Thehouse sparrow (Passer domesticus) is abird of thesparrow family Passeridae, found in most parts of the world. It is a small bird that has a typical length of 16 cm (6.3 in) and a mass of 24–39.5 g (0.85–1.39 oz). Females and young birds are coloured pale brown and grey, and males have brighter black, white, and brown markings. One of about 25 species in the genusPasser, the house sparrow is native to most of Europe, theMediterranean Basin, and a large part of Asia. Its intentional or accidentalintroductions to many regions, including parts of Australasia, Africa, and the Americas, make it the most widely distributed wild bird.
The house sparrow is strongly associated with human habitation, and can live in urban or rural settings. Though found in widely varied habitats and climates, it typically avoids extensivewoodlands,grasslands, polar regions, and hot, drydeserts far away from human development. For sustenance, the house sparrow routinely feeds at home and public bird feeding stations, but naturally feeds on the seeds of grains, flowering plants and weeds. However, it is an opportunistic, omnivorous eater, and commonly catches insects, their larvae, caterpillars, invertebrates and many other natural foods.
Because of its numbers, ubiquity, and association with human settlements, the house sparrow is culturally prominent. It is extensively, and usually unsuccessfully, persecuted as an agricultural pest. It has also often been kept as a pet, as well as being a food item and a symbol of lust, sexual potency, commonness, and vulgarity. Though it is widespread and abundant, its numbers have declined in some areas.[3] The animal'sconservation status is listed asleast concern on theIUCN Red List.
The house sparrow is typically about 16 cm (6.3 in) long, ranging from 14 to 18 cm (5.5 to 7.1 in).[4] The house sparrow is a compact bird with a full chest and a large, rounded head. Its bill is stout and conical with aculmen length of 1.1–1.5 cm (0.43–0.59 in), strongly built as an adaptation for eating seeds. Its tail is short, at 5.2–6.5 cm (2.0–2.6 in) long. Thewing chord is 6.7–8.9 cm (2.6–3.5 in), and thetarsus is 1.6–2.5 cm (0.63–0.98 in).[5][6] Wingspan ranges from 19–25 centimetres (7.5–9.8 in).[5]
In mass, the house sparrow ranges from 24 to 39.5 g (0.85 to 1.39 oz). Females usually are slightly smaller than males. The median mass on the European continent for both sexes is about 30 g (1.1 oz), and in more southerly subspecies is around 26 g (0.92 oz). Younger birds are smaller, males are larger during the winter, and females are larger during the breeding season. Birds at higher latitudes, colder climates, and sometimes higher altitudes are larger (underBergmann's rule), both between and withinsubspecies.[7][8][9][10]
Theplumage of the house sparrow is mostly different shades of grey and brown. The sexes exhibit strongdimorphism: the female is mostlybuffish above and below, while the male has boldly coloured head markings, a reddish back, and grey underparts.[9] The male has a dark greycrown from the top of its bill to its back, and chestnut brown flanking its crown on the sides of its head. It has black around its bill, on its throat, and on the spaces between its bill and eyes (lores). It has a small white stripe between the lores and crown and small white spots immediately behind the eyes (postoculars), with black patches below and above them. The underparts are pale grey or white, as are the cheeks, earcoverts, and stripes at the base of the head. The upper back and mantle are a warm brown, with broad black streaks, while the lower back, rump and upper tail coverts are greyish brown.[11]
The male is duller in fresh nonbreeding plumage, with whitish tips on many feathers. Wear and preening expose many of the bright brown and black markings, including most of the black throat and chest patch, called the "bib" or "badge".[11][12] The badge is variable in width and general size, and may signal social status or fitness. This hypothesis has led to a "veritable 'cottage industry'" of studies, which have only conclusively shown that patches increase in size with age.[13] The male's bill is dark grey, but black in the breeding season.[4]
Heads of a male (left) and an immature or female (right)
The female has no black markings or grey crown. Its upperparts and head are brown with darker streaks around the mantle and a distinct palesupercilium. Its underparts are pale grey-brown. The female's bill is brownish-grey and becomes darker in breeding plumage approaching the black of the male's bill.[4][11]
Juveniles are similar to the adult female, but deeper brown below and paler above, with paler and less defined supercilia. Juveniles have broader buff feather edges, and tend to have looser, scruffier plumage, like moulting adults. Juvenile males tend to have darker throats and white postoculars like adult males, while juvenile females tend to have white throats. However, juveniles cannot be reliably sexed by plumage: some juvenile males lack any markings of the adult male, and some juvenile females have male features. The bills of young birds are light yellow tostraw, paler than the female's bill. Immature males have paler versions of the adult male's markings, which can be very indistinct in fresh plumage. By their first breeding season, young birds generally are indistinguishable from other adults, though they may still be paler during their first year.[4][11]
Most house sparrowvocalisations are variations on its short and frequent chirping call. Transcribed aschirrup,tschilp, orphilip, this note is made as a contact call by flocking or resting birds; or by males to proclaim nest ownership and invite pairing. In the breeding season, the male gives this call repetitively, with emphasis and speed, but not much rhythm, forming what is described either as a song or an "ecstatic call" similar to a song.[14][15] Young birds also give a true song, especially in captivity, a warbling similar to that of theEuropean greenfinch.[16]
