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Tetraclinis

From Wikipedia, the free encyclopedia
Genus of conifers

Tetraclinis
Scientific classificationEdit this classification
Kingdom:Plantae
Clade:Tracheophytes
Clade:Gymnospermae
Division:Pinophyta
Class:Pinopsida
Order:Cupressales
Family:Cupressaceae
Subfamily:Cupressoideae
Genus:Tetraclinis
Mast.
Species:
T. articulata
Binomial name
Tetraclinis articulata
Natural range
Synonyms
  • Callitris quadrivalvisRich. & A.Rich.
  • Thuja articulataVahl
Tetraclinis forest atAl Hoceima National Park

Tetraclinis (also called arar,[2] araar[3] or Sictus tree) is agenus ofevergreenconiferous trees in the cypress familyCupressaceae, containing only one species,Tetraclinis articulata, also known asThuja articulata,[4]sandarac,sandarac tree[5] orBarbary thuja,[6]endemic to the westernMediterranean region, particularly insouthern Europe andNorth Africa.

Distribution and habitat

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It is native to northwesternAfrica in theAtlas Mountains ofMorocco,Algeria, andTunisia, with two small outlying populations onMalta, and nearCartagena in southeastSpain inEurope. It grows at relatively low altitudes in a hot, drysubtropicalMediterranean climate.[7]The species has reportedly become aninvasive species in the eastern Mediterranean drylands.[8] The genus once had a much broader range during theOligoceneMiocene, with some species occurring inNorth America.[9]

Taxonomy

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Its closest relatives arePlatycladus,Microbiota, andCalocedrus, with the closest resemblance to the latter. In older texts, it was sometimes treated inThuja orCallitris, but it is less closely related to those genera.[7]

Description

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Tetraclinis cones atAl Hoceima National Park

It is a small, slow-growing tree, to 6–15 m (rarely 20 m) tall and 0.5 m (rarely 1 m) trunk diameter, often with two or more trunks from the base. The foliage forms in open sprays with scale-likeleaves 1–8 mm long and 1–1.5 mm broad; the leaves are arranged in opposite decussate pairs, with the successive pairs closely then distantly spaced, so forming apparent whorls of four. Thecones are 10–15 mm long, green ripening brown in about 8 months from pollination, and have four thick scales arranged in two opposite pairs. Theseeds are 5–7 mm long and 2 mm broad, with a 3–4 mm broad papery wing on each side.[7][10]

It is one of only a small number of conifers able tocoppice (regrow by sprouting from stumps), an adaptation to survivewildfire and moderate levels of browsing by animals. Old trees that have sprouted repeatedly over a long period form large burls at the base, known aslupias.[7]

Uses and symbolism

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It is thenational tree ofMalta, where it is known asgħargħar (derived from the Arabicعَرْعَرʿarʿar). It is now being used locally in afforestation projects.[citation needed]

Theresin, known assandarac, is used to makevarnish andlacquer; it is particularly valued for preservingpaintings.

Thewood, known as thuya wood,[11] citron wood,[4] and alerce,[12] and historically also known asthyine wood, is used for decorative woodwork, particularly wood from burls at the base of the trunk. It has been used thus since antiquity (Ancient Greek:θύον,[13][14]Latin:citrus[15]), and was used to make valuable furniture in the time of theRoman Empire.[16] The market in Morocco is unsustainable, focusing as it does on the burl, and has resulted in mass deforestation of the species. The species is also threatened byovergrazing, which can kill the coppice regrowth before it gets tall enough to be out of the reach of livestock.[7]

Cultivation

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The species is cultivated to be grown as anornamental tree, valued in hot, dry climates. It is also pruned in ahedge form, for privacy and security.[10] The plant can be trained for use asbonsai specimens.

