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Horseshoe Canyon Formation

Coordinates:51°25′24″N112°53′18″W / 51.42333°N 112.88833°W /51.42333; -112.88833 (Horseshoe Canyon)
From Wikipedia, the free encyclopedia
Geological formation in Canada
Horseshoe Canyon Formation
Stratigraphic range:Campanian-Maastrichtian
~73.1–68 Ma[1]
Horseshoe Canyon Formation atHorsethief Canyon, nearDrumheller. The dark bands are coal seams.
TypeGeological formation
Unit ofEdmonton Group
Sub-unitsStrathmore Member, Drumheller Member, Horsethief Member, Morrin Member, Tolman Member, Carbon Member, Whitemud Member
UnderliesBattle Formation,Scollard Formation
OverliesBearpaw Formation
Lithology
PrimarySandstone
OtherShale,coal
Location
Coordinates51°25′24″N112°53′18″W / 51.42333°N 112.88833°W /51.42333; -112.88833 (Horseshoe Canyon)
Region Alberta
Western Canadian Sedimentary Basin
Country Canada
Type section
Named forHorseshoe Canyon
Named byE.J.W. Irish, 1970

TheHorseshoe Canyon Formation is astratigraphic unit of theWestern Canada Sedimentary Basin in southwesternAlberta.[2][3] It takes its name fromHorseshoe Canyon, an area ofbadlands nearDrumheller.

The Horseshoe Canyon Formation is part of theEdmonton Group. In its type section (Red Deer River Valley atDrumheller), it is ~250 metres (820 ft) thick, but further west the formation is older and thicker, exceeding 500 metres (1,600 ft) nearCalgary.[4] It is of LateCretaceous age,Campanian to earlyMaastrichtian stage (EdmontonianLand-Mammal Age), and is composed ofmudstone,sandstone,carbonaceousshales, andcoal seams. A variety of depositional environments are represented in the succession, includingfloodplains,estuarine channels, and coal swamps, which have yielded a diversity of fossil material. Tidally-influenced estuarine point bar deposits are easily recognizable as Inclined Heterolithic Stratification (IHS). Brackish-water trace fossil assemblages occur within these bar deposits and demonstrate periodic incursion of marine waters into the estuaries.

The Horseshoe Canyon Formation crops out extensively in the area around Drumheller, as well as farther north along theRed Deer River nearTrochu and along theNorth Saskatchewan River inEdmonton.[2] It is overlain by the Battle andScollard formations.[4] The Drumheller Coal Zone, located in the lower part of the Horseshoe Canyon Formation, was mined forsub-bituminous coal in the Drumheller area from 1911 to 1979, and theAtlas Coal Mine in Drumheller has been preserved as a National Historic Site.[5] In more recent times, the Horseshoe Canyon Formation has become a major target forcoalbed methane (CBM) production.

Contact (red arrow) between the underlying marine shales of the Bearpaw Formation and the coastal Horseshoe Canyon Formation. Coal beds (black bands) are common in the Horseshoe Canyon Formation and were formed in coastal swamps.

Dinosaurs found in the Horseshoe Canyon Formation includeAlbertavenator,Albertosaurus,Anchiceratops,Anodontosaurus,Arrhinoceratops,Atrociraptor,Epichirostenotes,Edmontonia,Edmontosaurus,Hypacrosaurus,Ornithomimus,Pachyrhinosaurus,Parksosaurus,Saurolophus, andStruthiomimus. Other finds have includedmammals such asDidelphodon coyi, non-dinosaurreptiles,amphibians,fish, marine and terrestrialinvertebrates and plant fossils. Reptiles such as turtles and crocodilians are rare in the Horseshoe Canyon Formation, and this was thought to reflect the relatively cool climate which prevailed at the time. A study by Quinney et al. (2013) however, showed that the decline in turtle diversity, which was previously attributed to climate, coincided instead with changes in soil drainage conditions, and was limited by aridity, landscape instability, and migratory barriers.[6]

Oil/gas production

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The Drumheller Coal Zone has been a primary coalbed methane target for industry. In the area between Bashaw and Rockyford, the Coal Zone lies at relatively shallow depths (about 300 metres) and is about 70 to 120 metres thick. It contains 10 to 20 metres of cumulative coal, in up to 20 or more individual thin seams interbedded with sandstone and shale, which combine to make an attractive multi-completion CBM drilling target. In total, it is estimated there are 14 trillion cubic metres (500 tcf) of gas in place in all the coal in Alberta.

