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Thescelosauridae

From Wikipedia, the free encyclopedia
Extinct family of dinosaurs

Thescelosaurids
Temporal range:EarlyLate Cretaceous,130–66 Ma
Mounted specimen ofThescelosaurus,Burpee Museum of Natural History
Scientific classificationEdit this classification
Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Dinosauria
Clade:Ornithischia
Clade:Pyrodontia
Family:Thescelosauridae
Sternberg,1937
Subfamilies[2]
Synonyms[2]
  • ParksosauridaeBuchholtz, 2002
  • Jeholosauridae?Hanet al., 2012

Thescelosauridae is a clade ofneornithischians from theCretaceous ofEast Asia andNorth America. The group was originally used as a name byCharles M. Sternberg in1937, but was not formally defined until2013, where it was used by Brown and colleagues as the group unitingThescelosaurus andOrodromeus, based on their phylogenetic results.[2][3] During a phylogenetic revision of neornithischians by Clint Boyd in2015, the authorship of Thescelosauridae was given to Brown and colleagues, which meant that the similar nameParksosauridae, informally defined in2002 by Buchholz, would have had priority over Thescelosauridae. The two clades had slightly different definitions, with Parksosauridae referring to all animals closer toParksosaurus thanHypsilophodon, but they contained the same taxa so Boyd used Parksosauridae under the assumption it had priority.[4] However, in formalizing the clade following the regulations of thePhyloCode, Madzia, Boyd, and colleagues identified in2021 that Sternberg was the proper authority for Thescelosauridae, giving it priority over Parksosauridae. As well, they gave Thescelosauridae the definition of the largest clade containingThescelosaurus neglectus but notIguanodon bernissartensis, as long asHypsilophodon foxii was not in the group, modifying previous definitions for Thescelosauridae in order to maintain its modern use, so that the clade was not applied ifThescelosaurus fell withinHypsilophodontidae, a family that has not been recently used but may be revived if the systematic position ofHypsilophodon was solidified at some point in the future.[2] Madziaet al. identified the analysis of Madziaet al. in2018 as the reference analysis for the name Thescelosauridae, an analysis based on a revised version of the 2015 Boyd analysis.[2][5]

Neornithischia

History

[edit]

AlthoughHypsilophodontidae was interpreted as a natural group in the early 1990s,[6][7] this hypothesis has fallen out of favor and Hypsilophodontidae has been found to be an unnatural family composed of a variety of animals more or less closely related to Iguanodontia (paraphyletic), with various smallclades of closely related taxa.[8][9][10][11][12] "Hypsilophodontidae" and "hypsilophodont" are better understood as informal terms for anevolutionary grade, not a true clade.Thescelosaurus has been regarded as both very basal[7] and veryderived[10] among the hypsilophodonts. One issue that has potentially interfered with classifyingThescelosaurus is that not all of the remains assigned toT. neglectus necessarily belong to it.[13] Clint Boyd and colleagues found that while the cladeThescelosaurus included the genusBugenasaura and the species that had been assigned to that genus, there were at least two and possibly three species withinThescelosaurus, and several specimens previously assigned toT. neglectus could not yet be assigned to a species within the genus.[8] It appears to be closely related toParksosaurus,[8][10][11][14][15] although this relationship has been called into question.[16]

The dissolution ofHypsilophodontidae was followed by the recognition of the distinct family Parksosauridae by Buchholz in 2002, defined as the most inclusive clade containingParksosaurus warreni, but notHypsilophodon foxii,Dryosaurus altus, orIguanodon bernissartensis.[17][4] Boydet al. (2009) and Brownet al. (2011) found North American "hypsilophodonts" of Cretaceous age to sort into two related clusters, one (Orodrominae) consisting ofOrodromeus,Oryctodromeus, andZephyrosaurus, and the other (Thescelosaurinae) consisting ofParksosaurus andThescelosaurus.[8][14] Brownet al. (2013) recovered similar results, with the addition of the new genusAlbertadromeus to theOrodromeus clade (calledOrodrominae) and several long-snouted Asian forms (previously described underJeholosauridae)[9] to theThescelosaurus clade (calledThescelosaurinae).[15]

Paleoecology

[edit]

Orodromines may haveburrowed, as is known forOryctodromeus which was found in burrows, based upon the packing of their bones in situations where they typically would have been scattered.[12]

