Protorthoptera Temporal range:318–299 Ma PreꞒ Ꞓ O S D C P T J K Pg N MiddleCarboniferous
Hadentomum americanum
Scientific classification
Kingdom:	Animalia
Phylum:	Arthropoda
Class:	Insecta
Cohort:	Polyneoptera
Order:	†Protorthoptera Handlirsch, 1906
Families
See text
Synonyms
Cacurgida Paraplecoptera Protoblattodea Protoblattoidea

TheProtorthoptera are an extinct order ofPalaeozoic insects, and represent awastebasket taxon andparaphyletic assemblage ofbasalneoptera^{[citation needed]}. They appear during the MiddleCarboniferous (lateSerpukhovian or earlyBashkirian), making them among the earliest known winged insects in the fossil record.Pronotal lobes may be expanded to form a shield. The group includes the ancestors of all otherpolyneopterous insects.

Protorthoptera^[1]

• †AdeloneuridaeCarpenter, 1938
  • †AdeloneuraCarpenter, 1938
• †AnthracoptilidaeHandlirsch, 1922
  • †AnthracoptilusLameere, 1917
• †AnthracothremmidaeHandlirsch, 1906
  • †AnthracothremmaScudder, 1885
  • †MelinophlebiaHandlirsch, 1911
  • †PericalypheHandlirsch, 1911
  • †SilphionHandlirsch, 1911
• †ApithanidaeHandlirsch, 1911
  • †ApithanusHandlirsch, 1911
• †AsyncritidaeHandlirsch, 1911
  • †AsyncritusHandlirsch, 1911
• †BlattinopsidaeBolton, 1925 (synonym OryctoblattinidaeHandlirsch, 1906)
  • †BlattinopsisGiebel, 1867
  • †GlaphyrokorisRichardson, 1956
  • †GlaphyrophlebiaHandlirsch, 1906
  • †ProtoblattiniellaMeunier, 1912
• †CheliphlebidaeHandlirsch, 1911
  • †CheliphlebiaScudder, 1885
• †CymbopsidaeKukalova, 1965
  • †CymbopsisKukalova, 1965
• †Epimastacidae
• †EucaenidaeHandlirsch, 1906 (synonym Teneopteridae)
  • †EucaenusScudder, 1885
• †HadentomidaeHandlirsch, 1906 (synonym HadentomoideaHandlirsch, 1906)
  • †Fabreciella
  • †Fayoliella
  • †HadentomumHandlirsch, 1906
• †HapalopteridaeHandlirsch, 1906 (synonym AenigmatodidaeHandlirsch, 1906 & HapalopteroideaHandlirsch, 1906)
  • †AenigmatodesHandlirsch, 1906
  • †HapalopteraHandlirsch, 1906
• †HeteroptilidaeCarpenter, 1977
  • †HeteroptilonCarpenter, 1977
• †HomalophlebiidaeHandlirsch, 1906
  • †HomalophlebiaBrongniart, 1893
  • †ParahomalophlebiaHandlirsch, 1906
• †Homoeodictyidae
  • †HomoeodictyonCarpenter, 1992
• †Pachytylopsidae
  • †PachytylopsisBorre, 1875
  • †ProtopachytylopsisLaurentiaux & Laurentiaux-Vieira, 1981
  • †SymballophlebiaHandlirsch, 1904
• †Stenoneuridae
  • †EoblattinaBolton, 1925
  • †StenoneuraBrongniart, 1893
• †StereopteridaeCarpenter, 1950
  • †StereopterumCarpenter, 1950
• †StrephocladidaeMartynov, 1938
  • †HomocladusCarpenter, 1966
  • †ParacladusCarpenter, 1966
  • †StrephocladusScudder, 1885
• †StygnidaeHandlirsch, 1906
  • †StygneHandlirsch, 1906
• †Thoronysididae
  • †ThoronysisCarpenter, 1992
• †Incertae sedis
  • †AcriditesGermar, 1842
  • †AdiphlebiaScudder, 1885
  • †AgogoblattinaHandlirsch, 1906
  • †AnthrakorisRichardson, 1956
  • †ArchaeologusHandlirsch, 1906
  • †ArchimastaxHandlirsch, 1906
  • †AtavaSellards, 1909
  • †AxiologusHandlirsch, 1906
  • †ChrestotesScudder, 1868
  • †CommentryaLameere, 1917
  • †DidymophlepsScudder, 1885
  • †EndoiasmusHandlirsch, 1906
  • †GeraroidesHandlirsch, 1906
  • †GyrophlebiaHandlirsch, 1906
  • †HemeristaDana, 1864
  • †LecopterumSellards, 1909
  • †MegalometerHandlirsch, 1906
  • †MetacheliphlebiaHandlirsch, 1906
  • †MiamiaDana, 1864
  • †PalaeocixiusBrongniart, 1893
  • †PolyetesHandlirsch, 1906
  • †PruvostiaBolton, 1921
  • †PseudogerarusHandlirsch, 1906
  • †PseudopolyernusHandlirsch, 1906
  • †PtenoderaBolton, 1922
  • †SchuchertiellaHandlirsch, 1911

• Carpenter, F. M. 1992. Superclass Hexapoda. Volume 3 of Part R, Arthropoda 4;Treatise on Invertebrate Paleontology, Boulder, Colorado, Geological Society of America.
• Grimaldi, David & Engel, Michael S. (2005-05-16).Evolution of the Insects.Cambridge University Press.ISBN 0-521-82149-5.

• Carboniferous insects
• Extinct insect orders
• Taxa named by Anton Handlirsch
• Paraphyletic groups
• Polyneoptera

• Articles with short description
• Short description matches Wikidata
• Articles with 'species' microformats
• All articles with unsourced statements
• Articles with unsourced statements from October 2021