Biostratigraphy

[edit]

The timeline below follows work by David A. Eberth and Sandra L. Kamo published in 2019.[7]

Dinosaurs

[edit]

Ornithischians

[edit]

Ankylosaurs

[edit]
Ankylosaurs reported from the Horseshoe Canyon Formation
GenusSpeciesStratigraphic positionMaterialNotesImages

Anodontosaurus[8][9]

A. lambei[8][9]

Horsethief, Morrin, and lowest Tolman[8][9]

[Five] skulls, mandibles, cervical vertebra, caudal vertebrae, dorsal vertebrae, sacrocaudal vertebrae, sacrum, illium, ischium, partial pelvis, coossified ribs, femur, ?manual phalanx, tail clubs, numerous cervical half-rings and osteoderms.[8][9]

Anankylosaurineankylosaurid also known from the middleDinosaur Park Formation and closely related toAnkylosaurus.[10]

Edmontonia[11]

E. longiceps[11]

Upper Horsethief[11]

A skull with mandible, dorsal vertebrae, dorsal vertebrae, sacral vertebrae, caudal vertebrae, cervical ribs, dorsal ribs, humerus, radius, ulna, radius, ilia, ischia, both pubes (?), femur, tibia, fibula and osteoderms.[11]

Apanoplosaurinnodosaurid also known from the lower Dinosaur Park Formation and closely relatedDenversaurus.[12]

Euoplocephalus

E. tutus

Walter Coombs (1971) synonymisedAnodontosaurus lambei withE. tutus. However, recent studies suggest thatAnodontosaurus is distinct enough fromEuoplocephalus to be placed in its own genus and species.[8][13] Furthermore, all Horseshoe Canyon Formationankylosaurine specimens were suggested to be reassigned toAnodontosaurus.[9]

Hadrosaurs

[edit]
Hadrosaurs reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Edmontosaurus

E. regalis

Horsethief; likely present in Drumheller.

Hypacrosaurus

H. altispinus

Morrin and Tolman.

"[Five to ten] articulated skulls, some associated with postcrania, isolated skull elements, isolated postcranial elements, many individuals, embryo to adult."[14]

Saurolophus

S. osborni

Upper Morrin and Tolman.

"Complete skull and skeleton, [two] complete skulls."[14]

Marginocephalians

[edit]
Ceratopsians
[edit]
Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes
Uncertain or tentative taxa are insmall text;crossed out taxa are discredited.
Ceratopsians reported from the Horseshoe Canyon Formation
GenusSpeciesStratigraphic positionMaterialNotesImages

Anchiceratops[15][16]

A. ornatus[15][16]

Horsethief, Morrin, and Tolman; may have been present in Drumheller[16]

[Two] nearly complete skulls and [seven] partial skulls.[16]

A chasmosaurine ceratopsid contemporaneous and closely related toArrhinoceratops.[16]

Arrhinoceratops[17]

A. brachyops[17]

Horsethief[17]

A nearly complete skull.[17]

Achasmosaurineceratopsid restricted to the holotype specimen as Farke (2007) assigned the referred specimen ROM 1439 toTorosaurus.[18]

Eotriceratops[19]

E. xerinsularis[19]

Carbon[19]

Premaxillae, maxillae, rostral, supraorbital horncore with lacrimal, prefrontal, frontal, postorbital, jugal, epijugal, quadratojugal, quadrate, partial parietal, squamosal frill, braincase, syncervical, cervical vertebrae, dorsal vertebrae, ribs and fragments of ossified ligaments.[19]

