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Goode's thornscrub tortoise

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Species of tortoise

Goode's thornscrub tortoise

Conservation status
Vulnerable (IUCN 3.1)^[1]
CITES Appendix II ^[2]
Scientific classification
Kingdom:	Animalia
Phylum:	Chordata
Class:	Reptilia
Order:	Testudines
Suborder:	Cryptodira
Family:	Testudinidae
Genus:	Gopherus
Species:	G. evgoodei
Binomial name
*Gopherus evgoodei* Edwards et al., 2016

TheGoode's thornscrub tortoise, also known as theSinaloan thornscrub tortoise,Sinaloan desert tortoise orGoode's desert tortoise (Gopherus evgoodei), is a species of tortoise that is native to the Sinaloan desert region. First described in 2016,G. evgoodei inhabits Tropical Deciduous Forest and Sinaloan Desertscrub biomes in Mexico.^[3] Its range may overlap in the north withG. morafkai, the Morafka's orSonoran desert tortoise.

Goode's thornscrub tortoise is named forEric V. Goode, a conservationist, naturalist, and founder of theTurtle Conservancy.^[4]

Taxon history

[edit]

Bogert and Oliver were the first researchers to notice a distinct morphology in the southern range of what was then consideredG. agassizii, but could not confirm a new species due to limitations of small sample size.^[5] Edwards et al. sampled 233 tortoises that represented Sonoran and Sinaloan lineages ofG. morafkai.^[6] The authors then conducted a large-scale genetic analysis that when combined with significant ecological and morphological differentiation, suggested that the southernmostGopherus "Sinaloan" population constituted a newly described species,G. evgoodei.^[3]

Hybrid zones forG. morafkai andG. evgoodei have been found in a narrow ecotone between the Sonoran Desert scrub and the Sonoran Thorn scrub in Mexico.^[3]

Description

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Gopherus evgoodei differs fromG. morafkai andG. agassizii in that it is flatter in its shell profile, has rounded foot pads and multiple enlarged spurs on the radial-humeral joint. It also has orange tones in the integument (skin) and shell and a distinctly shallower concavity on the plastron of males.^[4]

Habitat

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Gopherus evgoodei lives in hills and low mountains, and prefers slopes, boulders and rock outcrops. Like the Mojave and Sonoran desert tortoises, it digs burrows in the soil, but prefers to excavate under boulders.^[3]

Activity

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Gopherus evgoodei is active from June until November and it is assumed this corresponds with monsoonal rains and corresponding plant growth.^[3]

Conservation

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Gopherus evgoodei resides in threatened habitat and has a smaller distribution thanG. agassizii orG. morafkai.^[3] In 2018, the IUCN Tortoise and Freshwater Turtle Specialist Group recommended a re-assessment and re-classification of all sixGopherus species. This reclassification would moveG. evgoodei from Near Endangered (NE) to Vulnerable (VU).^[7]

References

[edit]

^Edwards, T.; Karl, A.E.; Rosen, P.C.; Vaughn, M.; Melendez Torres, C.; Kiester, A.R.; Goode, E.V. (2018)."Gopherus evgoodei".IUCN Red List of Threatened Species.2018 e.T90377823A144764819.doi:10.2305/IUCN.UK.2018-2.RLTS.T90377823A144764819.en. Retrieved19 November 2021.
^"Appendices".CITES. Convention on International Trade in Endangered Species of Wild Fauna and Flora. 4 May 2023. Retrieved9 May 2023. Note: Protected as a member of the Testudinidae.
^^a ^b ^c ^d ^e ^fEdwards, Taylor; Karl, Alice; Vaughn, Mercy; Rosen, Philip; Torres, Christina Meléndez; Murphy, Robert (2016-10-02)."The desert tortoise trichotomy: Mexico hosts a third, new sister-species of tortoise in the Gopherus morafkai–G. agassizii group".ZooKeys (562):131–158.Bibcode:2016ZooK..562..131E.doi:10.3897/zookeys.562.6124.ISSN 1313-2970.PMC 4768471.PMID 27006625.
^^a ^b"Gopherus evgoodei Edwards, Karl, Vaughn, Rosen, Meléndez-Torres & Murphy".The Reptile Database. Retrieved26 April 2021.
^Bogert CM, Oliver JA. (1945). "A preliminary analysis of the herpetofauna of Sonora".Bulletin of the American Museum of Natural History.83: 301−425.
^Edwards, Taylor; Vaughn, Mercy; Rosen, Philip C.; Meléndez Torres, Cristina; Karl, Alice E.; Culver, Melanie; Murphy, Robert W. (March 2016)."Shaping species with ephemeral boundaries: the distribution and genetic structure of desert tortoise (Gopherus morafkai) in the Sonoran Desert region".Journal of Biogeography.43 (3):484–497.Bibcode:2016JBiog..43..484E.doi:10.1111/jbi.12664.
^Rhodin, Anders G.J.; Stanford, Craig B.; Dijk, Peter Paul Van; Eisemberg, Carla; Luiselli, Luca; Mittermeier, Russell A.; Hudson, Rick; Horne, Brian D.; Goode, Eric V.; Kuchling, Gerald; Walde, Andrew (2018-12-18)."Global Conservation Status of Turtles and Tortoises (Order Testudines)"(PDF).Chelonian Conservation and Biology.17 (2): 135.doi:10.2744/CCB-1348.1.ISSN 1071-8443.S2CID 91937716.

