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Sequoia sempervirens

From Wikipedia, the free encyclopedia
Species of tree
"California Redwoods" redirects here. For the professional lacrosse team, seeCalifornia Redwoods (PLL). For the defunct UFL football team, seeSacramento Mountain Lions.

Sequoia sempervirens
Sequoia sempervirens alongUS 199

Apparently Secure (NatureServe)[2]
Scientific classificationEdit this classification
Kingdom:Plantae
Clade:Tracheophytes
Clade:Gymnospermae
Division:Pinophyta
Class:Pinopsida
Order:Cupressales
Family:Cupressaceae
Genus:Sequoia
Species:
S. sempervirens
Binomial name
Sequoia sempervirens
Natural range of California subfamilySequoioideae
green -Sequoia sempervirens
Trunk in sectional view
Redwood cone scales begin to open mid November, with seeds dispersing by the wind.

Sequoia sempervirens (/səˈkwɔɪ.əˌsɛmpərˈvrənz/)[3] is the sole livingspecies of thegenusSequoia in the cypress familyCupressaceae (formerly treated inTaxodiaceae). Common names includecoast redwood,coastal redwood andCalifornia redwood. It is anevergreen, long-lived,monoecious tree living 1,200–2,200 years or more.[4] This species includes thetallest living trees on Earth, reaching up to 115.9 m (380.1 ft) in height (without theroots) and up to 8.9 m (29 ft) indiameter at breast height. These trees are also among thelongest-living trees on Earth. Before commercial logging and clearing began by the 1850s, this massive tree occurrednaturally in an estimated 810,000 ha (2,000,000 acres)[5][6][7] along much of coastal California (excluding southern California where rainfall is not sufficient) and the southwestern corner of coastal Oregon within the United States. Being the tallest tree species, with a small range and an extremely long lifespan, many redwoods are preserved in various state and national parks; many of the largest specimens have their own official names.

The namesequoia sometimes refers to the subfamilySequoioideae, which includesS. sempervirens along withSequoiadendron (giant sequoia) andMetasequoia (dawn redwood). Here, the termredwood on its own refers to the species covered in this article but not to the other two species.

Description

[edit]

The coast redwood normally reaches a height of 60 to 100 m (200 to 330 ft), but can be more than 110 m (360 ft) in some cases,[8] with a trunk diameter of 9 m (30 ft).[9] Historically, theAmericannaturalist,physician and founder member of theCalifornia Academy of Sciences,William P. Gibbons (1812–1897) described in 1893 the hollow shell of a coast redwood in theOakland Hills ofAlameda County with diameter of 9.9 metres (32 ft) at breast height.[10] This tree's girth is rivalled by the "Fieldbrook Stump" ofHumboldt County with a diameter of 9.8 metres (32 ft) at 1.5 metres (4.9 ft) from the ground.[11] This stump is still intact, with numerous healthystump shoots encircling it.

Coast redwoods have a conicalcrown, with horizontal to slightly drooping branches. The trunk is remarkably straight. Thebark can be very thick, up to 35 cm (1.15 ft), and quite soft and fibrous,[9] with a bright red-brown color when freshly exposed (hence the name redwood),weathering darker. Theroot system is composed of shallow, wide-spreadinglateral roots.

The leaves are variable, being 15–25 mm (58–1 in) long and flat on young trees and shaded lower branches in older trees. The leaves are scalelike,5–10 mm (1438 in) long on shoots in full sun in the upper crown of older trees, with a full range of transition between the two extremes. They are dark green above and have two blue-whitestomatal bands below. Leaf arrangement is spiral, but the larger shade leaves are twisted at the base to lie in a flat plane for maximum light capture.

The species ismonoecious, with pollen and seed cones on the same plant. Theseed cones are ovoid,15–32 mm (9161+14 in) long, with 15–25 spirally arranged scales;pollination is in late winter with maturation about 8–9 months after. Each cone scale bears three to sevenseeds, each seed3–4 mm (18316 in) long and0.5 mm (132 in) broad, with two wings1 mm (116 in) wide. The seeds are released when the cone scales dry and open at maturity. The pollen cones are ovular and4–6 mm (31614 in) long.