A largechasmosaurineceratopsid that may represent asenior synonym ofOjoceratops from theOjo Alamo Formation.[20]

Montanoceratops[21][22]

M. cerorhynchus[21][22]

Upper Tolman[22]

An isolated braincase.[22]

Aleptoceratopsidceratopsian also known from theSt. Mary River Formation.[21]

Pachyrhinosaurus[23]

P. canadensis[23]

Drumheller and Horsethief[23]

[Two] partial skulls.[23]

Acentrosaurineceratopsid also known from theSt. Mary River Formation.[23]

Pachycephalosaurs
[edit]
Pachycephalosaurids reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Sphaerotholus[24][25]

S. edmontonense[25]

Tolman[24]

A frontoparietal.[24][25]

A pachycephalosaurinepachycephalosaurid also known from theHell Creek andKirtland Formation.[25]

Sphaerotholus
Sphaerotholus

Thescelosaurs

[edit]
Thescelosaurids reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Parksosaurus

P. warreni

Tolman

A partially articulated skeleton and partial skull.Athescelosauridneornithischian.
Parksosaurus
Parksosaurus

Theropods

[edit]

Maniraptorans

[edit]
Maniraptors reported from the Horseshoe Canyon Formation
GenusSpeciesStratigraphic positionMaterialNotesImages

Albertavenator[26]

A. curriei[26]

Horsethief[26]

[Two] frontals.[26]

Atroodontidtheropod that adds to the diversity of North American troodontids.[26]

Albertonykus[27]

A. borealis[27]

Upper Tolman[27]

Ulna, tibiae, metatarsals, manual ungual, pedal phalanxes and unidentified pedal phalanges.[27]

An mononykinalvarezsaurid specialized for digging insect nests.[27]

Apatoraptor[28]

A. pennatus[28]

Horsethief[28]

A palatine, mandibles, ceratobranchials, axis, cervical vertebrae, dorsal vertebrae, pectoral girdles, sternal plate, forelimb, ilium, femur, tibia and fibula.[28]

Acaenagnathidoviraptorosaur recovered as sister taxon toElmisaurus .[28]

Atrociraptor[29]

A. marshalli[29]

Lower Horsethief[29]

Premaxillae, maxilla, dentaries, teeth and numerous bone fragments.[29]

Adromaeosaurid; teeth indicate it may have been present across all members.[29]

Epichirostenotes[30]

E. curriei[30]

Horsethief, Morrin, and Tolman[30]

A maxilla, probable palatine, partial braincase, cervical vertebrae, dorsal vertebrae, caudal vertebrae, synsacrum, an anterior and a posterior cervical rib, a mid-dorsal rib, fragments of ilia, ischium, both pubes, partial tibia, and unidentifiable bones.[30]

A caenagnathid oviraptorosaur known from material previously assigned toChirostenotes.[30]


Ornithomimids

[edit]
Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes
Uncertain or tentative taxa are insmall text;crossed out taxa are discredited.
Ornithomimids reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Dromiceiomimus

D. brevitertius

Several specimens.type specimen

Anornithomimid

Ornithomimus

O. currelli

Junior synonym ofO. edmontonicus

O. edmontonicus

Drumheller, Horsethief, Morrin, and Tolman

Several specimens,type specimen

An ornithomimid

Struthiomimus

S. altus

Drumheller, Horsethief, and Morrin

Anornithomimid


Tyrannosaurs

[edit]
Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes
Uncertain or tentative taxa are insmall text;crossed out taxa are discredited.
Tyrannosaurids reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Albertosaurus

A. arctunguis

Junior synonym ofA. sarcophagus

Albertosaurus

A. sarcophagus

Horsethief, Morrin, and Tolman; likely present in Drumheller and Carbon.