Aggressive males give a trilled version of their call, transcribed as "chur-chur-r-r-it-it-it-it". This call is also used by females in the breeding season, to establish dominance over males while displacing them to feed young or incubate eggs.[17] House sparrows give a nasal alarm call, the basic sound of which is transcribed asquer, and a shrillchree call in great distress.[18] Another vocalisation is the "appeasement call", a softquee given to inhibit aggression, usually given between birds of a mated pair.[17] These vocalisations are not unique to the house sparrow, but are shared, with small variations, by all sparrows.[19]
An immature of the Indian subspecies (P. d. indicus) in Rajasthan, India
Some variation is seen in the 12 subspecies of house sparrows, which are divided into two groups, theOrientalP. d. indicus group, and thePalaearcticP. d. domesticus group. Birds of theP. d. domesticus group have grey cheeks, whileP. d. indicus group birds have white cheeks, as well as bright colouration on the crown, a smaller bill, and a longer black bib.[20] The subspeciesP. d. tingitanus differs little from the nominate subspecies, except in the worn breeding plumage of the male, in which the head is speckled with black and underparts are paler.[21]P. d. balearoibericus is slightly paler than the nominate, but darker thanP. d. bibilicus.[22]P. d. bibilicus is paler than most subspecies, but has the grey cheeks ofP. d. domesticus group birds. The similarP. d. persicus is paler and smaller, andP. d. niloticus is nearly identical but smaller. Of the less widespreadP. d. indicus group subspecies,P. d. hyrcanus is larger thanP. d. indicus,P. d. hufufae is paler,P. d. bactrianus is larger and paler, andP. d. parkini is larger and darker with more black on the breast than any other subspecies.[21][23][24]
The house sparrow can be confused with a number of other seed-eating birds, especially its relatives in the genusPasser. Many of these relatives are smaller, with an appearance that is neater or "cuter", as with theDead Sea sparrow.[25] The light brown-coloured female can often not be distinguished from other females, and is nearly identical to those of theSpanish andItalian sparrows.[11] TheEurasian tree sparrow is smaller and slenderer with a chestnut crown and a black patch on each cheek.[26] The male Spanish sparrow and Italian sparrow are distinguished by their chestnut crowns. TheSind sparrow is very similar but smaller, with less black on the male's throat and a distinct pale supercilium on the female.[11]
The bird's scientific name and its usual English name have the same meaning. TheLatin wordpasser, like the English word "sparrow", is a term for small active birds, coming from a root word referring to speed.[31][32] The Latin worddomesticus means "belonging to the house", like the common name a reference to its association with humans.[33] The house sparrow is also called by a number of alternative English names, includingEnglish sparrow, chiefly in North America;[34][35] andIndian sparrow orIndian house sparrow, for the birds of theIndian subcontinent andCentral Asia.[36] Dialectal names includesparr,sparrer,spadger,spadgick, andphilip, mainly in southern England;spug andspuggy, mainly in northern England;spur andsprig, mainly inScotland;[37][38] andspatzie orspotsie, from the GermanSpatz, in North America.[39]
The genusPasser contains about 25 species, depending on the authority, 26 according to theHandbook of the Birds of the World.[40] MostPasser species are dull-coloured birds with short, square tails and stubby, conical beaks, between 11 and 18 cm (4.3 and 7.1 in) long.[9][41]Mitochondrial DNA studies suggest thatspeciation in the genus occurred during thePleistocene and earlier, while other evidence suggests speciation occurred 25,000 to 15,000 years ago. WithinPasser, the house sparrow is part of the "Palaearctic black-bibbed sparrows" group and a close relative of the Mediterranean "willow sparrows".[40][42]
The taxonomy of the house sparrow and its Mediterranean relatives is complicated. The common type of "willow sparrow" is the Spanish sparrow, which resembles the house sparrow in many respects.[43] It frequently prefers wetter habitats than the house sparrow, and it is often colonial and nomadic.[44] In most of the Mediterranean, one or both species occur, with some degree ofhybridisation.[45] In North Africa, the two species hybridise extensively, forming highly variable mixed populations with a full range of characters from pure house sparrows to pure Spanish sparrows.[46][47][48]
In most of Italy, the breeding species is theItalian sparrow, which has an appearance intermediate between those of the house and Spanish sparrows. Its specific status and origin are the subject of much debate, but it may be a case of long-agohybrid speciation.[47][49] In theAlps, the Italian sparrowintergrades over a narrow roughly 20 km (12 mi) strip with the house sparrow, and some house sparrows migrate into the Italian sparrow's range in winter.[50] On the Mediterranean islands ofMalta,Gozo,Crete,Rhodes, andKarpathos, other apparently intermediate birds are of unknown status.[47][51][52]
A male of the subspeciesP. d. balearoibericus inIstanbulA male of the migratory subspeciesP. d. bactrianus (with aEurasian tree sparrow and young house or Spanish sparrows) inBaikonur, Kazakhstan