Genetics

[edit]

Itsgenetic makeup (chromosome count is 2n=66) is unusual among conifers, being ahexaploid (6n) and possiblyallopolyploid (AAAABB).[12] Both themitochondrial andchloroplast genomes of the redwood arepaternally inherited.[13] According toGuinness the Coast Redwood has the largest knowngenome of any plant examined up to the year 2020, totaling 26.5 gigabases.[14]

Taxonomy

[edit]

Scottish botanistDavid Don described the redwood asTaxodium sempervirens, the "evergreenTaxodium", in his colleagueAylmer Bourke Lambert's 1824 workA description of the genus Pinus.[15] Austrian botanistStephan Endlicher erected the genusSequoia in his 1847 workSynopsis coniferarum, giving the redwood its current binomial name ofSequoia sempervirens.[16] It is unknown how Endlicher derived the name Sequoia. SeeSequoia Etymology.

The redwood is one of three living species, each in its own genus, in the subfamilySequoioideae. Molecular studies have shown that the three are each other's closest relatives, generally with the redwood and giant sequoia (Sequoiadendron giganteum) as each other's closest relatives.

However, Yang and colleagues in 2010 queried the polyploid state of the redwood and speculate that it may have arisen as an ancienthybrid between ancestors of the giant sequoia and dawn redwood (Metasequoia). Using two different single copynuclear genes,LFY and NLY, to generatephylogenetic trees, they found thatSequoia was clustered withMetasequoia in the tree generated using the LFY gene, but withSequoiadendronin the tree generated with the NLY gene. Further analysis strongly supported the hypothesis thatSequoia was the result of ahybridization event involvingMetasequoia andSequoiadendron. Thus, Yang and colleagues hypothesize that the inconsistent relationships amongMetasequoia,Sequoia, andSequoiadendron could be a sign ofreticulate evolution (in which two species hybridize and give rise to a third) among the three genera. However, the long evolutionary history of the three genera (the earliest fossil remains being from theJurassic) make resolving the specifics of when and howSequoia originated once and for all a difficult matter—especially since it in part depends on an incompletefossil record.[17]

Names

[edit]

The species name "sempervirens" means "evergreen", thought to be because of its previous placement in the same genus asTaxodium distichum (baldcypress) of the southeastern USA. Unlike coast redwood, baldcypress loses its leaves in winter.[18] The common name "redwood", applied to both the coast redwood and the giant redwood, is a reference to the redheartwood of the trees.[19] Common names that refer toSequoia sempervirens alone include "California redwood", "coastal redwood", "coastal sequoia", and "coast redwood".[20]

Distribution and habitat

[edit]

Coast redwoods occupy a narrow strip of land approximately 750 km (470 mi) in length and 8–75 km (5–47 mi) in width along the Pacific coast of North America; the most southerly native grove is inMonterey County, California, and the most northerly groves are in extreme southwestern Oregon. The aforementioned qualification of "native" is because the species was introduced to various locations inVictoria, Australia, in the 1930s for experimental reasons and have since flourished. The prevailing native elevation range is 30–750 m (100–2,460 ft) above sea level, occasionally down to 0 and up to about 900 m (3,000 ft).[21] They usually grow in the mountains where precipitation from the incoming moisture off the ocean is greater. The tallest and oldest trees are found in deep valleys and gullies, where year-round streams can flow, and fog drip is regular. Theterrain also made it harder for loggers to get to the trees and to get them out afterfelling. The trees above the fog layer, above about 700 m (2,300 ft), are shorter and smaller due to the drier, windier, and colder conditions. In addition,Douglas-fir,pine, andtanoak often crowd out redwoods at these elevations. Few redwoods grow close to the ocean, due to intense salt spray, sand, and wind.Coalescence of coastal fog accounts for a considerable part of the trees' water needs.[22] Fog in the 21st century is, however, reduced from what it was in the prior century, which is a problem that may be compounded byclimate change.[23]

Painting of a hillside of redwoods with the edge sun seen behind a hill
Albert Bierstadt,California Redwoods, 1872, Oil on paper mounted on canvas

The northern boundary of its range is marked by two groves on mountain slopes along the north side of theChetco River,[24] which is on the western fringe of theKlamath Mountains, near the California–Oregon border.[25][26] The northernmost grove is located withinAlfred A. Loeb State Park andSiskiyou National Forest at the approximate coordinates 42°07'36"N 124°12'17"W. The southern boundary of its range is theLos Padres National Forest'sSilver Peak Wilderness in theSanta Lucia Mountains of theBig Sur area of Monterey County, California. The southernmost grove is in theSouthern Redwood Botanical Area, just north of the national forest's Salmon Creek trailhead and near theSan Luis Obispo County line.[27][28]

The largest and tallest populations are in California'sRedwood National and State Parks (Del Norte andHumboldt counties) andHumboldt Redwoods State Park, with the overall majority located in the large Humboldt County.