Several skeletons and partial skeletons,type specimen

Atyrannosaurid which was the most common large carnivore in the area.[31][32]

Daspletosaurus

D. sp.

Suggested from the skeleton of an immature tyrannosaurid (CMN 11315), thorough analysis of this specimen supports a referral toA. sarcophagus.[33] An isolatedmaxilla and teeth from anEdmontosaurus bonebed were also mistakenly referred toDaspletosaurus, however all the tyrannosaurid material in the bonebed was confirmed to belong toA. sarcophagus.[34]

Other animals

[edit]

Mammals

[edit]
Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes
Uncertain or tentative taxa are insmall text;crossed out taxa are discredited.
Mammals reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Didelphodon[35]

D. coyi

astagodontidmetatherian

Didelphodon
Didelphodon

Other reptiles

[edit]
Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes
Uncertain or tentative taxa are insmall text;crossed out taxa are discredited.
Reptiles reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages

Stangerochampsa[36]

S. mccabei

"a skull, partial lower jaws, and partial postcranial skeleton"

Analligatoroid

Champsosaurus[37]

C. albertensis

"partial skeleton with partial skull"

achoristodere

Champsosaurus
Champsosaurus

Leurospondylus[38]

L. ultimus

"a partial skeleton"

aplesiosaur of uncertain classification

Basilemys[39]

B. morrinensis

"nearly complete shell"

ananhsiungchelyidturtle

Fish

[edit]
Color key
TaxonReclassified taxonTaxon falsely reported as presentDubious taxon or junior synonymIchnotaxonOotaxonMorphotaxon
Notes
Uncertain or tentative taxa are insmall text;crossed out taxa are discredited.
Fish reported from the Horseshoe Canyon Formation
GenusSpeciesLocationStratigraphic positionMaterialNotesImages
Boreiosturion[40]B. labyrinthicusEdmonton.Partial skull.Asturgeon.

Horseshoeichthys[41]

H. armaserratus

Anellimmichthyiform

See also

[edit]