A large number of subspecies have been named, of which 12 were recognised in theHandbook of the Birds of the World. These subspecies are divided into two groups, the PalaearcticP. d. domesticus group, and the OrientalP. d. indicus group.[40] Several Middle Eastern subspecies, includingP. d. biblicus, are sometimes considered a third, intermediate group. The subspeciesP. d. indicus was described as a species, and was considered to be distinct by many ornithologists during the 19th century.[20]
Migratory birds of the subspeciesP. d. bactrianus in theP. d. indicus group were recorded overlapping withP. d. domesticus birds without hybridising in the 1970s, so the Soviet scientists Edward I. Gavrilov and M. N. Korelov proposed the separation of theP. d. indicus group as a separate species.[29][53] However,P. d. indicus group andP. d. domesticus group birds intergrade in a large part ofIran, so this split is rarely recognised.[40]
In North America, house sparrow populations are more differentiated than those in Europe.[8] This variation follows predictable patterns, with birds at higher latitudes being larger and darker and those in arid areas being smaller and paler.[9][54][55] However, how much this is caused by evolution or by environment is not clear.[56][57][58][59] Similar observations have been made in New Zealand[60] and in South Africa.[61] The introduced house sparrow populations may be distinct enough to merit subspecies status, especially in North America and southern Africa,[40] and American ornithologistHarry Church Oberholser even gave the subspecies nameP. d. plecticus to the paler birds of western North America.[54]
P. d. domesticus group
P. d. domesticusLinnaeus, 1758, thenominate subspecies, is found in most of Europe, across northern Asia toSakhalin andKamchatka. It is the most widely introduced subspecies.[27]
P. d. tingitanus(Loche, 1867), described fromAlgeria, is found in theMaghreb fromAjdabiya inLibya toBéni Abbès in Algeria, and toMorocco's Atlantic coast. It hybridises extensively with the Spanish sparrow, especially in the eastern part of its range.[62]
P. d. niloticusNicoll and Bonhote, 1909, described fromFaiyum, Egypt, is found along theNile north ofWadi Halfa, Sudan. It intergrades withbibilicus in theSinai, and withrufidorsalis in a narrow zone aroundWadi Halfa. It has been recorded inSomaliland.[62][63]
P. d. persicusZarudny and Kudashev, 1916, described from theKarun River inKhuzestan, Iran, is found in the western and central Iran south of theAlborz mountains, intergrading withindicus in eastern Iran, and Afghanistan.[40][62][64]
P. d. biblicusHartert, 1910, described fromPalestine, is found in the Middle East fromCyprus and southeastern Turkey to the Sinai in the west and fromAzerbaijan toKuwait in the east.[40][62]
P. d. indicus group
P. d. hyrcanusZarudny and Kudashev, 1916, described fromGorgan, Iran, is found along the southern coast of theCaspian Sea from Gorgan to southeastern Azerbaijan. It intergrades withP. d. persicus in the Alborz mountains, and withP. d. bibilicus to the west. It is the subspecies with the smallest range.[40][62]
P. d. bactrianusZarudny and Kudashev, 1916, described fromTashkent, is found in southern Kazakhstan to theTian Shan and northern Iran andAfghanistan. It intergrades withpersicus inBaluchistan and withindicus across central Afghanistan. Unlike most other house sparrow subspecies, it is almost entirely migratory, wintering in the plains of the northern Indian subcontinent. It is found in open country rather than in settlements, which are occupied by the Eurasian tree sparrow in its range.[40][62] There is an exceptional record from Sudan.[63]
By a nest in asaguaro cactus inArizona, U.S.House sparrows perching on a roof, during winter in theSouthern Alps of New Zealand
The house sparrow originated in theMiddle East and spread, along with agriculture, to most of Eurasia and parts of North Africa.[67] Since the mid-19th century, it has reached most of the world, chiefly due to deliberate introductions, but also through natural and shipborne dispersal.[68] Its introduced range encompasses most ofNorth America (includingBermuda),[69]Central America, southern South America,southern Africa, part ofWest Africa,Australia,New Zealand, and islands throughout the world.[70] It has greatly extended its range in northern Eurasia since the 1850s,[71] and continues to do so, as was shown by its colonisation around 1990 ofIceland andRishiri Island, Japan.[72] The extent of its range makes it the most widely distributed wild bird on the planet.[70]
The house sparrow has become highly successful in most parts of the world where it has been introduced. This is mostly due to its early adaptation to living with humans, and its adaptability to a wide range of conditions.[73][74] Other factors may include its robust immune response, compared to theEurasian tree sparrow.[75] Where introduced, it can extend its range quickly, sometimes at a rate over 230 km (140 mi) per year.[76] In many parts of the world, it has been characterised as a pest, and poses a threat to native birds.[77][78] A few introductions have died out or been of limited success, such as those toGreenland andCape Verde.[79]
The first of many successful introductions to North America occurred when birds from England were released in New York City, in 1852,[80][81] intended to control the ravages of thelinden moth.[82] In North America, the house sparrow now occurs from theNorthwest Territories of Canada to southernPanama,[5] and it is one of the most abundant birds of the continent.[77] The house sparrow was first introduced to Australia in 1863 atMelbourne and is common throughout the eastern part of the continent as far north asCape York,[79] but has been prevented from establishing itself inWestern Australia,[83] where every house sparrow found in the state is killed.[84] House sparrows were introduced in New Zealand in 1859, and from there reached many of the Pacific islands, includingHawaii.[85]