Fossil record

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A relatedextinct species,Tetraclinis salicornioides, has leaf and conefossils ofMessinian age (ca. 5.7 Ma) that have been uncovered in Monte Tondo andBorgo Tossignano, northernApennines,Italy.[17]

Gallery

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Wikimedia Commons has media related toTetraclinis.
  • Tetraclinis articulata in the mountains of Cartagena, Spain
    Tetraclinis articulata in the mountains of Cartagena, Spain
  • A cup made of root burl wood from the Essaouira area of Morocco
    A cup made of root burl wood from the Essaouira area of Morocco
  • Illustration from Koehler's Medicinal-Plants (1887)
    Illustration from Koehler'sMedicinal-Plants (1887)
  • Tetraclinis articulata
    Tetraclinis articulata

References

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  1. ^Sánchez Gómez, P.; Stevens, D.; Fennane, M.; Gardner, M.; Thomas, P. (2011)."Tetraclinis articulata".IUCN Red List of Threatened Species.2011 e.T30318A9534227.doi:10.2305/IUCN.UK.2011-2.RLTS.T30318A9534227.en. Retrieved19 November 2021.
  2. ^"Tetraclinis articulata".The Gymnosperm Database. Retrieved2011-02-13.
  3. ^but it is ambiguous Arabic name also given toJuniperus phoenicea
  4. ^abMemidex: sandarac (wood) Retrieved 2012-05-16
  5. ^Collins: sandarac and sandarac tree Retrieved 2012-05-16
  6. ^Jacques Blondel & James Aronson:Biology and Wildlife of the Mediterranean Region, Oxford University Press 1999 Retrieved 2012-05-16
  7. ^abcdeFarjon, A. (2005).Monograph of Cupressaceae and Sciadopitys. Royal Botanic Gardens, Kew.ISBN 1-84246-068-4
  8. ^Cohen, O.; Sheffer, E.; Bar-Shmuel, N.; Weber, G.; Guy, A.; Avioz, T.; Osem, Y. (2025)."Determinants of naturalization and invasion ofTetraclinis articulata in eastern Mediterranean drylands: Is it a "superinvader"?".Biol. Invasions.27 (3) 94.Bibcode:2025BiInv..27...94C.doi:10.1007/s10530-025-03538-z.
  9. ^Kvaček, Zlatko; Manchester, Steven R.; Schorn, Howard E. (2000)."Cones, Seeds, and Foliage of Tetraclinis salicornioides (Cupressaceae) from the Oligocene and Miocene of Western North America: A Geographic Extension of the European Tertiary Species".International Journal of Plant Sciences.161 (2):331–344.doi:10.1086/314245.ISSN 1058-5893.
  10. ^abRushforth, K. (1999).Trees of Britain and Europe. CollinsISBN 0-00-220013-9.
  11. ^Arc-genesis: Thuya WoodArchived 2011-08-08 at theWayback Machine Retrieved 2012-05-16
  12. ^"Definition of ALERCE".
  13. ^Liddell & Scott."θύον".A Greek-English Lexicon – viaLogeion.
  14. ^Beekes, Robert S. P. (2009). "θύον".Etymological Dictionary of Greek. Brill. p. 565.
  15. ^de Vaan, Michiel (2008)."citrus".Etymological Dictionary of Latin and the other Italic Languages. Brill. p. 116.
  16. ^Lewis & Short."citrus".A Latin Dictionary – viaLogeion.
  17. ^Palaeoenvironmental analysis of the Messinian macrofossil floras of Tossignano and Monte Tondo (Vena del Gesso Basin, Romagna Apennines, northern Italy) - Vasilis Teodoridis, Zlatko Kvacek, Marco Sami and Edoardo Martinetto - December 2015 DOI: 10.14446/AMNP.2015.249.
ExtantCupressaceae species
SubfamilyAthrotaxidoideae
Athrotaxis
SubfamilyCallitroideae
Actinostrobus
Austrocedrus
Callitris
Diselma
Fitzroya
Libocedrus
Neocallitropsis
Papuacedrus
Pilgerodendron
Widdringtonia
SubfamilyCunninghamioideae
Cunninghamia
SubfamilyCupressoideae
Callitropsis
Calocedrus
Chamaecyparis
Cupressus
Hesperocyparis
Fokienia
Juniperus
SectionJuniperus
SectionSabina
Microbiota
Platycladus
Tetraclinis
Thuja
Thujopsis
Xanthocyparis
SubfamilySequoioideae
Metasequoia
Sequoia
Sequoiadendron
SubfamilyTaiwanioideae
Taiwania
SubfamilyTaxodioideae
Cryptomeria
Glyptostrobus
Taxodium
Tetraclinis
Tetraclinis articulata
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