References

[edit]
  1. ^Eberth, David A.; Kamo, Sandra L. (October 2020)."High-precision U–Pb CA–ID–TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada".Canadian Journal of Earth Sciences.57 (10):1220–1237.Bibcode:2020CaJES..57.1220E.doi:10.1139/cjes-2019-0019.ISSN 0008-4077.
  2. ^abPrior, G. J., Hathaway, B., Glombick, P.M., Pana, D.I., Banks, C.J., Hay, D.C., Schneider, C.L., Grobe, M., Elgr, R., and Weiss, J.A. (2013)."Bedrock Geology of Alberta. Alberta Geological Survey, Map 600". Archived fromthe original on 2013-07-05. Retrieved2013-08-13.{{cite web}}: CS1 maint: multiple names: authors list (link)
  3. ^Mossop, G.D. and Shetsen, I., (compilers), Canadian Society of Petroleum Geologists (1994)."The Geological Atlas of the Western Canada Sedimentary Basin, Chapter 24: Upper Cretaceous and Tertiary strata of the Western Canada Sedimentary Basin". Archived fromthe original on 2013-07-21. Retrieved2013-08-01.{{cite web}}: CS1 maint: multiple names: authors list (link)
  4. ^abEberth, David A.; Braman, Dennis R. (September 2012)."A revised stratigraphy and depositional history for the Horseshoe Canyon Formation (Upper Cretaceous), southern Alberta plains".Canadian Journal of Earth Sciences.49 (9):1053–1086.Bibcode:2012CaJES..49.1053E.doi:10.1139/e2012-035.ISSN 0008-4077.
  5. ^"Mine History". Atlas Coal Mine National Historic Site. Archived fromthe original on 23 August 2010. Retrieved9 June 2010.
  6. ^Quinney, Annie; Therrien, François; Zelenitsky, Darla K.; Eberth, David A. (2013). "Palaeoenvironmental and palaeoclimatic reconstruction of the Upper Cretaceous (late Campanian–early Maastrichtian) Horseshoe Canyon Formation, Alberta, Canada".Palaeogeography, Palaeoclimatology, Palaeoecology.371:26–44.Bibcode:2013PPP...371...26Q.doi:10.1016/j.palaeo.2012.12.009.
  7. ^Eberth, David A.; Kamo, Sandra (2019)."High-precision U-Pb CA-ID-TIMS dating and chronostratigraphy of the dinosaur-rich Horseshoe Canyon Formation (Upper Cretaceous, Campanian–Maastrichtian), Red Deer River valley, Alberta, Canada".Canadian Journal of Earth Sciences.57 (10):1220–1237.Bibcode:2020CaJES..57.1220E.doi:10.1139/cjes-2019-0019.S2CID 210299227.
  8. ^abcdePenkalski, P. (2013)."A new ankylosaurid from the late Cretaceous Two Medicine Formation of Montana, USA".Acta Palaeontologica Polonica.doi:10.4202/app.2012.0125.
  9. ^abcdeArbour, Victoria (2010). "A Cretaceous armoury: Multiple ankylosaurid taxa in the Late Cretaceous of Alberta, Canada and Montana, USA".Journal of Vertebrate Paleontology.30 (Supplement 2): 55A.doi:10.1080/02724634.2010.10411819.
  10. ^Paul Penkalski (2018). Revised systematics of the armoured dinosaur Euoplocephalus and its allies. Neues Jahrbuch für Geologie und Paläontologie - Abhandlungen 287(3): 261-306. doi:https://doi.org/10.1127/njgpa/2018/0717
  11. ^abcdSternberg, C.M. (1928). "A new armored dinosaur from the Edmonton Formation of Alberta".Transactions of the Royal Society of Canada. Series 3.22:93–106.
  12. ^Russell, L.S. (1940). "Edmontonia rugosidens (Gilmore), an armored dinosaur from the Belly River Series of Alberta".University of Toronto Studies, Geology Series.43:3–28.
  13. ^Penkalski, P.; Blows, W. T. (2013). "Scolosaurus cutleri (Ornithischia: Ankylosauria) from the Upper Cretaceous Dinosaur Park Formation of Alberta, Canada".Canadian Journal of Earth Sciences.50 (2):171–182.Bibcode:2013CaJES..50..171P.doi:10.1139/cjes-2012-0098.
  14. ^ab"Table 20.1," in Weishampel, et al. (2004). Page 441.
  15. ^abBrown, B (1914). "Anchiceratops, a new genus of horned dinosaurs from the Edmonton Cretaceous of Alberta. With a discussion of the origin of the ceratopsian crest and the brain casts ofAnchiceratops andTrachodon"".Bulletin of the American Museum of Natural History.33:539–548.