In southern Africa, birds of both the European subspecies (P. d. domesticus) and the Indian subspecies (P. d. indicus) were introduced around 1900. Birds ofP. d. domesticus ancestry are confined to a few towns, whileP. d. indicus birds have spread rapidly, reachingTanzania in the 1980s. Despite this rapid spread, native relatives such as theCape sparrow also occur and thrive in urban habitats.[79][86] In South America, it was first introduced nearBuenos Aires around 1870, and quickly became common in most of the southern part of the continent. It now occurs almost continuously fromTierra del Fuego to the fringes of theAmazon basin, with isolated populations as far north as coastal Venezuela.[79][87][88]
The house sparrow is closely associated with human habitation and cultivation.[89] It is not an obligatecommensal of humans as some have suggested: birds of the migratory Central Asian subspecies usually breed away from humans in open country,[90] and birds elsewhere are occasionally found away from humans.[89][91][92] The only terrestrial habitats that the house sparrow does not inhabit are dense forest andtundra. Well adapted to living around humans, it frequently lives and even breeds indoors, especially in factories, warehouses, and zoos.[89] It has been recorded breeding in an Englishcoal mine 640 m (2,100 ft) below ground,[93] and feeding on theEmpire State Building's observation deck at night.[94] It reaches its greatest densities in urban centres, but its reproductive success is greater in suburbs, where insects are more abundant.[89][95] On a larger scale, it is most abundant in wheat-growing areas such as theMidwestern United States.[96]
It tolerates a variety of climates, but prefers drier conditions, especially in moist tropical climates.[79][89] It has several adaptations to dry areas, including a high salt tolerance[97] and an ability to survive without water by ingestingberries.[98] In most of eastern Asia, the house sparrow is entirely absent, replaced by the Eurasian tree sparrow.[99] Where these two species overlap, the house sparrow is usually more common than the Eurasian tree sparrow, but one species may replace the other in a manner that ornithologistMaud Doria Haviland described as "random, or even capricious".[100] In most of its range, the house sparrow is extremely common, despite some declines,[1] but in marginal habitats such as rainforest or mountain ranges, its distribution can be spotty.[89]
The house sparrow is a very social bird. It is gregarious during all seasons when feeding, often forming flocks with other species of birds.[101] Itroosts communally while breeding nests are usually grouped together in clumps. House sparrows also engage in social activities such asdust or water bathing and "social singing", in which birds call together in bushes.[102][103] The house sparrow feeds mostly on the ground, but it flocks in trees and bushes.[102] At feeding stations and nests, female house sparrows are dominant despite their smaller size, and they can fight over males in the breeding season.[104][105]
House sparrows sleep with the bill tucked underneath thescapular feathers.[106] Outside of the reproductive season, they often roost communally in trees or shrubs. Much communal chirping occurs before and after the birds settle in the roost in the evening, as well as before the birds leave the roost in the morning.[102] Some congregating sites separate from the roost may be visited by the birds prior to settling in for the night.[107]
As an adult, the house sparrow mostly feeds on the seeds ofgrains andweeds, but it is opportunistic and adaptable, and eats whatever foods are available.[109] In towns and cities, it often scavenges for food in garbage containers and congregates in the outdoors of restaurants and other eating establishments to feed on leftover food and crumbs. It can perform complex tasks to obtain food, such as opening automatic doors to enter supermarkets,[110] clinging to hotel walls to watch vacationers on their balconies,[111] andnectar robbingkowhai flowers.[112] In common with many other birds, the house sparrow requiresgrit to digest the harder items in its diet. Grit can be either stone, often grains of masonry, or the shells of eggs orsnails; oblong and rough grains are preferred.[113][114]
Several studies of the house sparrow in temperate agricultural areas have found the proportion of seeds in its diet to be about 90%.[109][115][116] It will eat almost any seeds, but where it has a choice, it preferscorn:oats,wheat ormaize.[117] Rural birds tend to eat more waste seed from animal dung and seed from fields while urban birds tend to eat more commercial bird seed and weed seed.[118] In urban areas, the house sparrow also feeds largely on food provided directly or indirectly by humans, such as bread, though it prefers raw seeds.[116][119] The house sparrow also eats some plant matter besides seeds, includingbuds, berries, and fruits such as grapes and cherries.[98][116] In temperate areas, the house sparrow has an unusual habit of tearing flowers, especially yellow ones, in the spring.[120]
Animals form another important part of the house sparrow's diet, chieflyinsects, of whichbeetles,caterpillars,dipteran flies, andaphids are especially important. Various noninsectarthropods are eaten, as aremolluscs andcrustaceans where available,earthworms, and evenvertebrates such aslizards andfrogs.[109] Young house sparrows are fed mostly on insects until about 15 days after hatching.[121] They are also given small quantities of seeds, spiders, and grit. In most places,grasshoppers andcrickets are the most abundant foods of nestlings.[122]True bugs,ants,sawflies, and beetles are also important, but house sparrows take advantage of whatever foods are abundant to feed their young.[122][123][124] House sparrows have been observedstealing prey from other birds, includingAmerican robins.[5]