The ancient range of the genus is considerably greater, with relatives of the coast redwood living in Europe and Asia prior to theQuaternary geologic period. In recent geologic time there have been considerable shifts in the coast redwood's range in North America. Coast redwood bark has been found in theLa Brea Tar Pits, showing that 25,000–40,000 years before the present redwood trees grew as far south as the Los Angeles during thelast ice age.[29][30] The authors of a 2022 paper suggest, "Were it not for the remarkable ability to sprout after fire, many southern forests may have lost theirSequoia component long ago."[31] As to previous redwood range to the north, an upright fossil stump of a coast redwood on a beach in central Oregon was documented 257 km (160 mi) north of the current range.[32]

Assisted migration

[edit]
Coast redwood (2016 photo) planted in Seabeck, WA, in 1949.

The ability of Coast Redwood to live for more than a thousand years, along with its unusual capacity to resprout from its root crown when felled by natural or human causes, have earned this species the label of "carbon-sequestration champion."[33] Its potential to contribute towardclimate change mitigation, as well as its demonstrated ability to thrive in coastal regions of thePacific Northwest,[34][35] led to the formation of a citizen group in Seattle, Washington undertakingassisted migration of this species hundreds of miles north of itsnative range.[36][37]

In contrast to cautionary statements made by forestry professionals assessing other tree species for assisted migration,[38] the citizens involved with the group known as PropagationNation had met with little controversy until in 2023 a national news outlet published a lengthy article that cast a favorable light on their efforts. TheNew York Times Magazine wrote:

Not wanting to cause ecological problems by planting the trees across the Pacific Northwest, [Philip] Stielstra would eventually contact one of the foremost experts on the coast redwood, a botanist and forest ecologist namedStephen Sillett, at Cal Poly Humboldt, and ask if moving redwoods north was safe. Sillett thought planting redwoods around Seattle was a fantastic idea. ("It's not like it's going to escape and become a nuisance species," Sillett told me, before adding, "it just has so many benefits.") Another factor encouraged Stielstra too: Millions of years ago, redwoods — or their close relatives — grew across the Pacific Northwest. By moving them, Stielstra reasoned, he was helping the magnificent trees regain lost territory.[36]

In December 2023, theAssociated Press exclusively reported criticism from professionals in the region and nationally: While beginning to favor experiments inassisted population migration of more southerly genetics of the main native timber tree,Douglas fir, professionals were united against large-scale plantings of California redwoods into the Pacific Northwest.[39] The next month, January 2024, carried a regional news article that, once again, showed strong support as well as bold statements by the group's founder.[40]

Even before the controversy developed in Washington state, professionals in Canada were documenting horticultural plantings of the California species already in place in southwesternBritish Columbia. In 2022 a Canadian Forestry Service publication used northward horticultural plantings, along with a review of research detailing redwood'spaleobiogeography and current range conditions, as grounds for proposing that Canada'sVancouver Island already offered "narrow strips of optimal habitat" for extending the range of coast redwood.[41] The authors point to a topographical "bottleneck" north of the California border that could have impeded northward migration during the Holocene. The bottleneck entails a lack of lowland passages through theOregon Coast Range north of theChetco River, coupled with the absence of coastal landscapes beyond storm salt-spray andtsunami inundation — for which this conifer species is highly intolerant.

Ecology

[edit]

Fog and flood adaptations

[edit]
Fog is of major importance in coast redwood ecology.Redwood National Park.
Cross section of a Sequoia sempervirens showing tree rings
Young, 3 m (9.8 ft) coast redwood growing in sediment withbroadleaf cattails in flume pond, Spring Creek,Filoli Estate, June 2025.

The native area provides a unique environment with heavy seasonal rains up to 2,500 mm (100 in) annually. Cool coastal air andfog drip keep the forest consistently damp year round. Several factors, including the heavy rainfall, create a soil with fewer nutrients than the trees need, causing them to depend heavily on the entire biotic community of the forest, and making efficient recycling of dead trees especially important. This forest community includescoast Douglas-fir,Pacific madrone,tanoak,western hemlock, and other trees, along with a wide variety offerns,mosses,mushrooms, andredwood sorrel. Redwood forests provide habitat for a variety of amphibians, birds, mammals, and reptiles.Old-growth redwood stands provide habitat for the federally threatenedspotted owl and the California-endangeredmarbled murrelet.