  16. ^abcdeMallon, Jordan C.; Holmes, Robert; Eberth, David A.; Ryan, Michael J.; Anderson, Jason S. (2012). "Variation in the skull ofAnchiceratops (Dinosauria, Ceratopsidae) from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta".Journal of Vertebrate Paleontology.31 (5):1047–1071.doi:10.1080/02724634.2011.601484.S2CID 86349131.
  17. ^abcdParks, W.A. (1925). "Arrhinoceratops brachyops, a new genus and species of Ceratopsia from the Edmonton Formation of Alberta".University of Toronto Studies, Geology Series 19:1-15
  18. ^Farke, A.A., 2007, "Cranial osteology and phylogenetic relationships of the chasmosaurine ceratopsidTorosaurus latus", pp 235-257 in: K. Carpenter (ed.),Horns and Beaks: Ceratopsian and Ornithopod Dinosaurs, Bloomington, Indiana University Press
  19. ^abcdWu, X-C.; Brinkman, D.B.; Eberth, D.A.; Braman, D.R. (2007). "A new ceratopsid dinosaur (Ornithischia) from the uppermost Horseshoe Canyon Formation (upper Maastrichtian), Alberta, Canada".Canadian Journal of Earth Sciences.44 (9):1243–1265.Bibcode:2007CaJES..44.1243W.doi:10.1139/E07-011.
  20. ^Wick, S. L.; Lehman, T. M. (2013). "A new ceratopsian dinosaur from the Javelina Formation (Maastrichtian) of West Texas and implications for chasmosaurine phylogeny".Naturwissenschaften.100 (7):667–82.Bibcode:2013NW....100..667W.doi:10.1007/s00114-013-1063-0.PMID 23728202.S2CID 16048008.
  21. ^abcBrown, B.; Schlaikjer, E. M. (1942). "The skeleton ofLeptoceratops with the description of a new species".American Museum Novitates (1169):1–15.
  22. ^abcdP. J. Makovicky. 2001. A Montanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta. Mesozoic Vertebrate Life, D. H. Tanke and K. Carpenter (eds.), Indiana University Press, Bloomington 243-262
  23. ^abcdeSternberg, C. M. (1950). "Pachyrhinosaurus canadensis, representing a new family of the Ceratopsia, from southern Alberta".National Museum of Canada Bulletin.118:109–120.
  24. ^abcLongrich N. R.; Sankey J. T.; et al. (2010). "Texacephale langstoni, a new genus of pachycephalosaurid (Dinosauria: Ornithischia) from the upper Campanian Aguja Formation, southern Texas, USA".Cretaceous Research.31 (2):274–284.Bibcode:2010CrRes..31..274L.doi:10.1016/j.cretres.2009.12.002.
  25. ^abcdWilliamson Thomas E.; Carr Thomas D. (2002). "A new genus of highly derived pachycephalosaurian from western North America".Journal of Vertebrate Paleontology.22 (4):779–801.doi:10.1671/0272-4634(2002)022[0779:angodp]2.0.co;2.S2CID 86112901.
  26. ^abcdeEvans, D.C., Cullen, T.M., Larson, D.W., and Rego, A. "A new species of troodontid theropod (Dinosauria: Maniraptora) from the Horseshoe Canyon Formation (Maastrichtian) of Alberta, Canada." Canadian Journal of Earth Sciences Early Online: 813-826. DOI: dx.doi.org/10.1139/cjes-2017-0034
  27. ^abcdeLongrich, Nicholas R.; Currie, Philip J. (2009). "Albertonykus borealis, a new alvarezsaur (Dinosauria: Theropoda) from the Early Maastrichtian of Alberta, Canada: Implications for the systematics and ecology of the Alvarezsauridae".Cretaceous Research.30 (1):239–252.Bibcode:2009CrRes..30..239L.doi:10.1016/j.cretres.2008.07.005.
  28. ^abcdeGregory F. Funston; Philip J. Currie (2016)."A new caenagnathid (Dinosauria: Oviraptorosauria) from the Horseshoe Canyon Formation of Alberta, Canada, and a reevaluation of the relationships of Caenagnathidae".Journal of Vertebrate Paleontology.36 (4): e1160910.Bibcode:2016JVPal..36E0910F.doi:10.1080/02724634.2016.1160910.S2CID 131090028.
  29. ^abcdeCurrie, P. J. and D. J. Varricchio (2004). "A new dromaeosaurid from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada". Pp. 112–132 in P. J. Currie, E. B. Koppelhus, M. A. Shugar and J. L. Wright. (eds.),Feathered Dragons. Indianapolis: Indiana University Press.[1]
  30. ^abcdeRobert M. Sullivan; Steven E. Jasinski; Mark P.A. Van Tomme (2011)."A new caenagnathidOjoraptorsaurus boerei, n. gen., n. sp. (Dinosauria, Oviraptorosauria), from the Upper Ojo Alamo Formation (Naashoibito Member), San Juan Basin, New Mexico"(PDF).Fossil Record 3. New Mexico Museum of Natural History and Science Bulletin.53:418–428.