The gut microbiota of house sparrows differs between chicks and adults, withPseudomonadota (formerly Proteobacteria) decreasing in chicks when they get to around 9 days old, whilst the relative abundance ofBacillota increase.[125]
The house sparrow's flight is direct (not undulating) and flapping, averaging 45.5 km/h (28.3 mph) and about 15 wingbeats per second.[107][126] On the ground, the house sparrow typically hops rather than walks. It can swim when pressed to do so by pursuit from predators. Captive birds have been recorded diving and swimming short distances under water.[107]
Most house sparrows do not move more than a few kilometres during their lifetimes. However, limited migration occurs in all regions. Some young birds disperse long distances, especially on coasts, and mountain birds move to lower elevations in winter.[102][127][128] Two subspecies,P. d. bactrianus andP. d. parkini, are predominantlymigratory. Unlike the birds in sedentary populations that migrate, birds of migratory subspecies prepare for migration by putting on weight.[102]
A pair of the Indian subspecies (P. d. indicus) mating inKolkata
House sparrows can breed in the breeding season immediately following their hatching, and sometimes attempt to do so. Some birds breeding for the first time in tropical areas are only a few months old and still have juvenile plumage.[129] Birds breeding for the first time are rarely successful in raising young, and reproductive success increases with age, as older birds breed earlier in the breeding season, and fledge more young.[130] As the breeding season approaches, hormone releases trigger enormous increases in the size of thesexual organs and changes in day length lead males to start calling by nesting sites.[131][132] The timing of mating and egg-laying varies geographically, and between specific locations and years because a sufficient supply of insects is needed for egg formation and feeding nestlings.[133]
Males take up nesting sites before the breeding season, by frequently calling beside them. Unmated males start nest construction and call particularly frequently to attract females. When a female approaches a male during this period, the maledisplays by moving up and down while drooping and shivering his wings, pushing up his head, raising and spreading his tail, and showing his bib.[133] Males may try to mate with females while calling or displaying. In response, a female will adopt a threatening posture and attack a male before flying away, pursued by the male. The male displays in front of her, attracting other males, which also pursue and display to the female. This group display usually does not immediately result in copulations.[133] Other males usually do not copulate with the female.[134][135] Copulation is typically initiated by the female giving a softdee-dee-dee call to the male. Birds of a paircopulate frequently until the female is laying eggs, and the male mounts the female repeatedly each time a pair mates.[133]
The house sparrow ismonogamous, and typically mates for life, but birds from pairs often engage inextra-pair copulations, so about 15% of house sparrow fledglings are unrelated to their mother's mate.[136] Males guard their mates carefully to avoid being cuckolded, and most extra-pair copulation occurs away from nest sites.[134][137] Males may sometimes have multiple mates, and bigamy is mostly limited by aggression between females.[138] Many birds do not find a nest and a mate, and instead may serve ashelpers around the nest for mated pairs, a role which increases the chances of being chosen to replace a lost mate. Lost mates of both sexes can be replaced quickly during the breeding season.[134][139] The formation of a pair and the bond between the two birds is tied to the holding of a nest site, though paired house sparrows can recognise each other away from the nest.[133]
House sparrows in natural small populations, as can occur on islands, exhibitinbreeding depression.[140][141] Inbreeding depression is manifested as lower survival probability and production of fewer offspring, and can occur as a result of the expression of deleterious recessivealleles.[141] However sparrows in such populations do not appear toavoid inbreeding.[140]
Female bringing food for young in a nest made in a tree hole inCaliforniaSparrow in a ventilator
Nest sites are varied, though cavities are preferred. Nests are most frequently built in theeaves and other crevices of houses. Holes in cliffs and banks, andtree hollows, are also used.[142][143] A sparrow sometimes excavates its own nests in sandy banks or rotten branches, but more frequently uses the nests of other birds such as those ofswallows in banks and cliffs, and old tree cavity nests. It usually uses deserted nests, though sometimes it usurps active ones by driving away or killing the occupants.[142][144] Tree hollows are more commonly used in North America than in Europe,[142] putting the sparrows in competition withbluebirds and other North American cavity nesters, and thereby contributing to their population declines.[77]