Although fog is not required for redwoods growth,[42][43] the height ofS. sempervirens is closely tied to fog availability; taller trees become less abundant as fog becomes less frequent.[44] As tree height increases, transporting water viawater potential to the leaves becomes increasingly difficult due to gravity.[45][46][47] Despite the high rainfall that the region receives (up to 100 cm), the leaves in the upper canopy are perpetually stressed for water.[48][49] Thiswater stress is exacerbated by long droughts in the summer.[50] Water stress is believed to cause the morphological changes in the leaves, stimulating reduced leaf length and increased leaf succulence.[46][51] To supplement their water needs, redwoods use frequent summer fog events. Fog water is absorbed through multiple pathways. Leaves directly take in fog from the surrounding air through theepidermal tissue, bypassing thexylem.[52][53] Coast redwoods also absorb water directly through their bark.[54] The uptake of water through leaves and bark repairs and reduces the severity of xylem embolisms,[55][54] which occur whencavitations form in the xylem preventing the transport of water and nutrients.[54] Fog may also collect on redwood leaves, drip to the forest floor, and be absorbed by the tree's roots. This fog drip may constitute 30% of the total water used by a tree in a year.[50]

Redwoods often grow in flood-prone areas. Sediment deposits can form impermeable barriers that suffocate tree roots, and unstable soil in flooded areas often causes trees to lean to one side, increasing the risk of the wind toppling them. Immediately after a flood, redwoods grow their existing roots upwards into recently deposited sediment layers.[56] A second root system then develops from adventitious buds on the newly buried trunk and the old root system dies.[56] To counter lean, redwoods increase wood production on the vulnerable side, creating a supporting buttress.[56] These adaptations create forests of almost exclusively redwood trees in flood-prone regions.[57][56]

Pest and pathogen resistance

[edit]
A recently fallen canopy branch reveals thin bark and tunnels made by bark beetle larvae.

Coast redwoods are resistant to insect attack, fungal infection, and rot. These properties are conferred by concentrations ofterpenoids andtannic acid in redwood leaves, roots, bark, and wood.[57] Despite these chemical defenses, redwoods are still subject to insect infestations; none, however, are capable of killing a healthy tree. Bark is so thick on the bole that bark beetles cannot enter there. However, the canopy branches have thin bark (see photo at right) that native bark beetles are able to bore into for egg-laying and larval growth via tunnels.[57]

Redwoods also face herbivory from mammals:black bears are reported to consume the inner bark of small redwoods, andblack-tailed deer are known to eat redwood sprouts.[57]

The oldest known coast redwood is about 2,200 years old;[8] many others in the wild exceed 600 years. The numerous claims of older redwoods are incorrect.[8] Because of their seemingly timeless lifespans, coast redwoods were deemed the "everlasting redwood" at the turn of the century; inLatin,sempervirens means "ever green" or "everlasting". Redwoods must endure various environmental disturbances to attain such great ages.

Fire adaptations

[edit]

In response to forest fires, the trees have developed various adaptations. The thick, fibrous bark of coast redwoods is extremely fire-resistant; it grows to at least 30 cm (12 in) thick and protects mature trees from fire damage.[58][59] In addition, the redwoods contain little flammablepitch or resin.[59] Fires, moreover, appear to actually benefit redwoods by causing substantial mortality in competing species,[60] while having only minor effects on redwood. Burned areas are favorable to the successful germination of redwood seeds.[58] A study published in 2010, the first to compare post-wildfire survival and regeneration of redwood and associated species, concluded that fires of all severity increase the relative abundance of redwood, and higher-severity fires provide the greatest benefit.[61]

Two years after the 2020CZU fire that burned as high as the canopy inBig Basin Redwoods State Park. In addition to the customary sprouting of massive numbers of vertical shoots growing at ground-level around the charred trunks, thickepicormic greenery appears high up on some of the fire-pruned redwoods. (Photos were taken along the old growth loop trail, September 2022.)

Self-pruning of its lower limbs as height is gained is a crucial adaptation to prevent ground fires from rising into the canopy, where branch bark is thin and leaves are vulnerable. In the millions of years that predated human evolution, this level of protection worked well against natural fires originating from lightning strikes.[60]

When the first humans arrived in North America, redwoods thrived in regions where ground fires were intentionally set byindigenous populations on a seasonal basis.[62] However, when peoples arrived from other continents in the past few centuries,indigenous fire practices were disallowed — even in the few places where the first peoples were permitted to continue living. Flammable brush and young trees thus accumulated.[62]

Clearcut logging further hampered a return to a fire-resistant tall canopy.Governmental policies aimed atsuppressing all natural and human-caused fires, even in parks and wilderness areas, amplified the accumulation of dense undergrowth and woody debris. Thus, even a naturally occurring ground fire could threaten to spread upwards and become a canopy fire spreading uncontrollably over a widening area.[63][64]

Reproduction

[edit]
Lignotuber actively budding.
Lignotuber, Roosevelt Grove.
Basal regrowth after logging.
View upward from the center of an old, logged stump (Freshwater Creek).

