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Hemipsalodon

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Extinct genus of mammals

Hemipsalodon Temporal range:41.2–37.0 Ma PreꞒ Ꞓ O S D C P T J K Pg N ↓ Middle to LateEocene

Skull ofHemipsalodon grandis

Restoration ofH. grandis
Scientific classification
Domain:	Eukaryota
Kingdom:	Animalia
Phylum:	Chordata
Class:	Mammalia
Order:	†Hyaenodonta
Superfamily:	†Hyainailouroidea
Family:	†Hyainailouridae
Subfamily:	†Hyainailourinae
Genus:	†Hemipsalodon Cope, 1885
Type species
†Hemipsalodon grandis Cope, 1885
Species
†H. grandis(Cope, 1885)^[1] †H. viejaensis(Gustafson, 1986)^[2]
Synonyms
synonyms of species: H. grandis: Hemipsalodon cooki(Schlaikjer, 1935)^[3]

Hemipsalodon ("half-scissor tooth") is anextinct genus ofhyainailourid hyaenodonts from the subfamilyHyainailourinae that lived inNorth America during the middle to lateEocene.^[4]^[5]

Description

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The skull ofHemipsalodon grandis is 45 centimetres (18 in) in length, with a lower jaw length of 34 centimetres (13 in). The most complete skull (O.M.S.I. No. 619), belonged to an old individual. It is powerfully-built, and overall superficially resembles the mesonychidHarpagolestes. The nasal opening is quite large. Thecanines are greatly enlarged. The anterior portion of the skull is broad anteriorly, but narrows down considerably posteriorly.^[6]

References

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^E. D. Cope (1885.)"The White River beds of Swift Current River, Northwest Territory." American Naturalist 19(2):163
^E. P. Gustafson (1986.)"Carnivorous mammals of the Late Eocene and Early Oligocene of Trans-Pecos Texas." Texas Memorial Museum Bulletin 33:1-66
^Schlaikjer, Erich M. (1935)."Contributions to the stratigraphy and paleontology of the Goshen Hole area, Wyoming. III. A new basal Oligocene formation".Bulletin of the Museum of Comparative Zoology.76 (3):71–93.
^McKenna, Malcolm C.; Bell, Susan K. (1997).Classification of Mammals Above the Species Level. New York: Columbia University Press.ISBN 978-0-231-11012-9. Retrieved16 March 2015.
^G. F. Gunnell (1998.) "Creodonta". In C. M. Janis, K. M. Scott, and L. L. Jacobs (eds.),"Evolution of Tertiary Mammals of North America, Volume 1: Terrestrial Carnivores, Ungulates, and Ungulate like Mammals", Cambridge University Press, 703 pagesHemipsalodon
^Mellett, James Silvan (1969)."A skull ofHemipsalodon (Mammalia, Deltatheridia) from the Clarno Formation of Oregon".American Museum Novitates (2387).hdl:2246/2597.

Mammals of cladePan-Carnivora

Kingdom:Animalia
Phylum:Chordata
Class:Mammalia
Infraclass:Placentalia
Superorder:Laurasiatheria
Clade:Scrotifera
Grandorder:Ferungulata
Mirorder:Ferae

Pan-Carnivora

Clade:
Pan-Carnivora

Clade:
Carnivoramorpha

SeeCarnivoramorpha below ↓

Order:
†Hyaenodonta

SeeHyaenodonta below ↓

Order:
†Oxyaenodonta

Family:
†Oxyaenidae

Subfamily:
†Machaeroidinae

†Apataelurus †Diegoaelurus †Isphanatherium †Machaeroides
Incertae sedis	†Machaeroidinae sp. (CM 2386) †Machaeroidinae sp. (FMNH PM 1506) †Machaeroidinae sp. (USNM 173514)

Subfamily:
†Oxyaeninae

Subfamily:
†Palaeonictinae

†Ambloctonus
†Dipsalodon^{(paraphyletic genus)}
†Palaeonictis

Subfamily:
†Tytthaeninae

†Tytthaena

other
representatives

†Altacreodus
Family: †Simidectidae	†Simidectes
Family: †Wyolestidae	†Wyolestes

Carnivoramorpha

Clade:
Carnivoramorpha

Clade:
Carnivoraformes
(i.e. Clade "A")

Clade "B"

Family:
†Quercygalidae

†Quercygale

†Gracilocyon/Oodectes
clade

†Eogale
†Gracilocyon^{(paraphyletic genus)}
†Oodectes^{(paraphyletic genus)}
†Paramiacis
†Paroodectes

Incertae sedis	†"Miacis" sp. (CM 67873 &CM 77299)

Clade "C"

†Dormaalocyon
†Vulpavus clade	†Palaearctonyx †Vassacyon †Vulpavus^{(paraphyletic genus)} (including subgenus †Phlaodectes)
Incertae sedis	†"Miacis" deutschi †"Miacis" exiguus

Clade "D"

Order:Carnivora
(i.e. Clade "E")

Suborder:Caniformia	(SeeCaniformia)
Suborder:Feliformia	(SeeFeliformia)

Incertae sedis

other
representatives

†Africtis
†Dawsonicyon
†Miacis

Incertae sedis	†"Miacis" boqinghensis †"Miacis" hookwayi †"Miacis" latidens †"Miacis" petilus †Carnivoraformes undet. Genus A †Carnivoraformes undet. Genus B
ichnotaxa of Carnivoraformes	†Falcatipes