Especially in warmer areas, the house sparrow may build its nests in the open, on the branches of trees, especially evergreens and hawthorns, or in the nests of large birds such asstorks ormagpies.[133][142][145] In open nesting sites, breeding success tends to be lower, since breeding begins late and the nest can easily be destroyed or damaged by storms.[142][146] Less common nesting sites includestreet lights andneon signs, favoured for their warmth; and the old open-topped nests of other songbirds, which are then domed over.[142][143]Usually the couples repeat copulation many times. Every copulation is followed by some break of 3 to 4 seconds, and in that time both pair change their position by some distance.The nest is usually domed, though it may lack a roof in enclosed sites.[142] It has an outer layer of stems and roots, a middle layer of dead grass and leaves, and a lining of feathers, as well as of paper and other soft materials.[143] Nests typically have external dimensions of 20 × 30 cm (8 × 12 in),[133] but their size varies greatly.[143] The building of the nest is initiated by the unmated male while displaying to females. The female assists in building, but is less active than the male.[142] Some nest building occurs throughout the year, especially after moult in autumn. In colder areas house sparrows build specially created roost nests, or roost in street lights, to avoid losing heat during the winter.[142][147] House sparrows do not hold territories, but they defend their nests aggressively against intruders of the same sex.[142]
Clutches usually comprise four or fiveeggs, though numbers from one to 10 have been recorded. At least two clutches are usually laid, and up to seven a year may be laid in the tropics or four a year in temperate latitudes. When fewer clutches are laid in a year, especially at higher latitudes, the number of eggs per clutch is greater. Central Asian house sparrows, which migrate and have only one clutch a year, average 6.5 eggs in a clutch. Clutch size is also affected by environmental and seasonal conditions, female age, and breeding density.[151][152]
Some intraspecificbrood parasitism occurs, and instances of unusually large numbers of eggs in a nest may be the result of females laying eggs in the nests of their neighbours. Such foreign eggs are sometimes recognised and ejected by females.[151][153] The house sparrow is a victim of interspecific brood parasites, but only rarely, since it usually uses nests in holes too small for parasites to enter, and it feeds its young foods unsuitable for young parasites.[154][155] In turn, the house sparrow has once been recorded as a brood parasite of theAmerican cliff swallow.[153][156]
A juvenile, showing its pink bill and obvious nestlinggape—the soft, swollen base, which becomes harder and less swollen as the bird matures
The eggs are white, bluish white, or greenish white, spotted with brown or grey.[107] Subelliptical in shape,[9] they range from 20 to 22 mm (0.79 to 0.87 in) in length and 14 to 16 mm (0.55 to 0.63 in) in width,[5] have an average mass of 2.9 g (0.10 oz),[157] and an average surface area of 9.18 cm2 (1.423 in2).[158] Eggs from the tropical subspecies are distinctly smaller.[159][160] Eggs begin to develop with the deposition ofyolk in the ovary a few days before ovulation. In the day between ovulation and laying,egg white forms, followed byeggshell.[161] Eggs laid later in a clutch are larger, as are those laid by larger females, and egg size is hereditary. Eggs decrease slightly in size from laying to hatching.[162] The yolk comprises 25% of the egg, the egg white 68%, and the shell 7%. Eggs are watery, being 79% liquid, and otherwise mostly protein.[163]
The female develops abrood patch of bare skin and plays the main part inincubating the eggs. The male helps, but can only cover the eggs rather than truly incubate them. The female spends the night incubating during this period, while the male roosts near the nest.[151] Eggs hatch at the same time, after a short incubation period lasting 11–14 days, and exceptionally for as many as 17 or as few as 9.[9][133][164] The length of the incubation period decreases as ambient temperature increases later in the breeding season.[165]
Young house sparrows remain in the nest for 11 to 23 days, normally 14 to 16 days.[107][165][166] During this time, they are fed by both parents. As newly hatched house sparrows do not have sufficient insulation, they are brooded for a few days, or longer in cold conditions.[165][167] The parents swallow the droppings produced by the hatchlings during the first few days; later, the droppings are moved up to 20 m (66 ft) away from the nest.[167][168]
The chicks' eyes open after about 4 days and, at an age of about 8 days, the young birds get their firstdown.[107][166] If both parents perish, the ensuing intensive begging sounds of the young often attract replacement parents which feed them until they can sustain themselves.[167][169] All the young in the nest leave it during the same period of a few hours. At this stage, they are normally able to fly. They start feeding themselves partly after 1 or 2 days, and sustain themselves completely after 7 to 10 days, 14 at the latest.[170]
House sparrows in natural small populations, as can occur on islands, exhibitinbreeding depression.[140][141] Inbreeding depression is manifested as lower survival probability and production of fewer offspring, and can occur as a result of the expression of deleterious recessivealleles.[141] However sparrows in such populations do not appear toavoid inbreeding.[140]
In adult house sparrows, annual survival is 45–65%.[171] After fledging and leaving the care of their parents, young sparrows have a high mortality rate, which lessens as they grow older and more experienced. Only about 20–25% of birds hatched survive to their first breeding season.[172] The oldest known wild house sparrow lived for nearly two decades; it was found dead 19 years and 9 months after it wasringed in Denmark.[173] The oldest recorded captive house sparrow lived for 23 years.[174] The typical ratio of males to females in a population is uncertain due to problems in collecting data, but a very slight preponderance of males at all ages is usual.[175]
A male sparrow being eaten by adomestic cat. The cats are one of the main predators of the house sparrow.