Coast redwood reproduces both sexually by seed and asexually by sprouting of buds, layering, orlignotubers. Seed production begins at 10–15 years of age. Cones develop in the winter and mature by fall. In the early stages, the cones look likeflowers, and are commonly called "flowers" by professional foresters, although this is not strictly correct. Coast redwoods produce many cones, with redwoods in new forests producing thousands per year.[58] The cones themselves hold 90–150 seeds, but viability of seed is low, typically well below 15% with one estimate of average rates being 3 to 10 percent.[65][58] The viability does increase with age, trees under 20 years old have a viability of about 1%, and do not generally reach the highest levels of viability until age 250. The rates decrease as the tree starts to get very old, with trees over 1,200 not reaching viability rates over 3%.[66] The low viability may discourage seed predators, which do not want to waste time sortingchaff (empty seeds) from edible seeds. Successful germination often requires a fire or flood, reducing competition for seedlings. The winged seeds are small and light, weighing 3.3–5.0 mg (200–300 seeds/g; 5,600–8,500/ounce). The wings are not effective for wide dispersal, and seeds are dispersed by wind an average of only 60–120 m (200–390 ft) from the parent tree. Seedlings are susceptible to fungal infection and predation bybanana slugs,brush rabbits, andnematodes.[57] Most seedlings do not survive their first three years.[58] However, those that become established grow rapidly, with young trees known to reach 20 m (66 ft) tall in 20 years. When canopy space is not available, small trees can remain suppressed for up to 400 years before accelerating their growth rate.[67]

Coast redwoods can also reproduce asexually by layering or sprouting from the root crown, stump, or even fallen branches; if a tree falls over, it generates a row of new trees along the trunk, so many trees naturally grow in a straight line. Sprouts originate from dormant or adventitious buds at or under the surface of the bark. The dormant sprouts are stimulated when the main adult stem gets damaged or starts to die. Many sprouts spontaneously erupt and develop around the circumference of the tree trunk. Within a short period after sprouting, each sprout develops its own root system, with the dominant sprouts forming a ring of trees around the parent root crown or stump. This ring of trees is called a "fairy ring". Sprouts can achieve heights of 2.3 m (7 ft 7 in) in a single growing season.

Redwoods may also reproduce usingburls. A burl is a woodylignotuber that commonly appears on a redwood tree below the soil line, though usually within 3 m (10 ft) in depth from the soil surface. Coast redwoods develop burls as seedlings from the axils of theircotyledon, a trait that is extremely rare in conifers.[58] When provoked by damage, dormant buds in the burls sprout new shoots and roots. Burls are also capable of sprouting into new trees when detached from the parent tree, though exactly how this happens is yet to be studied. Shoot clones commonly sprout from burls and are often turned into decorative hedges when found in suburbia.

Cultivation and uses

[edit]
An example of abonsai redwood, from the Pacific Bonsai Museum

Coast redwood is one of the most valuable timber species in the lumbering industry. In California, 3,640 km2 (899,000 acres) of redwood forest are logged, virtually all of it second growth.[1] Though many entities have existed in the cutting and management of redwoods, perhaps none has had a more storied role than thePacific Lumber Company (1863–2008) of Humboldt County, California, where it owned and managed over 810 km2 (200,000 acres) of forests, primarily redwood. Coast redwood lumber is highly valued for its beauty, light weight, and resistance to decay. Its lack of resin makes it absorb water[68] and resist fire.

P. H. Shaughnessy, Chief Engineer of the San Francisco Fire Department wrote:

In the recentgreat fire of San Francisco, that began April 18th, 1906, we succeeded in finally stopping it in nearly all directions where the unburned buildings were almost entirely of frame construction, and if the exterior finish of these buildings had not been of redwood lumber, I am satisfied that the area of the burned district would have been greatly extended.

Because of its impressive resistance to decay, redwood was extensively used forrailroad ties andtrestles throughout California. Many of the old ties have been recycled for use in gardens as borders, steps, house beams, etc. Redwood burls are used in the production of table tops, veneers, and turned goods.