  31. ^Coulton, Richard L. (1977)."The Theropod Dinosaurs of the Edmonton Formation, Alberta, Canada".Rocks & Minerals.52 (4):176–181.Bibcode:1977RoMin..52..176C.doi:10.1080/00357529.1977.11761949. Retrieved21 March 2021.
  32. ^Tanke, Darren H.; Currie, Phillip J. (7 September 2010)."A history of Albertosaurus discoveries in Alberta, Canada".Canadian Journal of Earth Sciences.47 (9):1197–1211.Bibcode:2010CaJES..47.1197T.doi:10.1139/e10-057. Retrieved21 March 2021.
  33. ^Mallon, J.C.; Bura, J.R.; Currie, P.J. (2019)."A Problematic Tyrannosaurid (Dinosauria: Theropoda) Skeleton and Its Implications for Tyrannosaurid Diversity in the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta".The Anatomical Record.303 (4):673–690.doi:10.1002/ar.24199.PMC 7079176.PMID 31254458.
  34. ^Torices, A.; Reichel, M.; Currie, P.J. (2014). "Multivariate analysis of isolated tyrannosaurid teeth from the Danek Bonebed, Horseshoe Canyon Formation, Alberta, Canada".Canadian Journal of Earth Sciences.51 (11):1045–1051.Bibcode:2014CaJES..51.1045T.doi:10.1139/cjes-2014-0072.
  35. ^R. C. Fox and B. G. Naylor. 1986. A new species of Didelphodon Marsh (Marsupialia) from the Upper Cretaceous of Alberta, Canada: paleobiology and phylogeny. Neues Jahrbuch für Geologie und Paläontologie, Abhandlungen 172(3):357-380
  36. ^X.-C. Wu, D. B. Brinkman, and A. P. Russell. 1996. A new alligator from the Upper Cretaceous of Canada and the relationships of early eusuchians. Palaeontology 39(2):351-375
  37. ^K. -Q. Gao and R. C. Fox. 1998. New choristoderes (Reptilia: Diapsida) from the Upper Cretaceous and Palaeocene, Alberta and Saskatchewan, Canada, a phylogenetic relationships of Choristodera. Zoological Journal of the Linnean Society 124:303-353
  38. ^B. Brown. 1913. A new plesiosaur, Leurospondylus, from the Edmonton Cretaceous of Alberta. Bulletin of the American Museum of Natural History 32(40):605-615
  39. ^Mallon, Jordan C.; Brinkman, Donald B. (2018)."Basilemys morrinensis, a new species of nanhsiungchelyid turtle from the Horseshoe Canyon Formation (Upper Cretaceous) of Alberta, Canada".Journal of Vertebrate Paleontology.38 (2): e1431922.Bibcode:2018JVPal..38E1922M.doi:10.1080/02724634.2018.1431922.
  40. ^Murray, Alison M.; Nelson, Luke E.; Brinkman, Donald B. (2023-08-08)."A new sturgeon from the Upper Cretaceous Horseshoe Canyon Formation in central Alberta, Canada".Journal of Vertebrate Paleontology.43.doi:10.1080/02724634.2023.2232846.ISSN 0272-4634.
  41. ^Newbrey, Michael G.; Murray, Alison M.; Brinkman, Donald B.; Wilson, Mark V. H.; Neuman, Andrew G. (2010)."A new articulated freshwater fish (Clupeomorpha, Ellimmichthyiformes) from the Horseshoe Canyon Formation, Maastrichtian, of Alberta, CanadaThis article is one of a series of papers published in this Special Issue on the themeAlbertosaurus".Canadian Journal of Earth Sciences.47 (9):1183–1196.doi:10.1139/E10-041.

Bibliography

[edit]
  • Makovicky, P. J., 2001, AMontanoceratops cerorhynchus (Dinosauria: Ceratopsia) braincase from the Horseshoe Canyon Formation of Alberta: In: Mesozoic Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, pp. 243–262.
  • Varricchio, D. J. 2001. Late Cretaceous oviraptorosaur (Theropoda) dinosaurs from Montana. pp. 42–57 in D. H. Tanke and K. Carpenter (eds.), Mesozoic Vertebrate Life. Indiana University Press, Indianapolis, Indiana.
  • Weishampel, David B.; Dodson, Peter; and Osmólska, Halszka (eds.): The Dinosauria, 2nd, Berkeley: University of California Press. 861 pp. ISBN 0-520-24209-2.
Hydrocarbon history
Depositional regions
Central Alberta
Authority control databases: NationalEdit this at Wikidata
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