The subfamilyCaprinae,[1] also sometimes referred to as the tribeCaprini,[2] is part of theruminantfamilyBovidae,[3] and consists of mostly medium-sizedbovids. A member of this subfamily is called acaprine.[4]
Prominent members includesheep andgoats, with some other members referred to asgoat antelopes. Some earliertaxonomies considered Caprinae a separate family called Capridae (with the members being caprids), but now it is usually considered either a subfamily within the Bovidae, or a tribe within the subfamilyAntilopinae of the family Bovidae, with caprines being a type of bovid.
Although most goat-antelopes are gregarious and have fairly stocky builds, they diverge in many other ways – themuskox (Ovibos moschatus) is adapted to the extreme cold of thetundra; themountain goat (Oreamnos americanus) ofNorth America is specialised for very rugged terrain; theurial (Ovis orientalis) occupies a largely infertile area fromKashmir toIran, including much desert country. TheArmenian mouflon (Ovis gmelini gmelini) is thought to be the ancestor of the moderndomestic sheep (Ovis aries).
Many species have become extinct since the lastice age, probably largely because of human interaction. Of the survivors:
Five are classified as endangered,
Eight as vulnerable,
Seven as of concern and needing conservation measures, but at lower risk, and
Seven species are secure.
Members of the group vary considerably in size, from just over 1 m (3 ft) long for a full-growngrey goral (Nemorhaedus goral), to almost 2.5 m (8 ft 2 in) long for a musk ox, and from under 30 kg (66 lb) to more than 350 kg (770 lb). Musk oxen in captivity have reached over 650 kg (1,430 lb).[citation needed]
The lifestyles of caprids fall into two broad classes: 'resource-defenders', which are territorial and defend a small, food-rich area against other members of the same species; and 'grazers', which gather together into herds and roam freely over a larger, usually relatively infertile area.
The resource-defenders are the more primitive group: they tend to be smaller, dark in colour, males and females fairly alike, have long, tessellated ears, long manes, and dagger-shaped horns. The grazers (sometimes collectively known astsoan caprids, from the Hebrewtso'n meaning sheep and goats) evolved more recently. They tend to be larger, highly social, and rather than mark territory with scent glands, they have highly evolved dominance behaviours. No sharp line divides the groups, but a continuum varies from the serows at one end of the spectrum to sheep, true goats, and musk oxen at the other.
The goat-antelope, or caprid, group is known from as early as theMiocene, when members of the group resembled the modernserow in their general body form.[5] The group did not reach its greatest diversity until the recentice ages, when many of its members became specialised for marginal, often extreme, environments: mountains, deserts, and thesubarctic region.
The ancestors of the modern sheep and goats (both rather vague and ill-defined terms) are thought to have moved into mountainous regions – sheep becoming specialised occupants of the foothills and nearby plains, and relying on flight andflocking for defence against predators, and goats adapting to very steep terrain where predators are at a disadvantage.
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