The house sparrow's main predators arecats andbirds of prey, but many other animals prey on them, includingcorvids,squirrels,[176] and even humans—the house sparrow has been consumed in the past by people in many parts of the world, and it still is in parts of the Mediterranean.[177] Most species of birds of prey have been recorded preying on the house sparrow in places where records are extensive.Accipiters and themerlin in particular are major predators, though cats are likely to have a greater impact on house sparrow populations.[176] The house sparrow is also a common victim ofroadkill; on European roads, it is the bird most frequently found dead.[178]
The house sparrow is host to a huge number of parasites and diseases, and the effect of most is unknown. Ornithologist Ted R. Anderson listed thousands, noting that his list was incomplete.[179] The commonly recorded bacterial pathogens of the house sparrow are often those common in humans, and includeSalmonella andEscherichia coli.[180]Salmonella is common in the house sparrow, and a comprehensive study of house sparrow disease found it in 13% of sparrows tested.Salmonella epidemics in the spring and winter can kill large numbers of sparrows.[179] The house sparrow hostsavian pox andavian malaria, which it has spread to the native forest birds of Hawaii.[181] Many of the diseases hosted by the house sparrow are also present in humans and domestic animals, for which the house sparrow acts as areservoir host.[182]Arboviruses such as theWest Nile virus, which most commonly infect insects and mammals, survive winters in temperate areas by going dormant in birds such as the house sparrow.[179][183] A few records indicate disease extirpating house sparrow populations, especially from Scottish islands, but this seems to be rare.[184] House sparrows are also infected byhaemosporidian parasites, but less so in urban than in rural areas[185]Toxoplasma gondii has been detected in sparrows in northwestern China where they pose a risk due to their meat being consumed in the region.[186]
The house sparrow is infested by a number of external parasites, which usually cause little harm to adult sparrows. In Europe, the most commonmite found on sparrows isProctophyllodes, the most commonticks areArgas reflexus andIxodes arboricola, and the most common flea on the house sparrow isCeratophyllus gallinae.[148]Dermanyssus blood-feeding mites are also common ectoparasites of house sparrows,[187] and these mites can enter human habitation and bite humans, causing a condition known asgamasoidosis.[188] A number ofchewing lice occupy differentniches on the house sparrow's body.Menacanthus lice occur across the house sparrow's body, where they feed on blood and feathers, whileBrueelia lice feed on feathers andPhilopterus fringillae occurs on the head.[148]
House sparrows express strongcircadian rhythms of activity in the laboratory. They were among the first bird species to be seriously studied in terms of their circadian activity andphotoperiodism, in part because of their availability and adaptability in captivity, but also because they can "find their way" and remain rhythmic in constant darkness.[189][190]
Flocking and chirping together beneath afluorescent tube light in Germany
The house sparrow is closely associated with humans. They are believed to have become associated with humans around 10,000 years ago. The Turkestan subspecies (P. d. bactrianus) is least associated with humans and considered to be evolutionarily closer to the ancestral noncommensal populations.[191] Usually, the house sparrow is regarded as a pest, since it consumes agricultural products and spreads disease to humans and their domestic animals.[192] Evenbirdwatchers often hold it in little regard because of its molestation of other birds.[77] In most of the world, the house sparrow is not protected by law. Attempts to control house sparrows include the trapping, poisoning, or shooting of adults; the destruction of their nests and eggs; or less directly, blocking nest holes and scaring off sparrows with noise, glue, or porcupine wire.[193] However, the house sparrow can be beneficial to humans, as well, especially by eating insect pests, and attempts at the large-scale control of the house sparrow have failed.[40]
The house sparrow has long been used as a food item. From around 1560 to at least the 19th century in northern Europe,earthenware "sparrow pots" were hung from eaves to attract nesting birds so the young could be readily harvested. Wild birds were trapped in nets in large numbers, and sparrow pie was a traditional dish, thought, because of the association of sparrows withlechery, to haveaphrodisiac properties.[194] A traditional Indian medicine,Ciṭṭukkuruvilēkiyam inTamil, was sold with similar aphrodisiac claims.[195] Sparrows were also trapped as food forfalconers' birds and zoo animals. During the 1870s, there were debates on the damaging effects of sparrows in the House of Commons in England.[196] In the early part of the 20th century, sparrow clubs culled many millions of birds and eggs in an attempt to control numbers of this perceived pest, but with only a localised impact on numbers.[197] Sparrows were also persecuted in Germany from at least 1650 until 1970.[198] House sparrows have been kept as pets at many times in history, though they have no bright plumage or attractive songs, and raising them is difficult.[194] The house sparrow has an extremely large range and population, so it is assessed asleast concern for conservation on theIUCN Red List.[1]
