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Soomaspis

From Wikipedia, the free encyclopedia
Marine arthropods that lived during the late Ordovician

Soomaspis splendida
Temporal range:earlyHirnantian
Reconstruction ofSoomaspis splendida
Scientific classification
Kingdom:
Phylum:
Order:
Family:
Genus:
Soomaspis
Species:
S. splendida
Binomial name
Soomaspis splendida
Fortey & Theron, 1994

Soomaspis is agenus of small to average size (about 3 cm or 1.2-inch long) marinearthropods in the Liwiidae Family, that lived during the lateOrdovician[1] (early Hirnantian).[2]Fossil remains ofSoomaspis were collected from theSoom ShaleLagerstätte inWestern Cape,South Africa.Soomaspis looks like a large, softagnostidtrilobite. It has a headshield (orcephalon) wider than the tailshield (pygidium), and in between them threethoracic body segments (somites). The genus ismonotypic, its solespecies beingSoomaspis splendida.

Etymology

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The name of the genus is a compound word of the deposit where the species was collected (the Soom Shale), and the Greek word"aspis" (shield). The species epithet,splendida comes from the Latin word"splendere" (brightness).

Description

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Soomaspis splendida is estimated to be over 3 cm (1.2 inches) along the axis,[3]1+12 times longer than wide. Thedorsalexoskeleton consists of a cephalon, apygidium and two or three thoracicsomites with articulating half-rings, all non-calcified, supposedly of medium convexity. The axis is poorly defined. The cephalon is transversely oval, widest at midlength. The cephalon is wider than the pygidium. Eyes are absent. Antennas are not known. The body is constricted at the three thoracic somites, so the animal gives the impression to have a waist. The pygidium is slightly wider than long, with the greatest width at midlength. The pygidium has a mid-ridge and five segments divided by clear furrows on the outer parts of the pleural field. The back edge of these furrows curve backwards, ending at a sharp angle to the pygidial margin.[1]

Differences with other Liwiidae

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  • Soomaspis splendida differs fromLiwia by having 3 thoracic somites instead of 4.S. splendida has a mid-length ridge on the pygidium and furrows on the outer parts of the pleural field, which are unknown fromLiwia.Soomaspis also has a pygidium with an entire margin, whileLiwia has five pairs of modest marginal spines, and a concave posterior border.
  • S. splendida differs fromBuenaspis forteyi, that has a cephalon and pygidium that are not wider than its 6 thoracic somites. The pygidium ofB. forteyi is wider than long, with a straight anterior border.Buenaspis andSoomaspis have an entire margin and a mid-ridge on the pygidium in common.
  • S. splendida differs fromTariccoia arrusensis, that lacks visible segmentation of the pygidium. The surface of the exoskeleton ofS. splendida, particularly the pygidium, shows dense pitting, while it is smooth inT. arrusensis.Tariccoia andSoomaspis both have a broad doublure and lack a lengthwise mid-ridge on the cephalon. In both species the pygidium is about as long as wide, but inSoomaspis the widest point is around midlength, whileTariccoia is widest in the frontal half. The species share 3 thoracic somites, an entire margin and a mid-ridge on the pygidium.[4]

Distribution

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Soomaspis splendida has been collected from the Soom Shale (earlyHirnantian), Keurbos Farm, near Clanwilliam, Cape Province, South Africa.[1]

Habitat

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Soomaspis splendida was probably a marine bottom dweller. The Soom Shale is sometimes interfingered with the glacialtillite of the Pakhuis Formation, indicating thatSoomaspis lived in the open sea, near the edge of an ice sheet.[1]

References

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  1. ^abcdR.A. Fortey and J.N. Theron (1994). "A New Ordovician Arthropod,Soomaspis, and the Agnostid Problem arthropod".Palaeontology.37 (part 4):841–861 (PDF).
  2. ^Fossil Lagerstätten, a catalogue of sites of exceptional fossil preservation, produced by MSc Palaeobiology students atUniversity of Bristol's Department of Earth Sciences, web:lagerstattenArchived 2013-02-13 at theWayback Machine.
  3. ^L. Ramskold, J.-Y. Chen, G.D. Edgecombe, and G.-Q. Zhou (1996). "Preservational folds simulating tergite junctions in tegopeltid and naraoiid arthropods".Lethaia29: 15-20.ISSN 0024-1164, web:Preserv-folds.
  4. ^E. Bonino & C. Kier (2010).The Back to the Past Museum Guide to Trilobites. pp. 18–19, fig. 9.
Protosutura
Cheloniellida
Aglaspidida
Tremaglaspididae
Aglaspididae
Xandarellida
Nektaspida
Emucarididae
Naraoiidae
Liwiidae
Conciliterga
Trilobita
Cheloniellon calmani

TriarthrusAglaspis barrandei

Retifacies abnormalis
Soomaspis
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