TheSkyline-to-the-Sea Trail passing through a cut-out in a fallen California redwood tree

TheYurok people, who occupied the region before European settlement, regularly burned ground cover in redwood forests to bolstertanoak populations from which they harvested acorns, to maintain forest openings, and to boost populations of useful plant species such as those for medicine or basketmaking.[57]

Extensive logging of redwoods began in the early nineteenth century. The trees were felled by ax and saw onto beds of tree limbs and shrubs to cushion their fall.[57] Stripped of their bark, the logs were transported to mills or waterways by oxen or horse.[57] Loggers then burned the accumulated tree limbs, shrubs, and bark. The repeated fires favoredsecondary forests of primarily redwoods as redwood seedlings sprout readily in burned areas.[57][69] The introduction of steam engines let crews drag logs through long skid trails to nearby railroads,[57] furthering the reach of loggers beyond the land near rivers previously used to transport trees.[69] This method of harvesting, however, disturbed large amounts of soil, producing secondary-growth forests of species other than redwood such as Douglas-fir,grand fir, and western hemlock.[57] After World War II, trucks and tractors gradually replaced steam engines, giving rise to two harvesting approaches:clearcutting and selection harvesting. Clearcutting involved felling all the trees in a particular area. It was encouraged by tax laws that exempted all standing timber from taxation if 70% of trees in the area were harvested.[57] Selection logging, by contrast, called for the removal 25% to 50% of mature trees in the hopes that the remaining trees would allow for future growth and reseeding.[69] This method, however, encouraged growth of other tree species, converting redwood forests into mixed forests of redwood, grand fir,Sitka spruce, and western hemlock.[69][57] Moreover, the trees left standing were often felled bywindthrow; that is, they were often blown over by the wind.

The coast redwood isnaturalized in New Zealand, notably atWhakarewarewa Forest,Rotorua.[70] Redwood has been grown in New Zealand plantations for more than 100 years, and those planted in New Zealand have higher growth rates than those in California, mainly because of even rainfall distribution through the year.[71]

Other areas of successful cultivation outside of the native range include Great Britain, Italy, France,[72]Haida Gwaii, middle elevations of Hawaii,Hogsback in South Africa, the Knysna Afromontane forests in the Western Cape, Grootvadersbosch Forest Reserve near Swellendam, South Africa and the Tokai Arboretum on the slopes of Table Mountain above Cape Town, a small area in central Mexico (Jilotepec), and the southeastern United States from eastern Texas to Maryland. It also does well in the Pacific Northwest (Oregon, Washington, and British Columbia), far north of its northernmost native range in southwestern Oregon. Coast redwood trees were used in a display at Rockefeller Center and then given to Longhouse Reserve in East Hampton, Long Island, New York, and these have now been living there for over twenty years and have survived at 2 °F (−17 °C).[73]

This fast-growing tree can be grown as an ornamental specimen in those large parks and gardens that can accommodate its massive size. It has gained theRoyal Horticultural Society'sAward of Garden Merit.[74][75]

Statistics

[edit]
Driedresin
The foliage of an "albino" exhibiting lack of chlorophyll
"Albinos" lackchlorophyll

Fairly solid evidence indicates that coast redwoods were the world's largest trees before logging, with numerous historical specimens reportedly over 122 m (400 ft).[76]: 16, 42  The theoretical maximum potential height of coast redwoods is thought to be limited to between 122 and 130 m (400 and 427 ft), asevapotranspiration is insufficient to transport water to leaves beyond this range.[45] Further studies have indicated that this maximum requires fog, which is prevalent in these trees' natural environment.[77]

A tree reportedly 114.3 m (375 ft) in length was felled in Sonoma County by the Murphy Brothers saw mill in the 1870s,[78] another claimed to be 115.8 m (380 ft) and 7.9 m (26 ft) in diameter was cut down near Eureka in 1914,[79][80] and theLindsay Creek Tree was documented to have a height of 120 m (390 ft) when it was uprooted and felled by a storm in 1905. A tree reportedly 129.2 m (424 ft) tall was felled in November 1886 by the Elk River Mill and Lumber Company in Humboldt County, yielding 79,736 marketable board feet from 21 cuts.[81][82][83] In 1893, a Redwood cut at theEel River, nearScotia, reportedly measured 130.1 m (427 ft) in length, and 23.5 m (77 ft) in girth.[84][85][86] However, limited evidence corroborates these historical measurements.

Today, trees over 60 m (200 ft) are common, and many are over 90 m (300 ft). The current tallest tree is theHyperion tree, measuring 116.07 m (380.8 ft).[8] The tree was discovered in Redwood National Park during mid-2006 by Chris Atkins andMichael Taylor, and is thought to be the world's tallest living organism. The previous record holder was theStratosphere Giant in Humboldt Redwoods State Park at 112.84 m (370.2 ft) (as measured in 2004). Until it fell in March 1991, the "Dyerville Giant" was the record holder. It, too, stood in Humboldt Redwoods State Park and was 113.4 m (372 ft) high and estimated to be 1,600 years old. This fallen giant has been preserved in the park.

