.; et al. (2021)."main.dated.supermatrix.tree.T9.tre". Figshare.doi:10.6084/m9.figshare.14547354.v1.Archived from the original on 2024-01-01. Retrieved2023-03-30.{{cite journal}}:Cite journal requires|journal= (help)
  13. ^Knowlton, F.H. (1923). Fossil plants from the Tertiary lake beds of South-Сentral Colorado (Report). Professional Paper. Vol. 131-G. United States Geological Survey. pp. 183–197.doi:10.3133/pp131G.
  14. ^Wolfe, J.A.; Schorn, H.E. (1990). Taxonomic revision of the Spermatopsida of the Oligocene Creede flora, southern Colorado (Report). Bulletin. Vol. 1923. United States Geological Survey. pp. 1–40.doi:10.3133/b1923.
  15. ^Groth, Jacob (10 November 2000)."Monarch Migration Study". Swallowtail Farms.Archived from the original on 9 May 2022. Retrieved21 July 2014.
  16. ^"Monarch Migration". Monarch Joint Venture. 2013.Archived from the original on 2017-10-31. Retrieved2017-10-25.
  17. ^Hemingway, R. W. (2018-01-18). "Bark: Its Chemistry and Prospects for Chemical Utilization".Organic Chemicals from Biomass. CRC Press. pp. 189–248.ISBN 978-1-351-07525-1.
  18. ^Barton, George M. (2018-01-18). "Foliage".Organic Chemicals from Biomass. CRC Press. pp. 249–280.ISBN 978-1-351-07525-1.
  19. ^Jenkins, Steve (2023-09-03)."What is CLS timber and what DIY projects is it good for?".Homebuilding & Renovating. Retrieved2024-08-22.
  20. ^"Christian, T. (2021)Abies from the website Trees and Shrubs Online".Archived from the original on 2023-03-01. Retrieved2023-03-01.
  21. ^Schar, Douglas (2015)."Douglas Fir, Pseudotsuga menziesii".Archives. Doctor Schar.Archived from the original on 2015-10-05. Retrieved2015-10-04.
  22. ^Kershaw, Linda (2000).Edible and Medicinal Plants of the Rockies. Edmonton, AB: Lone Pine Publishing. p. 26.ISBN 978-1-55105-229-8.

Further reading

[edit]
  • Philips, Roger.Trees of North America and Europe, Random House, Inc., New YorkISBN 0-394-50259-0, 1979.

External links

[edit]
Wikimedia Commons has media related toAbies.
Classification ofArchaeplastida orPlantaes.l.
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