Tortoise family

Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Subclass:Anapsida
Order:Testudines
Suborder:Cryptodira
Superfamily:Testudinoidea
Family:Testudinidae

Genera

Species of thetortoise family

Agrionemys

Russian tortoise

Angulate tortoise

†Hadrianus

†Hesperotestudo percrassa

Pancake tortoise

Leopard tortoise

†Stylemys

Testudo

Phylogenetic arrangement of turtles based onturtles of the world 2017 update: Annotated checklist and atlas of taxonomy, synonymy, distribution, and conservation status. Key: †=extinct.

Testudines

Kingdom:Animalia
Phylum:Chordata
Class:Reptilia
Clade:Diapsida
Order:Testudines

Genus

Chelonioidea
(Sea turtles)

Cheloniidae	†Allopleuron Caretta †Carolinochelys Chelonia †Eochelone Eretmochelys †Gigantatypus †Glarichelys †Itilochelys Lepidochelys †Mexichelys †Miocaretta Natator †Pacifichelys †Syllomus †Tasbacka
Dermochelyidae	†Arabemys †Corsochelys †Cosmochelys Dermochelys †Eosphargis †Mesodermochelys †Psephophorus
	†Euclastes †Peritresius †Procolpochelys †Protosphargis †Puppigerus

Kinosternoidea

Dermatemydidae	Dermatemys
Kinosternidae	Claudius †Hoplochelys Kinosternon Staurotypus Sternotherus

Testudinoidea

Emydidae	†Acherontemys Chrysemys Clemmys Deirochelys Emys Actinemys Emydoidea Glyptemys Graptemys Malaclemys Pseudemys Terrapene Trachemys †Wilburemys
Geoemydidae	Batagur †Banhxeochelys Cuora Cyclemys Geoclemys Geoemyda Hardella Heosemys Leucocephalon Malayemys Mauremys Melanochelys Morenia Notochelys Orlitia Pangshura Rhinoclemmys Sacalia Siebenrockiella Vijayachelys
Platysternidae	Platysternon
Testudinidae	Aldabrachelys Astrochelys Centrochelys Chelonoidis Chersina Cylindraspis †Cymatholcus †Floridemys Geochelone Gopherus †Hadrianus †Hesperotestudo Homopus Indotestudo Kinixys Malacochersus Manouria †Megalochelys †Oligopherus Psammobates Pyxis †Solitudo Stigmochelys †Stylemys Testudo

Trionychia

Carettochelyidae	†Allaeochelys †Anosteira Carettochelys
Trionychidae	Amyda Apalone †Axestemys Chitra Cyclanorbis Cycloderma †Drazinderetes Dogania †Gilmoremys †Hutchemys †Khunnuchelys Lissemys Nilssonia Palea †Palaeoamyda Pelochelys Pelodiscus Rafetus Trionyx
	†Basilochelys †Sinaspideretes

Chelydridae	Chelydra †Chelydrops †Chelydropsis †Emarginachelys †Macrocephalochelys Macrochelys †Planiplastron †Protochelydra
†Nanhsiungchelyidae	†Anomalochelys †Basilemys †Jiangxichelys
†Protostegidae	†Alienochelys †Archelon †Atlantochelys †Bouliachelys †Calcarichelys †Cratochelone †Desmatochelys †Iserosaurus †Notochelone †Ocepechelon †Pneumatoarthrus †Protostega †Rhinochelys †Santanachelys †Terlinguachelys
	†Adocus †Argillochelys †Bashuchelys †Ctenochelys †Prionochelys †Toxochelys

Pleurodira

†Araripemydidae	†Araripemys
†Bothremydidae	†Araiochelys †Arenila †Azabbaremys †Bothremys †Cearachelys †Chedighaii †Chupacabrachelys †Eotaphrosphys †Foxemys †Galianemys †Ilatardia †Inaechelys †Itapecuruemys †Jainemys †Kinkonychelys †Kurmademys †Labrostochelys †Nigeremys †Phosphatochelys †Polysternon †Puentemys †Rosasia †Rhothonemys †Sankuchemys †Taphrosphys †Ummulisani †Zolhafah
Chelidae	Acanthochelys Chelodina Chelus Elseya Elusor Emydura Hydromedusa †Lomalatachelys Mesoclemmys Myuchelys Phrynops Platemys †Prochelidella Pseudemydura Rheodytes Rhinemys †Yaminuechelys
Pelomedusidae	Pelomedusa Pelusios
Podocnemididae	†Albertwoodemys †Bauruemys †Brontochelys †Caninemys †Carbonemys †Cerrejonemys †Cordichelys Erymnochelys †Lapparentemys †Latentemys Peltocephalus Podocnemis †Stupendemys
†Sahonachelyidae	†Sahonachelys †Sokatra

	†Caribemys †Caririemys †Tacuarembemys

Phylogenetic arrangement of turtles based onTurtles of the World 2017 Update: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status. † = extinct.

See alsoList of Testudines families

Taxon identifiers
Gopherus evgoodei	Wikidata:Q22813438 Wikispecies:Gopherus evgoodei BOLD:853066 CoL:88BH7 GBIF:9004290 iNaturalist:500757 ITIS:1016071 IUCN:90377823 NCBI:1825980 RD:evgoodei ZooBank:125138E1-31AC-4FE5-8971-2F3D0A5113B8

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