The largest known living coast redwood is Grogan's Fault, discovered in 2014 by Chris Atkins and Mario Vaden in Redwood National Park,[8] with a main trunk volume of at least 1,084.5 cubic meters (38,299 cu ft)[8] Other high-volume coast redwoods includeIluvatar, with a main trunk volume of 1,033 m3 (36,470 cu ft),[76]: 160  and theLost Monarch, with a main trunk volume of 988.7 m3 (34,914 cu ft).[87]

Albino redwoods aremutants that cannot manufacturechlorophyll. About 230 examples (including growths and sprouts) are known to exist,[88] reaching heights of up to 20 m (66 ft).[89] These trees survive likeparasites, obtaining food from green parent trees. While similar mutations occur sporadically in other conifers, no cases are known of such individuals surviving to maturity in any other conifer species.[citation needed] Recent research news reports that albino redwoods can store higher concentrations of toxic metals, going so far as comparing them to organs or "waste dumps".[90][91]

List of tallest trees

[edit]
See also:List of tallest trees

Heights of the tallest coast redwoods are measured yearly by experts.[8] Even with recent discoveries of tall coast redwoods above 100 m (330 ft), it is likely that no taller trees will be discovered.[8]

Ten tallestSequoia sempervirens[8]
RankNameHeightDiameterLocation
MetersFeetMetersFeet
1Hyperion115.85380.14.8415.9Redwood National Park
2Helios114.58375.94.9616.3Redwood National Park
3Icarus113.14371.23.7812.4Redwood National Park
4Stratosphere Giant113.05370.95.1817.0Humboldt Redwoods State Park
5National Geographic112.71369.84.3914.4Redwood National Park
6Orion112.63369.54.3314.2Redwood National Park
7Federation Giant112.62369.54.5414.9Humboldt Redwoods State Park
8Paradox112.51369.13.9012.8Humboldt Redwoods State Park
9Mendocino112.32368.54.1913.7Montgomery Woods State Natural Reserve
10Millennium111.92367.22.718.9Humboldt Redwoods State Park
TheDel Norte Titan

Diameter is measured at 1.4 m (4 ft 7 in) above average ground level (at breast height). Details of the precise locations for most of the tallest trees were not announced to the general public for fear of causing damage to the trees and the surrounding habitat.[8] The tallest coast redwood easily accessible to the public is the National Geographic Tree, immediately trailside in the Tall Trees Grove of Redwood National Park.[92]

List of largest trees

[edit]

The following list shows the largestS. sempervirens by volume known as of 2001.[76]: 186–7 

Five largestSequoia sempervirens[76]: 186–7 
RankNameHeightDiameterTrunk volumeLocation
MetersFeetMetersFeetCubic metersCubic feet
1Del Norte Titan93.63077.2323.71,04536,900Jedediah Smith Redwoods State Park
2Iluvatar91.43006.1420.11,03336,500Prairie Creek Redwoods State Park
3Lost Monarch97.83217.6825.298934,900Jedediah Smith Redwoods State Park
4Howland Hill Giant100.33296.0219.895133,600Jedediah Smith Redwoods State Park
5Sir Isaac Newton94.83117.0123.094033,000Prairie Creek Redwoods State Park

Calculating the volume of a standing tree is the practical equivalent of calculating thevolume of an irregular cone,[93] and is subject to error for various reasons. This is partly due to technical difficulties in measurement, and variations in the shape of trees and their trunks. Measurements of trunkcircumference are taken at only a few predetermined heights up the trunk, and assume that the trunk is circular incross-section, and that taper between measurement points is even. Also, only the volume of the trunk (including the restored volume of basal fire scars) is taken into account, and not the volume of wood in the branches or roots.[93] The volume measurements also do not take cavities into account. Most coast redwoods with volumes greater than 850 m3 (30,000 cu ft) represent ancient fusions of two or more separate trees, which makes determining whether a coast redwood has a single stem or multiple stems difficult.[94] Starting in 2014, more record-breaking coast redwood trees were discovered. The largest disclosed was a massive redwood called Grogan's Fault/Spartan,[95] which has been measured to have a volume of 38,300 cubic feet. In 2021 during a meeting presentation titled Redwoods 101 run by Henry Cowell Redwoods State Park, an even larger redwood was revealed, allegedly surpassed only by 3 giant sequoias in size.[96] This tree is popularly known as 'Hail Storm', and has a volume of 44,750 cubic feet.[97]

Details of the precise locations for most of the tallest trees were not announced to the general public for fear of causing damage to the trees and the surrounding habitat.[8] The largest coast redwood easily accessible to the public isIluvatar, which stands prominently about 5 meters (16 ft) to the southeast of the Foothill Trail of Prairie Creek Redwoods State Park.