References

[edit]
  1. ^Avrahami, Haviv M.; Makovicky, Peter J.; Tucker, Ryan T.; Zanno, Lindsay E. (2024-07-09)."A new semi-fossorial thescelosaurine dinosaur from the Cenomanian-age Mussentuchit Member of the Cedar Mountain Formation, Utah".The Anatomical Record.307 (12):3717–3781.doi:10.1002/ar.25505.ISSN 1932-8486.PMID 38979930.
  2. ^abcdeMadzia, D.; Arbour, V.M.; Boyd, C.A.; Farke, A.A.; Cruzado-Caballero, P.; Evans, D.C. (2021)."The phylogenetic nomenclature of ornithischian dinosaurs".PeerJ.9 e12362.doi:10.7717/peerj.12362.PMC 8667728.PMID 34966571.
  3. ^Brown, C.M.; Evans, D.C.; Ryan, M.J.; Russell, A.P. (2013). "New data on the diversity and abundance of small-bodied ornithopods (Dinosauria, Ornithischia) from the Belly River Group (Campanian) of Alberta".Journal of Vertebrate Paleontology.33 (3):495–520.Bibcode:2013JVPal..33..495B.doi:10.1080/02724634.2013.746229.S2CID 129160518.
  4. ^abBoyd, Clint A. (2015)."The systematic relationships and biogeographic history of ornithischian dinosaurs".PeerJ.3 e1523.doi:10.7717/peerj.1523.PMC 4690359.PMID 26713260.
  5. ^Madzia, D.; Boyd, C.A.; Mazuch, M. (2017)."A basal ornithopod dinosaur from the Cenomanian of the Czech Republic".Journal of Systematic Palaeontology.16 (11):967–979.doi:10.1080/14772019.2017.1371258.S2CID 90008574.
  6. ^Sues, Hans-Dieter; Norman, David B. (1990). "Hypsilophodontidae,Tenontosaurus, Dryosauridae". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.).The Dinosauria (1st ed.). Berkeley: University of California Press. pp. 498–509.ISBN 978-0-520-06727-1.
  7. ^abWeishampel, David B.; Heinrich, Ronald E. (1992)."Systematics of Hypsilophodontidae and Basal Iguanodontia (Dinosauria: Ornithopoda)"(PDF).Historical Biology.6 (3):159–184.Bibcode:1992HBio....6..159W.doi:10.1080/10292389209380426. Retrieved2007-03-10.
  8. ^abcdBoyd, Clint A.; Brown, Caleb M.; Scheetz, Rodney D.; Clarke, Julia A. (2009). "Taxonomic revision of the basal neornithischian taxaThescelosaurus andBugenasaura".Journal of Vertebrate Paleontology.29 (3):758–770.Bibcode:2009JVPal..29..758B.doi:10.1671/039.029.0328.S2CID 84273584.
  9. ^abHan, Feng-Lu; Paul M. Barrett; Richard J. Butler; Xing Xu (2012). "Postcranial anatomy ofJeholosaurus shangyuanensis (Dinosauria, Ornithischia) from the Lower Cretaceous Yixian Formation of China".Journal of Vertebrate Paleontology.32 (6):1370–1395.Bibcode:2012JVPal..32.1370H.doi:10.1080/02724634.2012.694385.S2CID 86754247.
  10. ^abcNorman, David B.; Sues, Hans-Dieter; Witmer, Larry M.; Coria, Rodolfo A. (2004). "Basal Ornithopoda". In Weishampel, David B.; Dodson, Peter; Osmólska, Halszka (eds.).The Dinosauria (2nd ed.). Berkeley: University of California Press. pp. 393–412.ISBN 978-0-520-24209-8.
  11. ^abWeishampel, David B.; Jianu, Coralia-Maria; Csiki, Z.; Norman, David B. (2003). "Osteology and phylogeny ofZalmoxes (n.g.), an unusual euornithopod dinosaur from the latest Cretaceous of Romania".Journal of Systematic Palaeontology.1 (2):1–56.Bibcode:2003JSPal...1...65W.doi:10.1017/S1477201903001032.S2CID 86339025.
  12. ^abVarricchio, David J.; Martin, Anthony J.; Katsura, Yoshihiro (2007)."First trace and body fossil evidence of a burrowing, denning dinosaur".Proceedings of the Royal Society B: Biological Sciences.274 (1616):1361–1368.doi:10.1098/rspb.2006.0443.PMC 2176205.PMID 17374596.
  13. ^Butler, Richard J.; Upchurch, Paul;Norman, David B. (2008). "The phylogeny of the ornithischian dinosaurs".Journal of Systematic Palaeontology.6 (1):1–40.Bibcode:2008JSPal...6....1B.doi:10.1017/S1477201907002271.S2CID 86728076.
  14. ^abBrown, Caleb M.; Boyd, Clint A.; Russell, Anthony P. (2011)."A new basal ornithopod dinosaur (Frenchman Formation, Saskatchewan, Canada), and implications for late Maastrichtian ornithischian diversity in North America".Zoological Journal of the Linnean Society.163 (4):1157–1198.doi:10.1111/j.1096-3642.2011.00735.x.
  15. ^abBrown, Caleb Marshall; Evans, David C.; Ryan, Michael J.; Russell, Anthony P. (2013). "New data on the diversity and abundance of small-bodied ornithopods (Dinosauria, Ornithischia) from the Belly River Group (Campanian) of Alberta".Journal of Vertebrate Paleontology.33 (3):495–520.Bibcode:2013JVPal..33..495B.doi:10.1080/02724634.2013.746229.S2CID 129160518.
  16. ^Herne, Matthew C.; Nair, Jay P.; Evans, Alistair R.; Tait, Alan M. (2019)."New small-bodied ornithopods (Dinosauria, Neornithischia) from the Early Cretaceous Wonthaggi Formation (Strzelecki Group) of the Australian-Antarctic rift system, with revision of Qantassaurus intrepidus Rich and Vickers-Rich, 1999".Journal of Paleontology.93 (3):543–584.Bibcode:2019JPal...93..543H.doi:10.1017/jpa.2018.95.
  17. ^Buchholz, P. W. (2002). "Phylogeny and biogeography of basal Ornithischia". In Brown, D. E. (ed.).The Mesozoic in Wyoming. Casper: Tate Geological Museum. pp. 18–34.
Avemetatarsalia
Ornithischia
    • see below↓
Heterodontosauridae
Thyreophora
Ankylosauria
Stegosauria
Thescelosauridae
Ornithopoda
Marginocephalia
Ceratopsia
Pachycephalosauria
Laquintasaura venezuelae

Heterodontosaurus tuckiScutellosaurus lawleriKulindadromeus zabaikalicus

Thescelosaurus neglectus
See also
Nomina dubia
Incertae sedis
Other taxa
Other articles
Thescelosauridae
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