TheIUCN estimates for the global population runs up to nearly 1.4 billion individuals, second among all wild birds perhaps only to thered-billed quelea in abundance (although the quelea is, unlike the sparrow, restricted to a single continent and has never been subject to human introductions).[1] However, populations have been declining in many parts of the world, especially near its Eurasian places of origin.[199][200][201] These declines were first noticed in North America, where they were initially attributed to the spread of thehouse finch, but have been most severe inWestern Europe.[202][203] Declines have even occurred in Australia, where the house sparrow was introduced recently.[204] While no serious declines had been reported fromEastern Europe by 2006,[204] as of 2023, a 50% decrease of the sparrow population has been registered inBulgaria.[205]
InGreat Britain, populations peaked in the early 1970s,[206] but have since declined by 68% overall,[207] and about 90% in some regions.[208][209] The RSPB lists the house sparrow's UK conservation status as red.[210] In London, the house sparrow almost disappeared from the central city.[208] The numbers of house sparrows in theNetherlands have dropped in half since the 1980s,[95] so the house sparrow is even considered anendangered species.[211] This status came to widespread attention after a female house sparrow, referred to as the "dominomus", was killed after knocking down dominoes arranged as part of an attempt to set a world record.[212] These declines are not unprecedented, as similar reductions in population occurred when the internal combustion engine replaced horses in the 1920s and a major source of food in the form of grain spillage was lost.[213][214]
Declines have been particularly apparent even in North America, where the house sparrow is invasive in some states. Introduced to Philadelphia initially in 1852 the house sparrow rapidly spread across the nation. However, the bird has largely disappeared from the city nowadays and overall, it is estimated to have declined in North America by 84% since 1966.[215]
InSouth Asia, the house sparrow has largely vanished from major cities such asKarachi,Kolkata,Mumbai,New Delhi, andLahore.[216] In general, the house sparrow population has been on the decline in many Asian countries, and this is quite evident in India.[217]
Various causes for the dramatic decreases in population have been proposed, including predation, in particular byEurasian sparrowhawks,[218][219][220] possibly facilitated by the elimination of bushes which the sparrows use to hide,[205] electromagnetic radiation from mobile phones;[221] and diseases[222] such as avian malaria.[223] A shortage of nesting sites caused by changes in urban building design is probably a factor, and conservation organisations have encouraged the use of special nest boxes for sparrows.[222][224][225][226] A primary cause of the decline seems to be an insufficient supply of insect food for nestling sparrows.[222][227] Declines in insect populations result from an increase of monoculture crops, the heavy use of pesticides,[228][229][230] the replacement of native plants in cities with introduced plants and parking areas,[231][232] and possibly the introduction ofunleaded petrol, which produces toxic compounds such asmethyl nitrite.[233]
Protecting insect habitats on farms[234][235] and planting native plants in cities benefit the house sparrow, as does establishing urban green spaces.[236][237] To raise awareness of threats to the house sparrow,World Sparrow Day has been celebrated on 20 March across the world since 2010.[238]To promote the conservation of sparrow, in 2012, the house sparrow was declared as the state bird of Delhi.[217]
To many people across the world, the house sparrow is the most familiar wild animal and, because of its association with humans and familiarity, it is frequently used to represent the common and vulgar, or the lewd.[239] One of the reasons for the introduction of house sparrows throughout the world was their association with the European homeland of many immigrants.[81] Birds usually described later as sparrows are referred to in many works of ancient literature and religious texts in Europe and western Asia. These references may not always refer specifically to the house sparrow, or even to small, seed-eating birds, but later writers who were inspired by these texts often had the house sparrow in mind.[40][239][240] In particular, sparrows were associated by the ancient Greeks withAphrodite, the goddess of love, due to their perceived lustfulness, an association echoed by later writers such asChaucer andShakespeare.[40][194][239][241] Jesus's use of "sparrows" as an example of divine providence in theGospel of Matthew[242] also inspired later references, such as that in Shakespeare'sHamlet[239] and theGospelhymnHis Eye Is on the Sparrow.[243]
The house sparrow is very rarely represented inancient Egyptian art, but anEgyptian hieroglyph is based on it. The sparrow hieroglyph had no phonetic value and was used as a determinative in words to indicate small, narrow, or bad.[244] An alternative view is that the hieroglyph meant "a prolific man" or "the revolution of a year".[245]
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