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Protathlitis

From Wikipedia, the free encyclopedia
Genus of spinosaurid theropod dinosaur

Protathlitis
Temporal range:Early Cretaceous,Barremian
129.4–125 Ma
Skeletal diagram of the relatedBaryonyx, showing the known bones ofProtathlitis in red
Scientific classificationEdit this classification
Domain:Eukaryota
Kingdom:Animalia
Phylum:Chordata
Clade:Dinosauria
Clade:Saurischia
Clade:Theropoda
Family:Spinosauridae
Subfamily:Baryonychinae
Genus:Protathlitis
Santos-Cubedoet al.,2023
Type species
Protathlitis cinctorrensis
Santos-Cubedoet al., 2023

Protathlitis (meaning "champion") is a genus ofspinosauriddinosaur from theEarly Cretaceous (Barremian)Arcillas de Morella Formation ofCastellón, Spain. Thetype andonly species isProtathlitis cinctorrensis, known from a partial skeleton. It was abasal member of theBaryonychinae.[1] Its discovery, as well as those of the spinosauridsCamarillasaurus,Iberospinus,Riojavenatrix, and the contemporaryVallibonavenatrix shows that the Iberian Peninsula held a diverse assemblage of spinosaurids during the Early Cretaceous.[2]

Discovery and naming

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Theholotype remains, themaxillary fragment 8ANA-109 andcaudal vertebrae 3ANA83, 4ANA43, 4ANA69, 4ANA76, and 5ANA78, were recovered from the ANA site of the Arcillas de Morella Formation, which was discovered in 1998 and remained unexplored until 2002. A tooth, 4ANA-11, possibly from the left mandible or right maxilla, was also referred.

The remains weredescribed as a new genus and species of spinosaurid in 2023,Protathlitis cinctorrensis. The genus name, "Protathlitis", is Greek for "champion", and is dedicated to the2020–21 UEFA Europa League won byVillareal C.F. and in light of the club's centenary in 2023. The specific name, "cinctorrensis", honorsCinctorres, the town where the remains were discovered.[1]

In 2024, Montealegre, Castillo-Visa & Sellés tentatively assigned specimen IPS919, a nearly complete tooth including a partialroot from the Arcillas de Morella Formation, tocf.Protathlitis.[3]

Classification

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Santos-Cubedoet al. (2023) performed aphylogenetic analysis, placingProtathlitis as the basalmost member the Baryonychinae. Their results are shown in thecladogram below:[1]

Speculative life reconstruction
Spinosauridae

Palaeoenvironment

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Protathlitis hails from theArcillas de Morella Formation, which has been dated to theBarremianstage of theEarly Cretaceousperiod, between 129.4 and 125 million years ago. It coexisted in this environment with other dinosaurs including theornithischiansIguanodon bernissartensis andMorelladon beltrani, an indeterminatesauropod, and the fellow spinosauridVallibonavenatrix.[4][5][6]

References

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  1. ^abcSantos-Cubedo, A.; de Santisteban, C.; Poza, B.; Meseguer, S. (2023)."A new spinosaurid dinosaur species from the Early Cretaceous of Cinctorres (Spain)".Scientific Reports.13 (1). 6471.doi:10.1038/s41598-023-33418-2.hdl:10234/203142.
  2. ^Isasmendi, E.; Cuesta, E.; Díaz-Martínez, I.; Company, J.; Sáez-Benito, P.; Viera, L. I.; Torices, A.; Pereda-Suberbiola, P. (2024)."Increasing the theropod record of Europe: a new basal spinosaurid from the Enciso Group of the Cameros Basin (La Rioja, Spain). Evolutionary implications and palaeobiodiversity".Zoological Journal of the Linnean Society.doi:10.1093/zoolinnean/zlad193.{{cite journal}}:|last4= has generic name (help)
  3. ^Montealegre, Adrian; Castillo-Visa, Oscar; Sellés, Albert (2024-01-31). "New theropod remains from the late Barremian (Early Cretaceous) of Eastern Iberian Peninsula".Historical Biology:1–11.doi:10.1080/08912963.2024.2308220.ISSN 0891-2963.
  4. ^Weishampel, David B; et al. (2004). "Dinosaur distribution (Early Cretaceous, Europe)." In: Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. Pp. 562.ISBN 0-520-24209-2.
  5. ^Verdú, F.J.; Godefroit, P.; Royo-Torres, R.; Cobos, A.; Alcalá, L. (2017). "Individual variation in the postcranial skeleton of the Early CretaceousIguanodon bernissartensis (Dinosauria: Ornithopoda)".Cretaceous Research.74:65–86.doi:10.1016/j.cretres.2017.02.006.
  6. ^Malafaia, E.; Miguel Gasulla, J.; Escaso, F.; Narváez, I.; Luis Sanz, J.; Ortega, F. (2019). "A new spinosaurid theropod (Dinosauria: Megalosauroidea) from the late Barremian of Vallibona, Spain: Implications for spinosaurid diversity in the Early Cretaceous of the Iberian Peninsula".Cretaceous Research.106: 104221.doi:10.1016/j.cretres.2019.104221.S2CID 202189246.
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