Superfamily:
†Viverravoidea

Family:
†Viverravidae

Subfamily:
†Didymictinae

†Bryanictis
†Didymictis
†Intyrictis
†Pristinictis
†Protictis^{(paraphyletic genus)} (including subgenus †Protictoides)
†Raphictis

Incertae sedis	†"Deltatherium" durini

Subfamily:
†Ictidopappinae

†Ictidopappus

Subfamily:
†Viverravinae

Incertae sedis

†Viverravidae sp. (CM 71188 &CM 71189)

Incertae sedis

†Hyaenodonta

Order:
†Hyaenodonta

Superfamily:
†Hyaenodontoidea

Family:
†Hyaenodontidae

†Cynohyaenodon/Quercytherium
clade

†Eurotherium clade

†Alienetherium
†Cartierodon
†Eurotherium^{(paraphyletic genus)}
†Paenoxyaenoides
†Prodissopsalis

†Leonhardtina clade

†Leonhardtina

†Matthodon clade

†Matthodon

†Oxyaenoides clade

†Oxyaenoides

Subfamily:
†Hyaenodontinae

†Consobrinus †Propterodon^{(paraphyletic genus)}
Tribe: †Hyaenodontini	†Hyaenodon (including subgenera †Neohyaenodon^{(paraphyletic subgenus)} and †Protohyaenodon^{(paraphyletic subgenus)})
Incertae sedis	†"Pterodon" hyaenoides

Clade:
†Proviverrinae

†Minimovellentodon †Morlodon †Parvagula
†Allopterodon/Proviverra clade	†Allopterodon †Lesmesodon †Proviverra

†Afro‑Arabian
clade

Superfamily:
†Hyainailouroidea
^{(polyphyletic superfamily)}

Family:
†Hyainailouridae
^{(paraphyletic family)}

Subfamily:
†Apterodontinae

†Apterodon
†Quasiapterodon

Subfamily:
†Hyainailourinae
^{(paraphyletic subfamily)}

†Akhnatenavus clade

†Maocyon/Orienspterodon
clade

†Maocyon
†Orienspterodon

Tribe:
†Hyainailourini
^{(polyphyletic tribe)}

†Bastetodon
†Exiguodon
†Falcatodon
†Hyainailouros^{(polyphyletic genus)}
†Parapterodon
†Sectisodon
†Sekhmetops
†Sivapterodon

Subtribe:
†Isohyaenodontina
^{(polyphyletic subtribe)}

†Isohyaenodon^{(polyphyletic genus)}

†Pterodon
clade

†Kerberos
Subtribe: †Pterodontina	†Pterodon

Incertae sedis

†"Pterodon" sp. (DPC 5036)

Tribe:
†Leakitheriini

†Leakitherium

Tribe:
†Metapterodontini

†Metapterodon

Tribe:
†Paroxyaenini

†Paroxyaena

Incertae sedis

Family:
†Prionogalidae

†Namasector
†Prionogale

Family:
†Teratodontidae

Subfamily:
†Teratodontinae

†Anasinopa †Brychotherium †Masrasector^{(paraphyletic genus)} †Metasinopa
Tribe: †Dissopsalini	†Buhakia †Dissopsalis
Tribe: †Teratodontini	†Teratodon
Incertae sedis	†Teratodontinae sp. (BC 2’08) †Teratodontinae sp. (CBI-1-614) †Teratodontinae sp. (Locality BQ-2, Fayum, Egypt)

other
representatives

†Furodon
†Glibzegdouia

†Kyawdawia clade

†Kyawdawia
†Paratritemnodon

†Lahimia clade

†Parvavorodon

Clade:
†Boualitominae
^{(paraphyletic clade)}

†Boualitomus †Lahimia
Incertae sedis	†Boualitominae sp. (Locality BQ-2, Fayum, Egypt) †Boualitominae sp. (Quarry L-41, Fayum, Egypt)

†Tritemnodon clade

†Tritemnodon

Clade:
†Koholiinae

†Koholia

Family:
†Indohyaenodontidae

†Indohyaenodon
†Yarshea

other
representatives

†Eoproviverra/Tinerhodon
clade

†Eoproviverra
†Tinerhodon

†Galecyon clade

Clade:
†Arfiinae

†Arfia

Clade:
†Limnocyoninae

†Iridodon †Limnocyon^{(paraphyletic genus)} †Oxyaenodon †Prolaena †Prolimnocyon^{(paraphyletic genus)} †Thinocyon
Incertae sedis	†"Thinocyon" sichowensis

Clade:
†Sinopidae

†Acarictis

†Sinopa clade

†Prototomus^{(paraphyletic genus)} †Sinopa
Incertae sedis	†Sinopidae sp. (FMNH PM 59529)

ichnotaxa
of Hyaenodonta

†Creodontipus †Dischidodacylus †Sarcotherichnus
Ichnofamily: †Sarjeantipodidae	†Hyaenodontipus †Quiritipes †Sarjeantipes

Taxon identifiers
Hemipsalodon	Wikidata:Q107547795 GBIF:4834344 Open Tree of Life:4945303 Paleobiology Database:40916
Hemipsalodon grandis	Wikidata:Q118787131 GBIF:4974135 Open Tree of Life:4945305 Paleobiology Database:46866

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