Canopy layers

[edit]
Clouded Salamander,Aneides ferreus

Redwood canopy soil forms from leaf and organic material litter shedding from upper portions of the tree, accumulating and decomposing on larger branches.[11] These clusters of soil require a lot of hydration, but they have an incredible amount of retention once saturated. Redwoods can send roots into these wet soils, providing a water source removed from the forest floor. This creates a unique ecosystem within old growth trees full of fungi, vascular plants, and small creatures. An example of a creature that lives there are the Clouded Salamanders that has been discovered up to 40 meters high. Evidence shows they breed and are born in the canopy soil of Redwood trees.[11] Due to the sheer mass height of these trees and the canopy layer, it was almost never explored for the last century. Due to the mass of these trees and the amount of trees in the surrounding area different molds of moss form on these canopies that are called epiphytes. These epiphytes have different characteristics but all of said species are very adaptable to the tough treetop weather and characteristics.[11] After hundreds of years these trees have been shaped into making it possible for these epiphytes to survive through the winter rain and the fall fog.[11]

Other notable examples

[edit]
  • TheBlossom Rock Navigation Trees were two especially tall sequoias located in theBerkeley Hills used as anavigational aid by sailors to avoid the treacherous Blossom Rock nearYerba Buena Island.[98]
  • The Crannell Creek Giant was documented to have a trunk volume of at least 1,744 m3 (61,573 cu ft) – about 32% larger than Grogan's Fault and 17% larger thanGeneral Sherman, the current largest tree. It was felled around 1945.[8][99]
  • TheLindsay Creek Tree was documented to have a height of 120 m (390 ft) and a trunk volume of at least 2,500 cubic meters (90,000 cu ft) when it was uprooted and felled by a storm in 1905. If these measurements are to be believed, the Lindsay Creek Tree was about 3 m (10 ft) taller than Hyperion, the current tallest tree, 213% larger than Grogan's Fault, and 171% larger than General Sherman.[8][100]
  • Old Survivor, also known as the Grandfather, is the last remaining old-growth coastal redwood of the redwood forest that populated theOakland Hills. The tree was seeded sometime between 1549 and 1554.[101]
  • One of the largest redwood stumps ever found, measuring 9.4 m (31 ft) in diameter, is inOakland, California, the Berkeley Hills, in theRoberts Regional Recreation Area section ofRedwood Regional Park.[102][103]

See also

[edit]

References

[edit]
  1. ^abFarjon, A.; Schmid, R. (2013)."Sequoia sempervirens".IUCN Red List of Threatened Species.2013 e.T34051A2841558.doi:10.2305/IUCN.UK.2013-1.RLTS.T34051A2841558.en. Retrieved11 November 2021.
  2. ^NatureServe. 2020.Sequoia sempervirens, Redwood. NatureServe, Arlington, Virginia. Availablehttps://explorer.natureserve.org/Taxon/ELEMENT_GLOBAL.2.140518/Sequoia_sempervirens. Accessed 11 November 2021.
  3. ^Sunset Western Garden Book, 1995: 606–607
    "sempervirent".Oxford English Dictionary (Online ed.). Oxford University Press. (Subscription orparticipating institution membership required.)
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Further reading

[edit]
  • Noss, R. F., ed. (2000).The Redwood Forest: history, ecology and conservation of the Coast Redwood. Island Press, Washington, D.C.ISBN 1-55963-726-9.

External links

[edit]
Sequoia sempervirens at Wikipedia'ssister projects
ExtantCupressaceae species
SubfamilyAthrotaxidoideae
Athrotaxis
SubfamilyCallitroideae
Actinostrobus
Austrocedrus
Callitris
Diselma
Fitzroya
Libocedrus
Neocallitropsis
Papuacedrus
Pilgerodendron
Widdringtonia
SubfamilyCunninghamioideae
Cunninghamia
SubfamilyCupressoideae
Callitropsis
Calocedrus
Chamaecyparis
Cupressus
Hesperocyparis
Fokienia
Juniperus
SectionJuniperus
SectionSabina
Microbiota
Platycladus
Tetraclinis
Thuja
Thujopsis
Xanthocyparis
SubfamilySequoioideae
Metasequoia
Sequoia
Sequoiadendron
SubfamilyTaiwanioideae
Taiwania
SubfamilyTaxodioideae
Cryptomeria
Glyptostrobus
Taxodium
Sequoia sempervirens
Taxodium sempervirens
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