These 'reversed' tone letters are used to indicatetone sandhi. For example,˧꜒ after a word or syllable indicates that it has a mid tone in isolation, but a high tone in the current environment.
When this letter occurs in ligature with other tone letters, they together indicate thepitch contour of the tone. For example,꜒ means a level high tone, and꜒꜖꜕ means one that begins high, goes low and ends mid-low.